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PAHHE-. 14 CPEAHEIOPCKILE : EEJIEMHVTTbI CEBEPACCCP MEGATEUTHINAE H
PSEUDODICOELITINAE
A H A A E M T . THf iA y r c c c c P c I { En P CK O E O T A EJ rE I r [ r E TPyALI nHCTATyTArEoJrornr.IR TEOOnBItKtr B H n y c a2 B g
A C A D E M Y O F S C I E N C E SO F T H E U S S R SIBENIAN BRANCH TRANSACTIONSOF THE INSTITUTE OF GEOLOGY AND GEOPHYSIC$ VoIume
2gg
V. N. SACHS,T. I. NALNJAEVA
THE EARLY AI.[D MIDDLE JURASSIC BELEMNITES OF THE NORTH OF THE USSR
PUBLISHING nOUSE tNAUKAT ' M o o K B at 9 ? 5
TI.H. CATtC,T. M: HAJIbHflEBA
PNIHE-
W CPEAHEroPCHIM BEJIEMHI{TbI CEBEPACCCP MEGATEUTHINAE N PSEUDODICOELITINAE
IISAATEJIbCTBO
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ateuthinae u
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(ceu. PasomrcaHbr 6eaeunnrsr noAcer.aeicra Megateuthinae pacnpocrpau Pseudodicoelitinae (ceu' Duvaliidae)'
""loi"uihid"") BeHIIbIenaCeaepecccPBoTJIoxeHI{gxBepxHeftqacruguxnefipPblr a sanse$cs 3a&flIoquTeaunoft cpeAneft ropbt r{ HtrsoB KeJuIoBes' IGnra 6eaeunnroB Iopbl }I paHomtca'l{'o nocBsue'Hblx aBTopoB, pa6br i.p",
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HeroMenaro6[raanuuxHareppl{ropsrCeaepaCCCP'AsropstAapro6u MeJIat cocraBJrgloulux HaAca' u,ylo KJraccnQuxaunrc 6eaer'rHnrog ropbl PaccvarpnBaloTcs cTpsTIr* Duvaliaceae. Cyiindroteuthaceae ,r.1"r"" u cperutefi lopbl Ha Qeaepe HuxHeft 6eaeprnuroe snaqeltrte rpa$nxecroe BosMoxHble ycJloBng I{x CCCP, $unoremra oElcblBaeMblx noAceMeficrn' pacpercfi"r, reorpa$nqecKoe ,rgrci'n, B qacrlrocrrr naJreoreMneparypnrtfi
IB
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Hacmfiuas KHr{na sBnsercs saBepuapuefi a cepnr{ pa6or aETo}oB, 11ocBsr[errr HbIx omrcaHlrlo lopcls{x tt paHHeMenoBblx 6enevnntoa Ceaepa CCCP. B neposlx AByx lgrf,rax (carc, HaacHseBa, 1964 1966) 6rran omcaHbr ropcf,[e u paHHeMerroBbre 6eaeuunru Ceaepa cccB nprnanrre)Karrue r cepreficray ,Cylindroteuihid"". Tp*", I(Hrra (Caxc, Hantunreaa, 19?O) saxrroqaer omrcanre rrepBbrx Tpex rpynn pagHgr cpeABelopcxilx 6eaeunrtog Ceaepa CCCP: Nannobelinae n Passaloteuthinae IGt ceM. Passaloteuthidae r ceM. Hastitidae. 3rr rpyrurbr cocT.ranruor ocgoagol @n ror"rnaexcoB 6eaerruroB B Toape n aaJreEe ceBeplrbrx o6nacreft cccp. Hnxe ollrcslBalo,cs noAceMef,crso Megateuthinae ns cev. Passaloteuthidae 11 rroAcer'retcrao Pseudodicoelitinae Duvaliidae, nan6onee ulrporco pacupocrpa"" ""t. HeHHbIe B cpeAHefi rcpe Ceaepa CCCP. Aaropu uMeJrr{ B cBoeM pacnopsxegrr roJulerurro ot(oJro 15OO pocrpoB HasBaHHbIx noAceMeficrs. paccuarproaeur,re 6e. JIeMHI{TII rpno6perapr nan6oaburee snaqenf,e Arrc crparxrpaftrrecroro pacvrene_ Hlrs cpeAlrercpcxl|x oTJtoxeHl{l Ceaepa r Aansnerp Bocrora CCCP. G1[sagbl raf,xce gonoJlHlrreJn'Ho cpeAHercpcf,re Cylindroteuthidae, roTopbre He Bouar B nepBbre ADe KHI{r.ll o qnarlrApoTe}Frf,Aarq Hcropns uswenns r{ MeroAlrxa uc-cJreAoBaH[s palrne- r cpeAteropcf,xx 6enev_ HItroB yxe 6rtmr A@Taroqno rro4)o6uo ocgeueHbr agTopaMu B [peAbrAyrqefi pa6ote.. JlonoaHl{Tenbllbre cBeAeE[c Aal1r,t H1;xe npn olrgcagrr11 cooTBeTcrB)nougx Taf,F corroB. B ynor..renyrofi Bbrue pa6ore aBtopoB lgZO r. qpnBegen crpararpaftrre_ cPfi oqepr ornoxeHnfi nrlcref, rophr rr rurxlref, qacTtr cpenref, rcprr Ceaepa E Bocroxa cccP Ilaronero c rPl{Bssrofi x paspesaM ocnoBnbrx c6opoa 6ener,rnrma. B Hacrocuefi f,nrre Mbr He co*mt ueaecoo6pasnblM AaBarb noAo6noe xe crparunpa$rrecroe oltrrcanrre 6onee Bbrcorqx nopnsonToB cpernefi rcpu Cesepa CCCR nocrorrbry oso 6srao roJrbxo uro ony6nutroBano GBMeaearrror (lgiC). floetorry sAecb MbI orpanrtl[Mcs rrnub nsrorteurrev o6uux crparlrpaSr.recrrx EhtBoAoB no 6eteulrf,Tarvr nnxnefi n cpeAuef, npn. Crparxrp"6usecrae pas6rara lulgel r cpeAnefi ropu Ea spycbl, rroAbspycbr f, sorrrr, BbrAerrseMhre Fns EopearuHoro [aJleqsooreorpa$r'recKono [osca (a parnefi rcpe o6nacnr), a sacrosqefi rcIune Bssra rpEHsras na Crparurpa$r'recroM coBeuagru rro Mesosop Gr64pr n llanssero Bocroxa s Hoeocr6rpcre n Lg12 n (caxc, ,earrc r ,qlr., tgizl. B pa6ore 1970 t. AAEo trotlnoe flepeqrrcreure f,crroJrbooBannbrx MarepgaJroB rlo paHlrb a cpeAHelopcruM 6enerrrturaM, Halmrrai or 3err,ran Qpanua-HocrQa ua sa[aAe no 6accelna Koaur"ot a ceBepnoro rr sanaAnoro uo6epercrt Onolcpro Mops Ea Bocrof,€. 3a [ocJIeAIIf,e nolbr (1968-19?3 nr.) aBTopbr AorlorrxnreJrbno o6pa6orarrn roJureruru 6erevnlroa, co6panHHe T.H. Hanaus."ol na no6eperGe Ana6apcrol ry6sr r Axa6apcf,ono oarrlrBa, Ha rr'oBe Yprcrrrr-Tyr.ryq B HxsoBbgx p. Gteuer I Ha p. I(ermu4Pr C.B.Meneqrrnofi rG p. tlepnorye6ertrofi (Bocrcrrorl Taft'Grp) II Ha p. Hlole (6acc. fleropar), T.H. I(npxHof, s Hrsogbsx Jlenrl, Ea p. Brurrcl c Ea ux 4pnrorax. ,0,ag cpasHeHlts ncrtoJrbgoBalor raror(e c6oprr palrneropcrorx 6ereMEEToB Ha reF pf,Topf,n Opanuru, Ee.nsnur u /lprceu6y?nq caerarnbre &FL Carcor..r s lg62 r. Ea glcrlpcnax Bo nperrre II MeloynapoAnono roJuror@H)rMa no ppe, n roJurerqf,s
u A.A.AaripaggeruuHc6axcp{x 6enevnuroB, co6pannaa s 1969 r. a lGpnarax A.C" caMH. BbIBOIaMH B rracTr{ KEI{1ll HapsAy co cTpaTlrrpasnvec6ltMn -El garntoqnteagnofi paclpgcTpaHefeorpa$u'recXOe pagcMarpnBarcTgq $uaorenus, OTAenrrEbIx rnaBax XI{3HI{' H, narOnet5 nOCKotIE'EUe Om{CbIBaeMbIX rpyln 6eaer'rfrnrOB I{ yCnOBIrg Hx paHEeMeJIoBbIx 6eneNaEuros n lopcxilx Ky AaEHOfi rnurofi saBepuaeTcs HsrIeHHe nnaccnsnxaurn yTOtIHeHHyIo noMecTuTb seo6xoAnMblM Mbr corrJtr{ ceaepa cccP, ony6alrroaapgas aBTopsrcpgprx rr Me1oBbrx 6eaeuEI{ToB. Tarcag 16,;raccnQutrAq11gt o6pa6orrr Mt{ B 1967 r., noltBepFnacb psAy yTOqHeHI{fi nocne r.aonorpa$nrlecKofi 6epaHgeMenoablx I{ topcffix rpylm CCCP o6flacTsx Bcex IIsBecTHbIx B ceBepgbrx EX nOqTU He 6bllo TO 6eleME[TOBr
[email protected] Xe KacaeTcs HTO aeMHltTOB. oorenrou ALs D CeBepEbrx pafionax Flpaeuu, n IIoeToMy oEI{ Re Moryr ctty)lc{Tb rro5r.renng. uouorpaSnqecroro omrcaHrK) paccMarprrBaeMblx s !|aE' flpexAe qeM flepelrn rcrcrreMaTrrrrecroMy' (nopo6rree eTo cAeJIaEo aBF yrasaTb Eeo6xoAI{Mo Bofi xprne rpy[n 6enevm'oa, ,ropaMu a pa6ore 197O r.), raE{e napaMerpbl pocrpoB ilcnoJrbsylorcs IIpI| oflllpocrpa y BepuI{HbI aJIEcaHngrq 3a 1OO% npuH[Masrcs clrHHO-6prOruUO[ nllaUetp napaMeTpblz fiIIt'F ocTarlbtlble Bce Beoabr ( CB ), no oTHolreHurc K HeMy paccqnTaHH qacTu 6oxosoil lIIaMeTpyBepulf,Ebtafrsseoilbl pocTpa-(nu), Ha nocJreaJlbBeoJlspgofi paArryc y Bepun'br arrbBeoJlbr, o6uat Arlr'a P@rPa, ne^'epenn41 G;)!-;p;Hofi (c6) AJIxHa npl{Bepunnnofi qacTn n AlaMsTpbI cn[Bno-6prcrunofi n [peAlloaaraeMan qacru y BepereHoBnAHbIX I|JII{ llpnBepuuHlrofi (66) y vecra nepexoAa r n o"*""fi pocrpoB y Mecra Han6ofirurero pacuupenlrs pocrpa. Ilag poctpoB BepereuoBf,AHol rra - AJMlrbI rrocJrearbBeollgptrofi qacru ''o orL 4opr*rr Bblqucne'o ra,o,.e s'arre',,e pOCTS g AIraMeTpy y MeCTa nar6Orn uen' pa.,u'pe',,s srm'E.rSpKlrltroMy H.ure'n,o lur@rpa. HeuepsJ1rcb aJrbBeoJrrlprorfi ynon y Bepur[Hbt aJIbBeoJIbI B cllltnHMplouttofi pocrpbl 6onsurefi xx rocronbry r(ocru r{ Bepurrl15rl yron -. AJre M"g"i"uthinae, n qacrb*t c)Karbr c 6oroa, s cmrnno-6p,ouBofi rurocrcocrt\ ptrs Psgudodicoelitinae rurocrocrtt. s 6oroaofi Cylindroteuthidae pocrprr B oaBr{crrMocrrr c,r pasMepo', rroApasAeJrsprcg Ha oqeEb xpFItIHe c p"""tpOr"r 6Oaee 2OO r"c"r, ryymrye Oor,a" IgaM€r,rpgr"r 6Oaee 30 r'1"r il ItIIIHot " I{ nrnrnofi MM 1O-2O c AraMerpoM cpeAlrne Mrvq 1OO-2OO u MM nrurnofi 2Cl€O rr'$"r, r'rendre 30-60 c AtraMerpor,r 5-1O MM rr pulrof uu, ue6onru[e 60-100 gnrrnofi r"rv' 30 MeBee c luaMerpoM Menee 5 r'tr"r n pa6ore Ee rIpItBoAI{Tcf,' Taf, rar Ao cl{x orscarue {pag,rorogoB B nacroguet B paMrax Hlrotu,,x rarcogoB - or ceuefi-' nop He yAaJrocb HaMernrb ux pas'rq,,s $parMoroHoB I| EbIsEIIeHse crBa 11 B11xe. Aaropsr cgr{Tarcrr rrlareJlbnoe ngfrerre sS nepBooqepeAtDrX 38r oAHOI ClqreMarrtor AtIg XCIIOIIbSOBaITRS UX EOSMO'(HOCTeI xqqneAgaa1ats 6e.1ei'rturr9a. Aaq AgtlbnefturerrO yrny6UeUgofp neoxpagsrcg s Hosocu6xpcxe, a Myeee Hucrzryra O6pa6oraglrbre rorurenrx @ A.y" cccP Uff Araner',rnn orAereHns e6rpcrono romx r reggnsrrr
AH C@P) rcs I.Ie87. flpr nonroroare pa6oru tr neqaru 6onauyn troMorqb aBTopaM orrasaJrlrJI.ELCaT'|poclpoB IpoIrsEoAEIocb s HIT CO QororpaqnpoBa'se r.-'--r @EoBa n M.B.)t*ascrag. AtI CCCP &o. fopryuoaUu. O6pa6orra Mareptara s pa6ora EaA pyrorncbp paqAe,rsracb Mer(A)r aETopaME M,esoteuthis, Hol,cobelus, creAlrrcruglro6pasor.r. Ornrcaus poAoB iiro"orlites) B.H.CarcoM, ottttcanf,s poAgB ParameLenobelus u pseudodicoclites DbrrroJrueHbr gateuthis s cylinikoteuthis EblnorEeHbtT.lL Hansnseaofi. o6rore rrIaEbI Banrcac EsrraBuuMtrcs Ebl aBropaMrr coBMecrEo, c pasAeaeBreM EX B COO|DEerCTBlru rpynxaMf, cr|creMarnqecMMr
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D ICavCO PaME s, tomercac
HAlICEMEIICTBO CYLINDROTEUTHACEAE SACHSET NALNJAEVA, I9?O
pAssALoTEUTHtDAENAEF, rgzz norcENtsftcrso MECATEUTHINAE SACHS ET NALNJAEvA, t96z cEMEficrBo
pocrpu oT crrJtbno ytrJMHeHHbrx Onncantre. Ao tropoTtorrq c aJlbB@rrotr, ea_ t{r{Marouefi oT L/2 no L/Lo ArnHbI pocrpa, c xopouo BtlpalreHHbIME rynBepurrHr HbIMI{ 6opoonav[ fi HepeAro c paArraJrstrofi urrpuxoBarocrbp B flpngepu'rnnof, .,a'Tlt'
1956; PseudohostitasNaefr- Lg22; LissJious,.igls. cpaB He Ef, e. Pocrrtt Mega-teutliin"" tvotr*"ecrct goprcr Moryr EorroMrHart pocrpar Passaloteuthinae' (pon orthobelus) r pocrprr Nannobelin"", orrF*lscb oT Tex u ApyTr|x pasEf,TEeM upnDepurrnrGtr 6oe€q. groT xe ryf,sgar otJrf,_ tlast cy6umlnplFreclse pocrpht Megateuthi nae or pocrpoB cy6urrunqprrecnol -catatcuthis). yroMsHltrblx Bttue norceuelco (poas- Braciybclr." pocrr. r,tf M.esateuthinaec ropoluo paaarrol " 6prournofi6opooaol (Acrococlitcs, 9u.belus) 6rursM no BHeurEefr@pue f, posrpaM Cylindroteuttriaaer,oarrgqaacbHolcoqr nocJreAlrrx npncyrcrBreM 6prcruno-6o[oBrtx-6opooa. lQorrre rod, Megateuthinae oTrll{qaprcs oT Passaloteuthinae n Cylindroteuthidae BarF1aeM cy6rornrqecloFo oTHOCHTerbHo
rppoTf,ono
poqrpa
Ha HAqa,IbHblx
CTAFFAX paSBETEl
n
a He ]rsro4 unporof, 6pnunrol'6opoonu. BospacT n reonpa$urectroe p a c r r p o c r p a H e E f , € . H r l c r s s r c p a( r _ e y r * ranr?-nnnnc6ax-toap) - cPeAHgs lopo - Hrsbr aepxnel ropbr (nrxnd reuoaefi). Fnpona, ceaepuan Aopnna, Iorgac Ar,aeprra, cn6nps, AancxHf, EbCrOr, ceaep nas Avepnra.'Ha ceBepe cccP poAbr salpin[oteuthis, pseudohastites n Homaloteilhis ne o6napyxeubr.
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Megateuthinae ra6anqapo4oBnoAcerueficrBa Onpegeanrenb6as
nrC C
c xopouo pasBlTblMf, f,ntr cy6UrnrEAplrsecxlte Pocrprr cy6rourrecxle orEoclrr'eJrbEo Kb craEf,sx HaqafibHbrx Ha rpf,BepuuEsru*ru 6opoeAaMu, pomxe cY6ronrrecxf,e.
1.
flonceveilcrgo
JIITH!
rur6o p)ryn
Megateuthinae Sachs et Nalniaeva, 1964
CTI'IE
3
6opOercf ' ' ' ' ' ' Z (S ). XOpOuO pagBnTbl llp,,Bepu,,B'bre cnnnrro-6oK.Bble pocrpbl cy6ro'n'rec'{e 6prcurnas 6opooAa or\cyrcrByer' nprrs"pnr'nnae 5 i4i: rml gy6tuJt[IrAplfrecxIe.
poru traF cMet
MEHI
Poa Masoteuthis Lissaious, l9l5 caa6o, Pocrpbl cy6rorur'recxile 4 (S ). npnsepu11pgas 6pnrunaa 6opoona EblpaxeHa I{JIu xoE(qecne. Pou P otamcgoteuthis Gustomesov, 1955 S ( 6 ). npnnepulrrgrrbre crurnnor6oroBue uEJIrrlrjll}rqeclg{e.
6opoenrr BblpaxeEbl caa6o, pocrpbt
cy6-
Pss Acrococlites Lissaious, l9l5 g ( 1O ). Pocrpsr cr{Jrb'o yIJrr{'eBHbIe BepeTeHoBHAHofi EJI'{ cy6rurnnunprrecrofi $opt*tst. Poa Pscudohastites Naef, 1922 emtp@TPr pocTpbl cru15go yAJIrHeBtrrIe cy6rUlrmllApmecKoil 1O(11),HueeTcs cY6xoruauecrofi QoPurt.
12.
( JIlIqU
xo{ oTc
6opoenaun xopouo pasBrTa 6 (12 ). Hapany c ,,p,nBepunEErME CTUTUUO:.S.K.EbIMH 6prournas 6opoeAa. sacTlr. ?(11,). Bprorunas 6opooAa HaxoAnTcs B Epl{Bepunnnofi s 6onsuluttcrBe $uep€Ecy6rrruutnAp]rqecpre' nnn g'(g). cy6xornrrecx[e eocrpsr Eo yA4nBeHHbIe.
AbI I Acrc Bepu KOT€
stit c e M (
t eutt AOnC F cpc 6orc
HATI€
utttt
6one NAAH
Salpingoteuthis Lissajous, 1915
6oposna [poTsrllBaeTcs 11(12).Eprourran Posrpa' PocrPbI cY6ronuveclole' pon Holcobelus Stolley, 1927
acul
I,ites
Psa Homaloteuthis StolleY' 1919
Pon
T 1O
qactu oT Bepu,'llbl Ao aJlbBeoflspnoft
cox ; BI{A E},}I OTM(
6opoenaMu IIp]rc]THapnAy c IIpIIBepunHHbIMH CnUUUO-SoKoBbIMIIn 6puunofi pocTpbl cy6rornuecxse 6opoerct, 6proruUo-6oKoBble cTByroT rp"".pt"Hgble oqeHb KpyIIHble. Pon lrlegateuthis Bayle et Zeiller, 1878
npnc ( Sch L84
t
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Pon
A c r o c o e l i le s L i s s a j o u s , l 9 l 5
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p. 54; StolMegateuthis (pars): Bayle et Zeiller, J819, pl. 26; Lissaious, 1906, S' 1ll' 1920, iliilow-Trummer; S. 36; ley, --"iiiiirtiti, 1919,
May.r-Eymar,1883,S.641,Werner,L912,S' 126' Megabelus(pars)',-n*ot,1914rcrp'13'1966'crp'1O8' Acrocoelit"Jrli""ajous, 1915,p. 10, 1925,p. 16,Kolb.,L942, S. 157'Roger, 1952,p. 710. (pars):,Abel,1916,S. 134,Stolley,1919,S. 35, Biilow-Trummer' Cuspiteuthis s . 1 0 1 . 1920, 1969,S' l8l' Naef,1922,5.238,Schwegler, Odontobelzs: Mesoteuthis(pars): crp. 158. 8
Ir(petvroasu,1932, ctp. I'
Lg4-7, crp' 201, 1958t
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19C AaM,
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poAa. Belemnites oxyconus Hehl in Zieten, 1ggo. .[|etac pxnofi Tun 11ar nr OPl. Onucanl{e. Pocrprt a 6oasufiHcrBe cpegHue, cy6roru.recrc{e, pexe cy6ruJlfiIrAplrqeclsre' c saocrpennoft nepuuHoft, oarl{Mapueft uenrpaJrbnoe notro)Kegr{e, rm6o cMerueHHofi r cnunnoft ctopone, co c)rclrbtv c 6oroB oBaJrbgbrM, pexe oKpyrJrblM I{Jr}r c}KarbIM B crmHHo-6plounoM HarrpaBJren[n rtofiepeqHbrM ceqegr{eM, q"T cunbHo yAJIrHeHHbrx Ao roporrs{x (tla - or 2OO-BOO go IOOO). y repuuusr xopotuo pasBl{Tbt ADe clrlrHHo-Sorosue 6opoegrt n 6prounas, rrporsnsBarouascs [IroDAa Ao L/3-L/2 flocJleaJlbBeonspnoft qacru. BepruuHa aJrbBeoJrbl r{ oceBas [r{Hr{s cMeIIIeHbI x 6p0uirofi cropoHe. Ha Ha.Iaabnbrx craAl{sx pocrpbr cy6ronnxecxfie, c MEI{bIIII{ME
tecxile
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TIA 'ICNA Y BsPOCJIbIX
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Buaoeoft cocras. ortrlcanbt 34 srAa, a rapKe s 3ananxofi Eapone oxoJro 10 Qopr'a nol AsofiIIbIMI{ EarMenoBarrrrsMu (rpynnrr qacrrrrrgo -CCCp $opr"r tripartitus, acuar.ius, compressus Quenstedt ). Ha Ceeep. BcrperreHbl g auAoe. CpanHeHI{e. Ot Apynrx PoAoa noAceueficrBa Megateuthlnss pocrprr AcrocoeI'ites orJrnqarcrrcs npn npeo6tanapuefi cy6roHnrecrofi cxerofi c 6oros Qopue ga_ JIr{q[eM Tpex xopouo Bbrpa)Kelrlrblx rpnBepurr{Hnbrx 6opo.aa _ 6prorunofi r uapot cF{It_ t{erxo Bbrpaxengaa 6prcuHas noSg*o"srx. 6opaaAa oTnrrraer pocrptt Acrocoelites or c)oAllblx c lruMu no @pve pocrpoo Mesoteuthis, y KoTopbrx 6prcunot 6opoe- \ AbI HeT I{JII4 oHa Ha)@AI{Tcs B saqaroqnoM cocrognrn. ?ror )Ke rpHgEaK orar{gaat ) Actocoelites oT cpeAnelopcrrx Paramegateuthis,uMepu[x caa6o pasBr{T}rronpr-/ Bepulrnnyro 6prorunyro 6oposAy, pocrpsr poAos pseud.ohastifes n Salpingoteuthis, KoTopbte TaK)Ke uMeror rrpr{Bepuxgnbre 6prorurcte 6opoealr, ornusarorcs,z p.seud,oha_ stif as - cuJrbHo yAtIIrHeHHoftnepereHoBr{AHofi - H€uMsHsopvo*, Salpingoteuthis eM CI{JIbHOyIUIHHeHHOnO pocrpa H npr{cyrcrBr{eM e[r{pocrpa. Pocrpsr poAa Megateuthis npu caa6oM pasBrrnr 6prcuuoft 6opoeAbr BbrAenrrrorcs ro rlp1acyrcrBgro Ao[oJurr{TenbHbrx [pr{BepuuHlrbl x 6oposg. Haan'rne [pnB€pturunoft 6prounoft 6opoearr c6ar>xaer pocrpbr poAa Acrocoelites c pocTpaMlr cevefic'rsa Cylindroteuthidae. y nocrre.Auro( Her crrrrrrur Oanaxo 6orcoaslx 6opaan u na6nrcAaerrcs yffrorlegr{e 6pnunrofi cToporrbr, pocrpbr Ha HaqaJrbHbD( cTqnuF( rrMelor cy6unnuuaprtrecKyro 6prcruxan 6opoOna $opuy, 6onee uupoKaq. 3aveqaHug. Pocrpbl c npl{Bepuunnofi 6prounofi 6opoegoft rosB.nsroTcs e 3anarnoft Fepone, a pxnoft qacru @Pf eue B paHHeM reTTartre ( Belernnites praecox Schwegler (Schwegler, 1939, 1962). Onrraro yBepemro orgecru yr(aoauxbrf, BI{A K poAy Acrocoelites nerssh us1"a oTcyrcrBns lanHbrx o BHyrpeHHeM crpoe_ Hur{ pocrpa. B rrosAHeM rerranre r{ cHHeMrope rrpeAcraBnTeJrn Acrocoelites He oTMerralorcs. B nuxHeu ruruHc6axe 3anatHoft Eapomr Acrocoelites yxe 6eccfiopgo rpl{cyrcrByrcr lA.trabecula Lissajous , Belemnites aff. apicicuruatus Schwegler, (Schwegler, 1969, crp.125l, B.paxillosus numismalis Quenstedt (Quensr"?;,' 1846-1849, crp. 39g, Ta6r. 23, Qnn. 2L, 22)1. Kar cauocrosreaslutfi BuA s cocraB poAa Acrocoel,ites HaMH He BKJrroqeH Belemnites tripartitus (Schlotheim, 1820, crp. 4g ). fiepasre Hso6paxeHrs B. tripartitus' Aa'r,"re Mnaaepov ( Miller, 1g26, crp.60, r. g, u EneuQur.1o-lg) Br{neM (Blainville,18 27, crp. g2, r.4, Qm.4)rrpunaArrexar HerroJr'oMye*seMrJrspy, xoroprrfi HeJrbss c yBepeHHocrbro oTHecrH laxe r poAy Acrocoelites. B gansHeitueM noA HasBaHfieM tripartitus pasHbIMH aBTopaMnolmcblBaJrucbcoBepueHHo p€rsJrHrrHbre 0opr"or, HanpuMep d'Op6NHsu (d'Orbigny, 1g4S, crp. 2g1, ra6n. 4;), (Dumortier, 1864- 1874, f.4, fliovop'be Sn-. 4) nprnaarexar uenortrroMy sK3eMnrurpy, (Phillips, 1865-187L, crp. 62, ra6r. 2, Sur.2gl, gnerueu (Janensch, 11":gr 1902, crp. 118, ta6r. 11, Qnr.6-8), 6eccnopHo npnnaAJrer(aruuepasHbrM BHAav. I-loeroMy BbTAeJIHTbrHn BHAa cefigac y)Ke HeBooMoxHo. B orneaunrtx c6opax Hs oAHotro cJlos Hapcay c pocrpaMr.r, HMerourr{Mr{xopoluo BbfpaxeHHble [pHBepuI[rHHbIe 6prournrte 6opos!.st, o6rrapyrcrBarorcs pocrpbl, JII{ru€Hnue 6prcuruofi 6opoencr. OrsocuTb HX K ApyroMy ellAy H poAy ocHoBaHnfi Her. Ilo-euaxvoMy B pesynbrare HHAHBI{AyanbHbIxoTrgroueHnfi npnBepurr{HHan 6prcuruan 6opoeaa Ha pocrpe Mo)Ker He pasBnBarbcs.
Mbr BKnIoIrrJru rapre pocrpbl c ,{opoluo pasantof B cocras pola Acroclites rlpgBepunggofi 6prorunofi 6opoeaoft, Ho cr(arble B cmruHol.6plorlrlroM EaIIpaBneExl{' gro ganaAuoeBponeilcrne Br{rr5r Belemnites suprapalatinus Kotb, B.unisulcatus n ABa BoBbrx ceBepocu6npcrcax BuAa: omolonensis u hedonensis. 3ru Blainniitesugbt HecoMHeHHo 11peAclsBiflsrcrr oco6yrc rpylmy, KoTopyP e 6ynyUeM 11pHIIo'lBMarepnaJra BotaMoxuo ueaecoo6pasHo BbIAeJII{Tb Jre'rrfi 6oaee fipeAcrasurerbnoro B HOBbrft poA. u n e o n p a Q n r r e c K o e p a c r t p o c r p a H e H I r e . H n x r r u f ir e r r a n n ( ? ) , Boepacr - Toap aareB suxsufi aalrex 3anaanofi Eaporrr, Bep)q{ nnuuc6axa ( ? ) nnunc6ax Ceaepuofi Cu6uPu.
25
26 27
Qnpe4eilnTeJtbHaflTafirJl;;qaBllAoB PoAa Arcrocoelites 1.
22 23 24
Pocrpcr c xoporuo paSBI{TbtMr{ [pnBepuIrHHbIM[ CmTHHO-6OI(OBbIMIIn nnn cy6tufllin.ryntrecrofi Oopt\'tbl. 6progrnof 6opoonar.,u4 cy6ronrqecrofi
28
Pq, Aeroc o e lit es Liss'aious' 1915 2 (53). g ig). 4 ( b ).
5 (8). G a? ).
? (B).
PoCrprr, CI(aTbte C 6oKOe C EB Megee lOO . . . . . . . . . . . o 3 ' o r ' 4 Poctprr, oqegb ctulbgo ygrurgegHbler fla orono 1OOO ... Pocrp cy6ronn recrol oopr*u c oBarI'EbrM nonepeqHblM ceseEreMt fla ortoao 9OO. - roaP A.grac.ilis (Hehl in Zieten, 1830, S. 28, Sabl, 22, Fig' 2) Pocrpu cy6rurnnrraprmecKol Sopr'rrr . . . . . . . - . - . . . . . . . 6 nouepeqrrbtM ceqeHl{eM, fla r orpyTtroMy c 6rgrsml{ fucrp orono 1000. - Hn)KHDrfi rrurnc6ax A.trabecufc,Lissaious, 1915, p. 10, fig.7-9 nonepeqHblM c 6oxoB otp)rmelrno-npsMoynoJrbllbtM Pocrp co cxriruu ceqeuueM, nla otrono 1OOO. A.pergrdcdlis Sachs sp. nov. - Hrrxnrrfi Toap . I o..... P o c r p r r , c l r r l b g oy n n n i e n r c r e , f l a o r o n o 6 0 0 - 8 0 0 . '1o ' ' ' ' ' ' ' ' ' ' ' ' ' ' ' ' ' Pocrpucy6xotmrecrofi0opr"{br' 11 ' ' ' ' ' ' ' ' ' ' ' ' ' ' nonepeqEoeceqeEl{epocrpaoBaltbHoe
8(L71. 1414). 10(13). 11 ( 12 ). Pocrp xopouro BbIpaxeBEOfi Cy6rOnrnrecrofi $opr.,6t. - rotlp A. stimulus (Dumortier, 1864-18? 4, v. 4, p. 38, tabl. 4, fig. 8-10) ue6oasuofi. cy6ropclecrof Bblpaxeggofi @pvu, ( caa6o 12 13 ). Pocrp
P. 63, pl. I ) A.boLetiLissaious ,Ig25, p. 60 (comb6morel,l9?1,
-
aePxnnfi ToaP - Hllrrnufi aaren. @pr.le' cpennnfi L2a(13 ). Poctp 6nngto{fi r cy6urannAprrlecxofi - BePFtrfi roap (Ig42, Taf. 161, S. 9, Fig' 5-8) A.subgrac.itis Kolb 1 3 L 4 l . flonepe.rnoe ceqeg11e pocTpa olpyrnegE9-npgMo]'ToJtbgoe. -rcap A,ilminstrensis (Phillips, 1865-18?1, p.64, pl'12, fig' 30) 15 . . . poctpu . . . . . . . . . . . . cy6rUnutrApr.Idciofi Qop*rgr L 7 l . L4 1 5 1 6 ) . flouepevHoe ceqeHue .pocTpa oBaJIbHoe, fla oKoJIo 7OO-8OO. A . s u - b t e n u d s( S i m p s o n , 1 8 5 5 , p , 2 6 , e m e n d . P h i l l i p s , 1 8 6 5 - 1 8 7 1 , p . 6 0 , pl. 10, fig. 27 ) - roap 1 6 ( 1 7 ) . flonepevnoe ceqenr{e poc"pa 6nneroe f, orp}'TtloMy. - nnxnnfi Toap A.loigiconus (Schwegler, 1969, S. 186, Abb.73) 16 . . . . .''' . . 400-600 . oxoao yAJIxHeHHbIe' fla ( 3 O ) . Pocrpu 17 L1 ' ' ' ' ..... L 8 ( 2 2 1 . P o c r p s r c y 6 r o r u r t r e c r o f i $ oo p. .u. b. .t . . 5OO-55O. oxoao fla pocTpa oBaJtbgoe' l e ( 2 o ) . flonepe,lroe ceqegre
- roap A . o x y c o n u (sH e h l i n Z i e t e n , 1 8 3 0 , 5 . 2 7 , T a f . 2 l , F i g ' 5 ) r 2 0 ( 2 L \ . llonepe.rnoe ceqegne pogTpa ca.a6OOBaJIbHoe,npn6nnxoeTgf, oKpyrnoMyr fla oroao 3 60-500. - roap' A, triscissus (Janensch, 1902,S. 113, Taf. 12, Fig.7 ) BogMoxHo Bepxn rurunc6axa 6prounas 2 L ( 2 2 1 . flonepe.rHoe cetlegge pocTpa o6pyrneHgo-TPaneq,elganbg@t cropona
t0
yxe
cmtunofi,
IIa
oroao
42O.
29
30 31 32 33
34 35
36
37 38 39 40
4L 42
I 44
45
46 47
I L tffi Drb
r(?), teB
) ) L
, . 1 6 3 ,T a f . 1 1 , F i g . l - 2 , 7 ) A . b a n z e n s i s( K o 1 b , 1 9 4 2 S
- nepxHr.rri
TOap 22 (3O). Pocrpsl cy6quauHApr{qecKoft @pruu . o . . . . . . . . . . . . ., 2.g 23 (261. llonepevHoe cerreHr{e pocrpa oBaJlbHoe o o . . . . . . . . . . . . 24 24 (25). Pocrp cpenHr{fi, fla oxono 500-600. A , q u . e n s t e d t r _ ( O p p e l 1, 8 5 6 - l B 5 B , S . 3 6 3 , Q u e n s t e d t , l 8 4 6 _ 1 8 4 9 , - rosp - HlrlKHHfiaaaen S.423, Taf. 27, Fig. 2-3) 25 (261. Pocrp rpynnufi, na oxoao 4OO. z{.trrscissiformis Kolb (L942, S. 164, Taf. ll, Fig. 3,9) - eepxHuri TOap 26 e9 ). flonepeqHoe cerre'ue pocrpa oKpynneH'o-npsMoyronbHoe . . . . 27 27 (28). Pocrp 4pynultfi, c yAnl{HenHofi npr.rBepu}rnHofi .racrrrc, BepuuHa cMerrleHa K crrnHHoft cropoHe. A , t r i s u l c o s z s ( S i m p s o n ,1 8 5 5 , p . 2 6 , e m e n d . P h i l l i p s , 1 8 6 5 - l B ? 1 , p.60, pl. l0,fig.27)- roap 28 (29 ). Pocrp ue6oasurofi, c yropo.reHHofi npuBepunHnoft vacrurc, BepurgHa SaHI{MAeT UeHTpanbHoe noJloxeHl{e. A.strictus Lissajous (I9ZT, p. 16, p\.2, fig. 2_4) _ roap 29 (3o). flonepevxoe ceqeHl{e pocrpa oKpyrnoe, cJrerra cxaroe c 6oKoe, np'BepuI{HHA-gqacTb KoporKas, BepurHHa cMeuleHa K cnHHHOfi CrOpoue, IIa oroao 4SO-SSO. A.polaris Sachs sp. nov. - Toap 30 lla oKoao 3OO-4OO . . . . . . . . . 31 !461. Poctprr caa6o yAnHHeHHbre, 3 1 ( 4 L l , . P o c r p uc y 6 x o r u . r e c rQo of ip r r r. g. .l . . . . . . . o. o..... . Sz 32F71. flonepe.moe cerreHr{epocrpa oBaJIbHO€.. . . . . . . . o . . . . . g3 33 (34). EprcruHae 6oposAa [poTsn'Baercs .qo nonoBurr.il fla. A.persulcatus (Janensch, 1902,S. l16, Taf. ll, Fig. 2) - BepxHHfi TOap 3 4 ( 3 7 1 . E p r c r u n 6a oa p o o nHa e 6 o a e eL / S f f a . . . . . . . . . . . . . . . . Bs 35 (36). Pocrp l9pynnatfi, c orHocnTeJrbHo roporxofi npxBepurHHofi qacrbro. A.subtriscr'ssus Kolb (1942, S. 160, Taf. ll, Fig. 6,8) _ eepxHuft ' Toap 3 6 ( 37 l, Pocrp cpeAnufi, c yaJrr{HeuHofi npuBepuunHofi qacruro. A"graciliformisKolb (1942,S. 16l ,Taf.T, Fig. Z, Taf. 9, Fig. 4,5) _ aepxunft Toap 3 7 ( 4 o l . flonepe.ruoe ceqerl{e pocrpa otpyrJlenHo-npgMoyfonbHoe . . . . . 3g 3 8 ( 3 9 ) . Pocrp Kpynnbrfi, c AlnHHoft 6prcuuofi 6oposnof,. A.uentralis (Phillips, l865-lBZl, p.80, pl. lZ, fig.44_45) __roap 3 e ( 4 0 ) . Pocrp cpeAHuf,, c KoporKoft 6prorunof, 6opa:aofi. A.uoltzii (Phillips, l865-lB7I, p,79, pl. l?, fig. 43) - roap 4 0 ( 4 L l . Ilonepevnoe ceqenne pocrpa orpyfaoe.
(Oppel, S. 363, Quenstedr, 1846_1g49, !.cgnoldezs .1856-1858, 5.424,Taf.27,Fig.4) - aepxuufiroap - aanen
4 L ( 4 6 ) . Pocrpst cy6unannapnqecKof, Oopr.,|bro . . . . . . . . . . . . . . . .42
42 (+sl. 4:! ( 461. 44 (451.
45 (46).
flonepe'rnoe ceqenxe pocrpa oBaJrbnoeo A.subspinaefornis Kolb (1942, S. 162, Taf. 5, Fig. 25 _27) nnfi roap flonepe.rHoe ceqenre pocrpa orpyraoe. Pocrp cnennnf c yruinHeHuofi npnBepur{Hnof, .racrsro.
_ Bepx_
A- iuaenis (werner,L9r2,s. 136, Quenstedr, I846-1949, s. 420f, Taf. 26, Fig. 12, 13) _ roap
Pocrp se6onsuofi, c oruoclrTeabno roporxofi npnBepurgunof,qacrbro. A.dillbersansisKolb (1942,S. l6t, Tuf. 5, f at. ZZ_ZL) - eepxHnfi Toap
4 6 ( s 3 ) . Pocrpsr KopoTK{e. cra6o cy6rorrrec[ile, 4 7 ( 5 L 1 . I-lonepevnoe ceqeHre pocrpa oBaabnoe, HOe noJloxeHue
lla rrenee goo . . . o 4 7 BepullHa saH[Maer UeHTpanE-
48
tl
48
(491.
Pocrp
A . t r a n , o y e n s i[s, i s s aj o u s ( 1 9 2 ? ,p . 1 , t rp, l . l , f i g . 1 , 1 - 1 5) nausc6ax
49 (5O). 50 (51).
SL
cpeAHufi, cnJIbHo c)Kar c 6oxoe'
(52
- nepxHufi
Pocrp cpeAHI'Ifi,caa6o cxar c 6oroe' - BePxHI{fi roaP A . s o l i d . u sL i s s a j o u s ( 1 9 2 7 , p . 2 0 , p l ' 2 , f i g ' 1 3 - l + ) Pocrp venxnft' cHrIbHo cx(ar c 6oros' - eePxHnfi 15-16) A . s p t n a e f o r n t i sL i s s a j o u s ( 1 9 2 7 , p ' 2 7 , p l ' 2 , f i g '
oKpyrneHgo-npsMoyronbHoe' ). flonepeuHoe ceqeHl{e pocrpa cMeureHa K c[IdHHofi c'roPoue. 9' l-1) A,jonenschi (Ernst , Ig2)\, S. 85, Taf' 12, I'ig' lvloxHo Bep)g4 naugc6axa
BepuI'IHa -
roap'
Bo3-
52 ( 53 ). flonepevHoe ceqeHr.repocrpa 6a'sxoe K oKpyrnoMy. 16) - eepxHnfi A.ttrniacensis Lissajous (1927, p' 15, pl' l, fig' roap tr Poc'rpot, cxarble B c[I'IHHo-6ptotuHov Ha[paBJIeHHH, EE ot 1OO 53. 6OAee. . . . . . . . . . . .
' ' | ' ' o ' ' ' ' ' t ' t ' '54 55 .''''''''''''
5 4 ( 5 e ) . P o c r p e t y A J I I , I H e H H fbl Iae o, x o a o 5 O O - 7 O O 5 s ( s 6 ) . Pocrp caa6o cy6xoHu'recrofi Sopuot.
A . u n i s u l c a t u s( B l a i n v i l l e I g 2 7 , p . B l , p l ' 5 , f i g ' Sopvst. cy6uuanHApnqecKoft ( 5 e ) . Poc'rptt 56
21) - r'oap
C yAnHHeHHOftnpNBepulIHHOtiqacruto, 6es 3aMeTHorO CnI{HHo6prcurgoro c>xarug. S. 164, Taf. 7, Fig' 4, LL ) - eepxA . s u p r a p a / a t r ' r r uKs o l b ( t g + 2 ,
5 ? ( 5 8 ) . Pocrp
58 (se). 59.
Huri roap. Pocrp c yKopoqeHuoft npllBepullUHoft qacTblo' CI{JIbHo c)KaTblfi e cnuHHo-SproruHoM HanpaBJIeHHll. A. hedonensis Sachs sp. nov. - roaP - aaJr€H (l-la oKorlo 25OJ50). Poctp caa6o yAnHHeHHbrfi - BePxHI{ftnnuHc6ax - ToaP nov. A.omolonensls Sachs sp. A c r o c o e l i t e s p e r g r a c i l i sS a c h s , s P . n o u . l Ta6n. 1, Sur.
l.-3
N 87-1, Myeeft Hfl CO AH CCCP, Hoaocu6upcx. Pera lleestfi f onornn Keao$ snxnuft Toap' sona Dactylioceras commune' Pocrp rpynnstf, oqenb cnnbHo yAII]IHeHHbrfi(tla ot(oro IOOO), cyG AnarHo3. qxnuHAp6qecroaft, c>xarsrfi c 6oxoa ( EE oroao 8O-9O ), c orpyrJreBHo-npsMoyronbXopouro BbIHbIM rronepeqHblM ceqeHueM. BepuruHa cMeulena K cmtHgofi cropone. 6ptounaR. u cnngHo-6oxoerte 6opoener paxegbt npnBepuugHble B geu Hrre npr{sHa Ku. flpunepruuHHagqacrb pocrpa BbITsHyra, cocraB.nser (ra6n. 1). flpuaepurunnan 6proruoKoJro Ll3 fla, nepundHrrfi yfon paBeH 2U24o 'racrn, 6opcaaa nay6oxaR, npnBepul{HHoft ecefi BAoab uaa 6opoeAa flporsr.r{Baercs s6rl{gH BepuHHbI AOBOAbHOrunpoKag, no Mepe yAa[eHI{s oT BepuIHHbI nepexoAl{T qacrl{. CmrHHo-SoKoBbIe B ynJlouegr{e, coxpanrrlouteecg BnJIorb Ao aJlbBeoJlspHofi
I,J
Anuui o6uiar
flnxnt r{acTu
[uavt Bepu) Aualr y Bep Paas: [nnut Auau npI{BC [nav TUHHHI
Ynoa Ho-61 Yroa 6ptou
j
Annr o6ut Amr tIacT
Atral Bepu
Auat YB€ Paq Anu, Aua npul Aua III}TH
Yno
Ta6nuua
1
I4svepeHurr pocrpoB A c r o c o e l i t e s p e r g r a c i l i s S a c h s . s p . n o v . Ne 87-2, P. Jlesbrft Kenos
llapaverprr
]ap
(
y.ait
AnuHa,l npeanoaaraeMas o6uraq lycrauoeneHHarr flauHa nocJreaJrbaeonspHofi qacTr{ Jluar'aer.p crnaHno-6pnurHofi y BepruuHbr aJrbBeoJIbI ,0,uar,terp 6oxosofi y
)4 i5
Fx-
BepUJI{Ebt
197,O(t-256) 193,O(]-229) 163,1(1039 )(83 ) L 5 , 7 ( 1 O O) 12,1(77)
L25,4( L740l L23,6(1124 ) 106,1( gos ) (as ) 1 1 ,O( 1 0 0 ) 9,2 (83)
AJIIrBeOnbr
Paruryc 6prournofi [nnna npnBepuunuoft qacrn Jlr,rar'aerp cnr{Huo-6proruHoft B npr{BepuuHgoft qacru Jlnar'aerp 6orosofi B npHBepruuHHorlqacru Yron aJrbBeoJrspHsrft s c[r{HHo-6pnruHofi urocKocrN, rpaA. Yroa Bepur{HHbrfi e cnuHxo6prcrunoft nJrocxocrr{, npaA,
6 ,9 ( 4 3 ) 54,9(35O) l s , o ( 9 7 ) ( 1 O O) 1 2 ,8 ( 8 2 ) (8 5 )
2T
41,6(378 )
10,5(e5 x 1OO) 9,7(88)(gZ)
20
lH-
Oxonqaune ra6nuusr
87-4, Kennr"rnp
2 1 O , O( r g e Z ) 1 8 2 , 0( r r e e ) 160,0(rZOS)(ZO)
Annsa Inp"rronanaeMarr o6qa.g Jy.m"o"".HxaJr Arnura no-cneargeeonrpuofi qacTu g yG' \&-IbH-
9? DI'II-
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y
6oroaofi Auaverp y Bepulil{br aJrbBeolbr Paaryc 6prcuruofi ,flrmna rrpun€pturuuroft .racrn fluarvrerp cnusso-SprourHoft a fipuBeprulranoft qactu fiuar',rerp 6oroeoft B rrpuBepuururoft llacrl{ Yroa aJrbBeojrspnsrfi s crrtrgHo-6prournofi nnocxocru, npaA. Vnon Beptulrrxbrfi e cnuxso6pnrunoft rrJlocKocru, npaA.
L22,stioo+) L22;5(1oo4 )(-)
1 5 , 2 (1 O O )
L2,2 ( 100 )
1 4 , O( 9 2 )
1 3 , O( 1 0 7 )
5,4(36) 65,0(4281 14 6( e6 ) (1oo)
4t4(361 44,O (3 61 ) 12,5(102I(1OO)
141(e3xe7)
L2,4(102 )(eg)
ZL
24
Mas' cJla6o HaolHyTa Ir npn6nnxena r 6prcurHof, cropone (p[c. 1). Ha Ha.rarbnbrx cra'ql|sx pasBl'Tns qpnn trlaMeTpe 41 5 rrrv pocrpbt r{Meror caa6o *{paxeggyrc cy6r(oHr{lrecKyto Qopr'ry rrptl sHaqeHnsx IIa oroao 640. IIpn nxarrreTpax nopsAra g rr,rv pocrp npno6peraeT yxe cy6urn[H.q)I*IecKyro Oopr,ry, snaqenxs na aoepacraror Ao 900-960. fipn AaasrreftureM pocre )Kr{BoTHorc Ao cmr'Hedprcrunoro A[aMerpa pocrpa L2-L6 r'o"r nafrrpaaer\cs yBenrtrenue fla no looo n 6onee H rrosanense
t3
grac i li s Sachs Puc. 1. flpononbHoe ceqenl{e pocrpa A croco e lit e s Per commune, eras P.IIesp. nov. I$ 87-,.3, nuxnnfi Toap, sona Dactylioc sbrfi Keaog (XO,7)
A
IITIJIHI ceqet xo{<
B cocTt rutfi
B Sopve pocrpa - €fo cy)Keuue aSJMeu
HeKoropoft seperexoBuanocru aepruniu alrl'BeonbI. H g lvreH q u Bo cr r. Meggeccs orHocurenbHas AnuHa npl{BepuunHofi l,lrer 6prcunroft 6opoeau (or 1/5 so 2/5 fla), cteneHb ee pacuupegus. 6oxoa (or c cxaroro pocrpoB MelrnrBbrM sBJIseTcs nonepequoe ceqeg1ae ao rroqrll oB€lJrbxono). Ocxoauble [apaMerpbl MeHscy6nptvoltronbgono roTcs cpaBnl{renbtlo Mano (na a ilpelenax 9OO-126C_, BE.77-92' HecronbKo BblAense.ncs pocTp c p.Kenuvsp(lS 87-4ls 66-85-97). (EE-107)' nepegnefi tIacTR cxatstfi B ctT[Hgo-6pnruuor'r HanpaBneHul{ cToJIb o6nagaroulux BlrAoB, C paBH egu e. Hs 3a[aggoeaponefiCxlrx (Hehl in Zieten ) A.gracilis pocTpoM, nefiacoaslfi xe BbrTcrryTbrM oTnlrqaetcn cy6xoHmecKgfi $opVofi DoCTDdr paHgenn6llc6axcruft A.trabecula Lissaious orryrcrBneM 6orosoro c)l(arns pocrpa. Effrsox no HI{xHer9 Toapa pxnofi qac"Tl{ oopr*re, Ho eute 6onee yAnuHeH pocTp tl3 g'rT,4t2, Opf, uso6paxe$Hsrft KgeHurreAToM (Quenstedt,l 846-1 849, gracilis. acuarius Ta6n. 25, 6nr. 4) rroA HasBaHueM Belemnites trlo BFyTpeHHee cTpoeHue er-o HeI|SBeCTHO'He [cKnloqeHo' flocxontry PocrpH ApyrHx 6nugxux BIIAOB oH o,THocuTcs K Solpingoteuthis. (Sinpson), /. ilminstrensis Aaocoelites [/. lsubstlicili"" Kolb, A. trisulcos,s (Phillips), A. suhtenuds(Simpson), d. longiconus (Schwegler), A' triqatti,tus , 184'6-184g, crp.42O, ta6n. 26, Qnr. 17)l ve;rffiirt; ({u"*t"At, no t}opHan6onee cxqxufi Hee yAnnHeHbI, IIMepT fla oxono 600-800. '205, A.bb. 86) (schwegler,,1969, s. Me pocTp A.trtpartitus gracilis (fla orono 8OO). rrs HuxHer.o Toapa OPI tarxe Merlee BbnslryT pagee onpeB xonnerUusx I{3 Ceaepnofi Cn6upu o[ucblBaeMrtfi euq I'lOlb. A.suhgracilrs xarc HanbHceBofi rr T.H. Aensrcs B.H. CarcoM o a u u a n b H a s n p u y p o q e H H o c T s . P o c v r p b lc o 6 p a x b l B f l p € I { l v f f , OvononurccrBeHxo tJrrrHrrcrbrx ocaAKax BAamr or rcpcKoro MarepEl(a'B flAru16tfi oTcyTCTByer C.osepuleHgo Otp11era. ctCOMpafiOxe 11B BllsoBbsx Ana6apo-Xa tf BllnoftcKoM a MerrxoBoaHbr,x oallltfl( BUA B llprl6perctbtx qTO XUBOTtIbIe Beml cao6ob TaHncKoM pafiouax. Monaro npeAnonoxl{Tb, Mope' HonnaBatounft o6pas )Kll3uu. B orKpbIToM p a c n p o c T p a H e H u e . H n x H n f iT o a p v reoFpa+HtIecKoe BospaCr
KOpo' 6oxot p03Al
BbD(Q Moxl
Cnns pocr. B ArrHll paAH nofi
rHyT pbl I1
350 AOCT
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sona Dactylioceras commune Ceaepnofi Cn6npn. 6 pocrpoB rrs ooubr Dactylioceras coffiInun€ 6acceftHa Koauust Marepuan (6accefiH p. Keanr"rrp (?) Ha roapa pocrp ns HHxHero l c6oprr A-C.[anrca, Onenera) - c6opst T.H. I(npnsofi.
A c r o c o e l i t e ss t t b t e n u i(sS i n , p s o en m e n d . P h i l l i p s ) Ta6a. ll,
Qnr. 1-3
s ub te n u i s :Si m p s o n , 1 8 5 5 , P' 2 6, 1884' p' t33' P hi l l i ps' B elem nit es 1865-1871, p. 60, pl. 10, fig. 27, 1875, p. 263, pl- 27, fig' 6' Acrocoelites subgracilis: Caxc, HaauxReoa,1967r Ta6I. 1, $nr. 4. hlesoteuthisaff. subgracilis: lGrnacoa,1968, crp. 133, Ta6I' 54, Onn' 1' HeusBecrex.I-lepnoenao6paxeHneBnaa AaHo:Phillips, 1865-187 L, f oaorxn 10, ra6r. $nr. 27. Toap AHrnnn, Caatentg e6flHgu Bur6u' 14
Puc
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lips rlac'
Poc min
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A n a r H o 3. Pocrp cpeAHI{% c}rabHo yAJrr{HeHHbrfi( I-la or(orro 7oo-goo ,, cy6qHnl{HApHuecKrft, c c)I(arbIM c 6oros (EB oxoao 75-80) onanunuM [onepeqHbrM ceqeHneM.. Bepurrna cMerqeHa K crrr{HHofi cropoHe. npuBepuulrnbre 6oposAbr crrr{H_ Ho.6oKoB6Ie n 6prcrunan He BglxoAsr sa npeAeJrbr npnBepullHHoft qacrr{. Bneul Hue ''pI{s HaKI{. flpueeprun'Has qacrb pocrpa yN'epe'Ho B'ITsHyTa, cocTaB,[ser oKoJro 3/8 nocneaJrbBeoJrspHofi qacrH. BepuuEa saocrpeHa, BepunH_ narf, yroa HaxoAr.rrcq B npeAerax Zt-iSo (ra6a.-21.' np;;p*;HHbre 6opoeaor KopoTt(xe, 6prcuHae 6opoeaa Bblpaxena JrJrque, uupoKas n nenay6oxas, annn"* 6orosbre 6opoeau qerKo Bblpaxenbl Jrr{rrrb y HeMHonux or(seMrrJrspoB. Epnrunar 6opq3lla no Mepe yAaneH[Is or Bepun[Ibl nepexoAr,rr, B y[nouleHr{e, Koropoe Toxe He BbD(QAItr sa [peAenbt rlpuBepurnnrofi trac.Tr{. Eoroarre cropoHbr ynJrorrlegbt, Ha Hrrx MoxHo na6nplatu ABe napaJureJlbHble noJrocbl, Tepgpu[ecg B [pr{BepugltHofi qacrr{. cnsgnag n 6prauHas cropoHbr BbrnyrJrbre. rlonepeqHoe cetreHre e eaanefi qacrr{ pocrpa Menee c)r€To c 6oron (EE_Zb_gg, 66_91_91). BnyrpeHHHe n p H s H a K I { u o H T o n e H e s . A r b s e o J r as a H H M a e r o x o n o Lf4 AJII{HbIpocrpat Bepu}IHa ee Snaq[TeJIbHo cMeIIIeHa r 6prourHofi cropoHe - 6prorurrofi paal{yc MeHsercs B npeAeaax 33-40% CE. AauaeoarpHrrfi ynoa B crrr{Hgo-6prcrunof, naocKocrl{ cocraBJtg er 22o. oceeag rrulris_ rroqTx npqMoruHefiHa, cJIerKa r{96 rHyra u npu6nxxeHa r 6prourHof, ctopone (pnc. 2r. Ha -Ha.ranbgbrx p*rpbl HMeIor orqeTJIHByIo cy6rorur'recKylo Qopr"ry, rpn AxaMerpe 4 uu "r"on"* fla oroao 3501 3areM pocrp cTaHoBHTCC60nee ynrr"""oiv (npn o".r"rp.7,5 ruru fla AocrHnaer 550-600) u nocreneHno npxo6peTaer cy6rurauHApuqecKyn (npn @pvy Ar{aMerpe oKoJro lO ruta r lla oroJro 650 ). Hel'{enqr{Bocrb. Hvercunecs B KoJrneKu[r,pocrpbr a o6uev qo_ BonbHo oaHcyrr{nHbr,Menrrrorcs sHaqeHuc fla (e npeaenax SSO_8OO), EE (or ZS ao gB), crenenb BbrrsHyrocru r{ saoqrpeHHocrl,r npHBep_ uuunoft qaqftI. Pagnl{qHa creneHb pasBr{Tr.rs npnBepurrHxsrx 60posa, eqrb pocrpbl, y KoropbD( npr{ HaJrrrqrm qerrofi 6plounoft 6opooau.c[ungo6oxoaste 6opoeasr He pasBurbt l{nu pasBnrbr oqenb cna6o. Ecrr pocrpbr, y Koropbx 6prcunan 6oposaa cBqqrrres x se6onsuofi nox6une EerocpeA_ crBeHHO y BepuflrHbr poqrpa. Bcrpe.rarc/Tcs pocrpbr c umpoKoft u xenrryr_ 6oxofi n nao6opor c y3Kofi 6prcurHofi 6oposaoft. flpu gToM Hepelro pocrpbr c pasnfiqHofi creuenu BbrpaxeHHocrbro npHBepur{Hxsx 60posa o6xapyxr{Barorcs coBMecrHo B oAHoM o6pasqe.
Il,fymlb rblft [&)b
E71,
Pttc. 2. flpotoauuoe cdqeHne pocrpa A c r o c o e l i t e . s s u b t enais (Simpson) It 85-12, HuxHnfi Toap, p. Buarofi(x o , g )
c pa B H e H rr e. onucbrBaeMbre pocrpbr HanoM[Haror pocrp A.subtenurs (simpson) I{3 Toapa AHnanu, 6oaee noapo6xo on".."orf, n nso6paxeHsrrf, @naanncoM (phillips' 1865-1871). Anrarf,crnf, pocrp nMeer MeHee yarruHeHHyroqpuBepunH'yro qacrb H HecKonuro npn6rurxapueeca r cy6npcMo]rronbuoMy nonepe{roe ceqesue. Poctpu ApyrHx 6nrsKrx sanaaHoeaponef,cror* r"alr [A. suigraci/is Kolb , A. ilminstrensts (Phillips), d. longiconu" tS.trwegler)l ,rp, otnHa*oBbrx s'ar{eunsx fla o"Jrnqarorcn cy6xoxn.recxof, Sopr.,roft. I-lpeanapnreJrr,Ho B.H. CarcoM H T.H. HaarHsesoft pocrpbl A' subtenuis us Ceeepnon cx6npx onpeAeJrsJrrcb rar A.subgraci/is Kolb u A.aff. subgracilis Kolb. floa ;; n o c J r e a H n MH a s B a H H e MA . s u b t e _ nuis onllcaH E B.fI. Knxacoaorv (196g) ne roapa 6accefiHa Konsrv.cr.
f5
Ta6nnua
2
(Simpson) Heuepenus PocrPoB A c r o c o e l i t e s s u b t e n z i s flapar.aerprt
Jlnuna o6ua.s
InpenuoaaraeMatr f v"t."o"tteHnas
Ns 87-5' p. Ileaatfi Kegon 1 1 0 , O( 1 0 2 8 ) 1 O 7 , 9 ( 1 O O 8)
IS 87-6, p. flenrtfi Keaon 1 O 3 ,O @ t z l 1 O O , 6 (9 4 9 )
.[nu rra nocJreaJlbeeonnPHoft tIacTu Jlnar.,rerp cnnnno-6Prcrunofi Y BepurI{HbI aJIbBeoJIbI 6oroeofi Y BePIUI{HbI llnaverp a.nbBeoJIbI PaAnyc 6prorurrofi Aauna npuBepunnnofi qacrl{ cmasno5Prorunoft a lluayetp trpuBeplunnnoft qacru
8 3 , 1 ( 7 7 7 x Z O)
7 7 , 7 ( 7 3 3 ) ( Z S)
10,7( 1OO)
10,6(1OO)
6oxoeoft B rPI{BePfluaverp unnnofi qacru Vroa aJIbBeoJIgPrDtfi e cnutlHo6prourttoft nJrocKocrl{' nPaA. Vron Bepul{Hnbtfi e cnunHo6prcunofi nJrocKocrl.Ir rPaA.
8,3(78)(83)
Oronqaulle
Beltr Bele
foac crp. 62, Ana: vecxHfi, il
oKp)'I'J
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8,5( 79 ) 3,9(36) 3 3 , I ( 3 O 9) 10,O(94) (1OO)
8,9 (83 )
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BHHbI
3,5(33 ) 32,O(gOZ ) 9 , 9 ( e 3 3 ) ( 1 O O)
cropoHel i'sKHe H '''x HHofi :'I;.KEHHII
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35
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91ro (zoo ) 83ro (og g X- )
1 O 2 , O ( g g O) 89,2( 7 63 ) 7s,L(671\3 2l
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crll{HHo-6prournoft y
BepurI{HbI
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Yron
13rO ( f OO )
1 0 , 9 ( 1 O O)
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8,2(75)
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5,2(40l 35,o(269) L2',!(93)(1OO)
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22
24
nocneA t;igrmi H)rc
H l t x < n u ft io a p ( s o pacnpocrpaHeHne' rt reorpaQuuecKoe Boapact (sona Collina mucronata) ua llu.po""ru" falcifer n errrue)n Hr{sbl Bepxueno roapa Ceeepnofi Cu6upu; ToaP Anrrun. L 5 p o c r p o B l I 3 H l I x H e I ' o H HusoB Bepxnero ToaPa 6accefixa KoMarepnaJl. JrbIM.bI- c6opot A.C. Aanuca, 7 poclpoB I{3 HllxHero roapa 6accefiga Bnaps c6optt T.H. HaasHceeofi u T.V.ltrpunofi. 16
rI[
P x c., i tson )
2 4 9
A c r o c o e l i t e .s t r i s u l c o s u s ( S i m p s o n ) Ta6n. II,
$nr. 4, ra6a. III,
Sur. 1-3
B e l e n n i t e s l r i s u l c o s z . sS i m p s o n , 1 8 5 5 , p . 2 6 1865-1871, p. 62, pl.11, B e l e m n i t e st r i p a r t i t u s P h i l [ i p s ,
fig. 28
H e r { s B e c r e } Lf l e p n o e u e o 6 p a x e n u e B H , q aA a H o : P h i l l i p s , 1 8 6 5 - 1 8 7 I , loaorun crp. 62, ra6fl. 2, Qur. 28. Toap AHnaun, Caarnux, e6ausu Bur6u. (tla oKorro 500-650), cy6urauuApl{Pocrp rpynrrsnt, yAJrHHeHHrrft EnarHos. .iecxufi, cxamrfi c 6oxon (EE oxoao 80-95 ), c BbrrrrHyror? npneepuruuHofi qacrblo ;{ oKpytJleHHo-[pnMoynoJrbtrblM noneperrHblM cerreHr.reM. Xopouro BbrpaxeHbl npt{BeprHHHbIe 6opo.aalr - 6prcurHas H cnuHHo-6oroeste. BHeunue n p u 3 H a K H . l l p n n e p u r n H H a tqr a c r b p o c r p a s a H r . r M a e ro K o J I o[ o J I o 3HHbt [ocJreaJrbBeorrrpHoftvacrn, BepuHHa ocrpal, cr,rJlbHocMeueHa r cnngHofi (ra6a. 3). I-lprnepuruHnble 6opoenst BepruxHHono ynna 27-350 cropoHe, Benr.rqr.rHa nonoBLIHbI npxBep6pnurHaa 6opoeaa npoTsrr{BaercrlAo n r.rs Hr.rx nay6orcle, i'3Kr.re :nngoft qacrl{, Aanee Ha6apaaercg ynJrouleHr.re,Koropoe flpocnexuBaercg Ha npoilorrru scefi nocJleanbBeoJrspHoft .racrl,t pocrpa. CnuHHo-6oKoBsIe 6opoealt "rr)KeHfir.r Kopoqe, Ho y cavofi BepurnHbl pocrpa Bpe3aHbI rny6xe 6pnurnofi 6opoellt. Eoro3bre cropoHbl ynno[IeHbr, Ha Hr{x HHonAa oTMeqarorcs caa6o BbIpax(eHHbIe napHble :toJrocbl, Tepgrcuuecrr B [pHBepuunnofi qacrr{ pocTpa. cnugHag cropoHa caa6o ert:r)'Knas, 6prcuruae oaMerl{o yflnolueua. Cnrnrsofi rpafi cna6o l{3orHyr B rrpxBep:irnsuoft qacTld, 6prounrofi l{goruyr suaqrrreJrbuo c}rnbuee. I-lonepe$roe ceqeul{e no aceft Anuue poc'Tpa coxpaurrer or(pyrneuxo-[psMoynonbuoe otrepraul{e. C:renenu 5oxoeolo c)Karus 6onsue y Bepur{Hbr arrbBeorrbr (EE 81-87, 66 84-98). BnyrpeHHr{e npr.rouaKH rr oHToreueg. A n b e e o n ag a l r u M a s r oKorro Or25-Or3 Anr{Hbr pocrpa, Bepur{Ha ee cMeueHa K 6prcumofi cTopoH€r 6prcunrofi pa[Hyc y Bepur{Hbr aJrbBeolrbrcocTaB,nser 1/3 CB. Ans-l Beonspublfi yrun paBeu 23o. Ocesag Jrrrur,rnnpu6nuxesa K 6pnuruofi oropoue, oqeub cna6o r{goruyr.a (puc. 3 ). Ha Hat{aJrbHbrxcraAr{rD( pocrf,br HMeIsr cy6xouuvecKylo $opuy, 6onee saocrpeHbr (nepuruunrrfi yron npu AuaMerpe pocTpa 615 MM paBeu 2Oo, flpn Ar{aMeTpe 15 vv 28o, rrpr{ AuaMerpe 21 r,av - 32o). Oruocurenbuas yAnr{Heuuocrb MM ila (npu guaMerpe 6rb c BospacroM Boopacraer oocrpa 634). npr truaMeTpe 15 r"n'r - 59O, rpl{ Ar{aMerpe 21 583, H sN,teH qI{ Bocrs. OcuoaHbre napaMerpbl pocTpoB A.trisulcosus noABepxeHbr rone6annsM, sHatreunrr fla s npelena:( or 4OO ao 7OO, e rcHoauoft Macce or 5OO Ao 650, EE e rpeAenalc 75-95, B ocHoBHoM cn 80 Ao 92. B paeuofi cTeneHH Bbrpaxenbr rrpr{Bepuruubre 6opoagtl, cnnnro-6oKoBble n 6prorunaq. Ecrs e[EHurrubre posnpbr, co6parnrbre coBMeqrrlo c pocrpaMl{ Tr{nulruoto Anrr AaHHoro Br{Aa o6nnra, Ho nuueuubre npnBepurlruoft 6promrofi 6opoaabr. y qflHorp r{s Tax}rx pocrpoB (Ne 8710) ua6npaaetcrr cnI{HHan 6opoeaa, He Aoxqflqas Ao Bepunrbr, o9eBr{AHO BTOpUqUOTOflpOHCXO)r(AeHHs. CpaBHeHue. Pocrprr,4.trisulcosus no ceoefi MaccuBuocru u cy6uI{JII{IIApI{trecrofi tlopr"re pesKo ornxqarsrcrr or Apyfux BrdroB Acrocoelites.. cpaaueuue c 6nueKHM Br{aov.A.polarjs sp. nov.a6119 npH orrucaHr{u nocneAuero. IloegueroapcKue pocrpat A.quenstedtt (Oppel) n .d.triscissit'ormis (Kolb) 6onee MeJTKI{e,6onee KoporKl{e, r{MeKvr 6onee yKopoqeuHyro npuBepurr{Huyro tracTb u 6onee yArrr{Hexrr}no 6prounryro 6opo.aay.
I
(sr r ta )
\ \
KoPuc. 3. I-lpoAoJrbnoeceqeHlre pocrpa Acrocoelites trisulcosus (Srmpson) Ne 87-11, roap, p. Keauvsp (xO,8)
2 4 9
\
I I
!r V l7
Boe;
Ta6nuqa 3
HH3bI
s t t l ( ' o s l r s( S i m p s o n ) HsuepeHusl PocrPoB A c r o c o e l i t e s t r i Ne 87-9, p. .Ileettfi Kelos
Anuna fnpeanoaanaeMas o6uras I VcraHoaJlenHatl lanna nocJleanbneoanPHofi qacTIr fluaruerp cnuHso-6Prculuofi Y BepurHHbl aflbBeoJrbl 6oroaofi Y BePutlHbI fiuavetp
1 5 6 ,O( 9 7 5 ) L 5 2 , 1 (9 5 6 )
11e,O ft++)(ZO)
87-10, Keanvap 1 8 O , O( g r + ) 1 5 5 , O( 7 8 7 ) 130,5( 6 ffi )( 73 )
1 6 , O (1 O O)
1 9 , 7 ( 1 O O)
1 3 , 9 (8 6 )
16,7(85)
5,8(36) 56,2(351) L 4 , 3 ( 8 9 ) ( 1 O O)
76,5( 338 ) 19,6(1OO)(1OO)
AJIbBEOJIbI
PaAuyc 6prcrunofi qacr}t Ilnnna npl{BePurrnnoft Jluar.,rerp cnunuo-6PmuHofi o npl{Bepuunnoft qacrn Jlnar.aerp 6oxoaofi B nPHBePtuHHHofi .Iactu Yroa anbBeorcPneifi B crII{HHo6prounoft nJrocKocrl{r trPa!. Yron Bepul{Hnbtfi e cnt{xHo6pnuuofi nJlocKocru, rPaA. Oronqanue
L ,O(8S) (98 )
16,4(84)(84)
Ansua IrR.att-araeMas o6uras lvctanonneHHas .[,mma nocneanbaeonnPuofi qgcTu Auar.,retp cnusHo-6prcrunofi Y
28
N 87-11, p. KeauuRp
N e 87-12' p. Bunnft
1 8 4 , 0 ( A Z O) L 7 2 , 7 ( 8 O 3) 133,2(6,3+lftZl
1 3 4 , O( e O S ) 1 2 3 , 0 ( Z g S ,s ) .
e 6 , o( o r g l ( t z l
L7,O(Ar)
1 3 . , 5 ( g Z)
PaalrXc 6puuuofi Anuua npuBepluuHuofi qacru Auaruerp cmrnso-6Pnurxofi a npr{Bepuunsofi qacru
7 , 7( 3 7 1 5 2 , OQ + Z l 1 9 ,1 ( 8 4 ) ( 1 o o)
6,1(39) 51,O(329) L 4 , 2 ( 9 2 ) ( 1 O O)
nrar'asrp 6oxosofi B nPuBePurItHnofi qactn Vrpn aJrbBeonspuufi B crlxllHo6prcunofi nnocKosrn' rPaA. Vron Bepurl{Hxbrfi s cnhrno6prcuxofi flnocKosrl{, rpaa.
L'7,2(8O)(9s)
L2,8(83 )(go )
Auavetp
6oxosofi Y BePUI{HbI
aJIlrBeOnbl
crr B np BaeTcg
oT
23 32
nyKIIag. qeM cT( r{acTH
Btt ) J.ill{Hbl I TaBnge'l 5pronrHo pbl HMe
xeHbl. I :xaMeTl pHqecK:
Hsw l.po lart !{brM (E l'€TpbI ile nopl
Hofi ssn qacTHOC x nepe)i
lia npcn (odp. }r
27
E o n b u r a sq a c r b p o c r p o B c o 6 p a n a B n p e nprrypoqeHHocrb. eaunaarrHas B Oprononcrou [MyIlecrBeHHo rJII{HrlcrbIx ocaArax BAaJIu oT lopcKoro Mareprrxa penXo (no g pafioge. Orens OneHeKcKoM n aMMOHnTaMn c paftOne CoBMeCTHO pocTpbl A.tricpaBgepr{ro c MaccoBbrM pasBHTHeM mytrHx sunos) BcTpeqalo1cc ."Lrro"r" u Ana6apcroBnnpficxoro ocaArax B MeJrKoBoalrbtx necqauo-rJmnucrblx IS{3HU H ro pafiouos. fIo-EtHA[MOMy, XJ{BOTHbIeBetr[ CSO6OAUOnnasaput{fi o6pae peAxo noAtuIbIBalI{ K 6ePerau. l8
BHe
AJIbBEOJ
Soxoerte
15,5(1OO)
anlrB@nbl
paxeHbt
:rpno6pe
2 L , O (1 O O)
BeplIJUHbI
qacTblo
pcsAa
ta6nuuut
flapaverpu
I o.ir xnxuttfi .[sa .recxufi,
xoae
28
BE'
Mar n ( . o ll i n 15 pocr c6oprt T Toapa II(
cpt enponei
s rssi/t HsrM cl
?nc. 4 : o r ' . N e{
I P olari
Bospacr u reonpa0HqecKoe p a c r r p o c r p a H e H t l € . H u x n u f tr o a p _ HH3bI BepxHetro'Toapa (eoHa Collina mucronata) Ceeepuofi Cr.r6r.rpu,Toap AHrauu. M arepuan. 30 pocrpoB Hs soH Harpocerasfalcifer, Dactylioceras commune x (.ollina mucronata no peKaM lleerlft Kenon n Mysyryaxax - c6ophr A.C.Laruca, 15 pocrpoB Hs roapa p. Kenuusp H 15 pocrpoB r{s roapa 6accefina p. Buarofi c6oprt T.H. KuptlHofi n T.H. HaruHneeofi, 3 pocrpa ns Bepxgefi .racru HHxHeno Toapa no6epexue AHa6apcxono saJrr.rBa- c6optr B.H.Caxia u T.V.Hansngesoft.
1 7 3)
A c r < t c o e l i t e sp o l a r j s S a c h s , . p . n o u l Ta6n. IV,
iloo) r4)
) i2)
)o) ))
B npF
d"r.
L-4
I o.ir o t u n Ne 8 7-1 4, Myeeft Hff CO AH CC.CP, Hoeocn6np"rq Pexa Map:
IOM 10 irtlpcKoIHII
I{
Puc. 4. rlporoabHoe ceqegre pocrpa Acrocoelites polari.s sachs, sp. nov.Ns 87-L5, unxnnf, roap, soua Harpoceras falcifer, p. Mapxa(x ci.a) I Polaris -nonapusrfi.
19
Ta6an ua 4
'sp' nov' I4errepenus pocrpoB A c r o c o e l i t e s p o l a r i s S a c h s '
N e 8 7 -1 4 ' P. MaPxa Jlnuna fnPennoaaFaeMas o6uas f Ycr"no-tenHas 'racru Ilnnrra nocJleaJlbBeonspHofi !,uar"rerp cnuuno-6Prcrunofi Y
1 2 9 , O( Z S g ) 114,8(675 ) 88,7( 522 x 69 ) 1 7 , O ( 1 O O)
Bepu[HbI aJIbEeoJIbl 6oroeofi Y BePIUI{HII
lluar'retp
N e 87-15, P. Mapxa
-aI
1 O 9 , O( O A r) 9 8 , 2 (6 1 4 ) 7 g , L ( 4 5 7x o z ) 16,0(r.oo)
16,3(gO)
1 4 , O( 8 8 )
2 s , A 1 7 s)
L 4 ,2 ( A g ) ( 1OO)
2 3 , 5 (L 4 7 l 1 3 , O ( 8 1 )( 1 O O)
13,O(ZO)(gz)
L 2 , 2 ( 76 l @ 4 1
AJlbBEOJIbI
Paanyc 6PtotuHofi Aanna rrpl{Bepulnttttoft qacru Auar'retp cmrnno-6Pnurnoft e npxaepuilHuofi qacru Jlnar'aetp 6oroeofi B nPI{BePtuHHHOfi qacru Yron aJII'BeoJIsPttltfi B cnunHo6prcuruofi lurocrocrnr rPa!. Vnqn BepuII{HHbtfi s cnHHEo6ptotunoft nJlocKocrl{r nPaA.
23 Q
46
yanugeggocrbp ero' He cu6upctclx KoBbrM cxarueM pocrpa n 6orourefi ornocuremHofi (Simpson), HecKoJIbKo6onee yAJrutrisulcosus A. pocTpbl BuIOB cxolqrbl no fupve qacTblo' Pocrpu A'trisBbrTslryrl.g&npueepurUruofi 6onee C rreHgbre, 6onee MaccHBIIble, (J"n"n""h) ;;r;;" ocmrqaxlrcs 6orree yxopoqe'uofi cy6xonuqecrofi fupuoft' E o a b u a s q a c T b p o c T p o a c o 6 p a H an QaUuaJrbHas npuypoqeHHocrs. e BHnIoficKoM lr Ana6apO-Xatanrcrov oCaAKax necqaHo-rn[HgcTblx MeJTKOBoAHbIx OCaAKaX pafiorra:L EnnBu.$rbte pocTpbr BcTpeqeHbl B npelrMylltecTBeHHo nJIHHI{CTbIX xItBoTHbIe pafione. flo-auguvouy' B4ann oT lopg11oro MaTepl{xa B OvoaOUcrou rpu6pexBeJlr{ cso6oAHonnaBaroutuft o6pae )ro{sHrr, Ho trpeArloqlaTaJrHMenKoBoAnrte o6ctaHogxr{. nsle p a c n p o c T p a H e H l ' I e . B e p x H s nq a c T b u reofpaouqecKoe Boepacr (aoHa Dactylioceras commune u sstue)' Cu6upu HuxHeI'o Toapa Ceneprroft Toapa 6accefiHa p. Bnnrce - c6opst Hlr)KHero pocrpoB r{3 12 MarepHaJr. 5 pocrpoB Hs HuxHero Toapa 6accefina KoartT.H. HaauHgeeoft u T.V.Itlpunoft, 14' pocrp H3 Hu)KHeFo roapa IrcBa YprcHr-TyMyc 1 MbI - c6optt A.C.Aaruca, T'H'Hansugeaofi' c6opsr p. KenrvsP pocrp c 1 A c r o c o el i t e s t r i s c i s s u s ( J a n e n s c h ) Ta6a. V, $ur.
icrt; : La|
l"-3
:=a-l-'
183O, S. 31, Taf' 24,?'ig' 3' B e l e m n i t e sf r i s u / c a t u s : H a r t m a n ni n Z i e t e n , B e l e m n i t e st r i s c r s s u s : J a n e n s c h , 1 9 O 2 , S . 1 1 3 , T a f ' 1 2 , F i g ' 7 ' 1932, cTp. L2, ta6a. 1, Our. 4, 5, Mesoteuthis triscrssa : I(ptlvroauq 2. 40, 1947, crp. 2O1, Ta6r. $ur. Acrocoelitestriscissus: Kolb, 1942, S- 157, Taf' 9, Fig' 5, 6 1, 2 Mesoteuthis oxycona: I(psrvroasU, 1953, crp. 87, ra6l. 13, Qxr. 5 4 ,Qxr'2' t a 6 l ' 1 3 3 , c t p . 1 9 6 8 , I ( u n a c o o , M e s o t e u t h i sc f . t r i s c i s s i / o r m i s :
_ /"n-fl
,-'
: . q - r t t
5-"
OnucaH flsenurev ( Janensch, 19O2, IoaoTHn. Hnrl roap BoctoqHofi QPaHurn.
20
Ta6r. 12, Qnr. 7).
Bepx-
l
Ne 87- L6, p. Kenuvsp
),lb87-17, P. Ilenstfi KeAon
1 O 9 , 2 ( 5 8 7| 109,2(587 ) 72,3(389xOO) 18,6(1OO)
-)
B O , O( O A Z) 75,O(625 ) 5 4 , O ( + S O) ( O e ) L 2 , O( 1 0 0 )
L 7 , O( g r )
l _ o ,o (8 3 )
21,3(115) 1 5 , O (8 1 ) ( 1 O O)
2 2 , 2 (r e s ) 1 1 ,O ( g Z l ( 1 0 0 )
9 , 7 (s 1 ) ( e a)
L4,3(ttl (gs)
46
FCrsrx :Tt|-
{. lris-
IJ\,{
rax e Fexb
r-5l-
n
tofi,
42
Pocrp cpeAnuft yAanHeHrrrft lluarHos. c yMepeHHo yIlnHHeHHoft npraepuuHHoft
Lr2,O(t qsl 9 6 , 0 ( O + O) 81,3(S+Zx 73 ) 15,O(rOO) 1 3 , O( A Z )
4,5(38)
o)
87-18, Buarorl
5,9(39) 26,6(L77,
1 3 , 5 (e o ) ( 1 o o)
L2,6(84)(93)
40
(fla orono 3OO-5OO), cy6roHr.recqacrbro H Beptur{Hofi, saHrvarcuteft
5aneroe
K uenTpanbHoMy noJroxeHr{e. flonepeqHoe cerreHue oKpyrnoe, cJrerKa c}K&c 6oxoe ( EE onono 86-94 ). npr,reepuuHHr,re 6opoegu qerKne, Ho KoporKlre. BueuHue npusHaKH. lpueepurr.rHHas qacrb 3aHuMaer oKoJlo L/2 nocJreqacrlt pocrpa atlbBeoflqpxofi H npu ero o6uteft cy6xoHn.recxofi Qopve HeorqernnBo oTAeJIeHa oT cpeAHefi qacru pocrpa. BepuruHa gaocrpeHa, 3aHr.rMaer ueHTpaJrbHoe roe
noJloxenue HJIH HecKoJrbKo-cMeUeHa K crrr{Hucift ctopoue. yron HaxoBepruusuufi (ra6a. 5 ). flpueepurnHnaa 6prouuarr 6opoeaa ray6oB npeAeJlax 3 4-370 txrcg Kas H ysKan' nporcfuBaercs no 1/ 3 nocJreaJrbBeorrspHoft vacru, AaJree BbrrroJraxsBaeTcrr H nepexoAHT B ynJroueHr.re, KoTopoe npocne)KHBaeTcc Ao Bepunlrbr aJrb_ BeoJIbt. CnuHHo-6oKoBbIe 6oposrut Bblpa)Kelrbr caa6ee H rlpocne)K[Barorcs ue 6oaee qeM Ha 1/5 qacru. nocaeaJlbBeoJlspHoft Eoxosue croponbr Hecyr napHbre noJrocbr, caa6o ynJlouet{bl. Cnnggag cropoHa BbrnyKJras. Qopua nonepeqHono ceqeHurr oKpyrnaflr cJrerKa c)Karag c 6oxon, ocraerctr 6oaee unu Meuee ndcrogHuofi no nceft Aluue pocrpa. CteneHs 6oxosoFo c)Karr{a y 6oasurnHcrBa pocrpoB cor(pauaercrr B npr.rBeprunHHofi nx qacrn. BHyrpeHHHe npHgHaKH n oHToneHes. A n b a e o J r ac o c r a B J r q e r o K o n o 3O-357o AJII{}IbI pocrpa, BepuHHa ee cnel'Ka npu6anxena K 6prourHoft cropoHe, 6piourHoft paAI,Iyc y BepuuHbr aJrbBeoJr.br paBeH 29% CB. Afluseoacpnrrfi yron B cnuHuo-6proruHori
2
paBeH 24o. nJlocKocrx Oceeag rrllug. norrrll npsMas, cJlenxa npu6auxeHa K 6prourHofi cropoHe (prc. 5 ). lOncre pocrpbr HMepr 6oaee npaBaJrbHyIo, qeM y B3pocJIbIx xIlBoTHbIx cy6xoHnvecKyto Qopvy, 6oaee 3aocrpeHbr (nepurnHHsrfi ynoJr npH AHaMeTpe pocrpa Zr5 vv paBeH L|o, npr.r Ar{aMerpe 20 vv 370),
Ho no cTeneHlt
orHoclrreauuoft
y[nHHeHHocrH
He orJrr.rqarorcrr or
B3pocJlbrx
oco6efi.
- 2 I-
HeveHqHBocru. ao 52O, qaue
MeHqrorcg 3HarreHua l-la, Bapbnpyroune B npeAeJrax or Bceno Ha6aroAaprcq [a nopaaKa oT 4OO no SOO.3naqeHHq EE oTMeqarorcrr B npeAenax or 76 ao g8, y 6onuuuHcrBa pocrpoB HaXo,asrcg Me)ruly 86 H 94. He ocraercrr nocrosHHofi $opr,la H creneHb Bbrpaxerrrro_ crN 6prcurnoi.i 6opo3A.br - y HeKoTopblx pocrpoB (l.fs g7-2 t) 6opoeqa 6rrcrpo rjr,r360
2l
(Ja5. flponoauuoe ceqeHr{e pocTpa Acrocoelites triscissus commune, soHa Dactylioceras Toap, Hrxgufi nensch) Ne 87-20, p. Mapxa (x O'8) puc.
!p) neH suI
86IT cBe
nonaxr{BaeTcs u pacuilpttsTcs' y Apynux ocTaeTcs yexOft n nny6O' qacrrrrpoc:rpa (]t87-22). Kofi Ha npgrrsxenul{ noqTlr Bcefi npnBepur,rtruoft OmacblBaeMbrepocrpbl He oTnl{qarorcs HuqeM CyCpaBHegue. llosAHe|,roapcKoro tIlec:rEeHHbIM oT pocTpOB sanalHoeBponeficKoro Acrocoelitestriscissus(Janensch),KoropblenuulbHecroJlbKo6onee OAnaxo n Cn6rpu Bcrpeqalorcs ygopotreubl (fla orono 35O-4OO). poqTpbr c nOAo6HbIMI{sgaqe6usMu fla, a qAg.1H@TI{ TaKOB P@TP' otrncanrrbrfi B.II.KunacoBbrM noa rrasBarlreM Mesote.uthiscf. triscissif ormis Kolb, cY!8 no BceM npusnaKaM rtp]IuaAnexautril K Bury triscdssus. flueuur ( Janensch, 19O2) rrpearloxnn HasEanne triscissus Bsa! uen 8 elemnites tlisulcatus Zieten - Mnarlu€ro crrrrolrl,i,iv.aBelemnites ('Zieien Salpingoteuthi s ) tri sulc atus Dlainville ( 1 827 l. Pocrp B.tri wlc atus 6onee ygnr{Herr, qeM y nonorruna triscissus (fla orono 43O) gToMy nplrsHal(y 6Onsrue npu6nuxaeTcs r cn6npcKl{M pocTpaM. tl nO pagBIrTOfi 6prcuuofi 6opoeqrt He ocTaBnee,t CoMHexopoluo Haruque Hr{s B ToMr tITo BI{A triscdssus otuocl{Tcs K poAy Acrocoelites. B cunonuMr{Ky gTofo Br4Aa Mbr BKJuoqHnI{TaKxe KaBKaocKruc nooAHeu MesoteToapcKroc .d.rriscissus |.fl. KpslMnonb[a ( 1932, t9471 ns roapa p. Bnnrofi. He srnocl{Tc9 K uthis oxycona |.fl. KpurMnorrbua (1953) t(Hyry6ugge, 1966, cTp. 161, ra6n' 38, Qrn. 8) AaHgortryBr{ry pocTp lri.trirrissa r{s Bepxgero roapa Ceoepuoro Kanraea, mrueulrblfi rrprBepunrmofi 6pnumot 6oPostrbl;
Ta6rrua
Op 3TX DET qaH F6t
r€p 6pe BAK
rGf 13 roI'
c60,
xrl p.)
5
HevepeHnnpocTpoB A c r o c o e l i t e s t r i s c i s s u s ( J a n e n s c h )
ls g 7-2 1,
llapaurerPrt
LnuHa fnp"nnot anaeMa.s o6uac lYctunoutteHHas Annna nocneanbBooirspHofi qacrl| ,tlnar'aetp crnanno-6prcuHof, y BePuIITHbIaJlb-
p. MaPxa
t& 87-22, P. MaPxa
111,otslo I 9 2 ,O (+ SS )
8 7, L (5 8 1 ) 75,9( 5 06 )
98,O (OOS) 90,?(560) 67,4(41 6) ( 69 )
2 O , 2 ( 1 O O)
1 5 , O( r O O )
1 6 , 2 ( 1 O O)
19,8( 98 )
1 3 , 9 (9 2 )
L 4 , L (8 7 )
5,8(29) 4 9 ,4 (2 1 5 )
33,3(222)
45,1( 278}
d
cnnnHo-6Plourlluaverp Hofi e npunepruuHHoft qacrl{ 6oxoeoft e lnavetp rIpI{Bepu}rHHofi qacrn Yron anbBeoJrspnstfi a cmanHo-6prcuruofi ruIocKocrH, rpaA. Yroa neprunnuufi e cnnsHo-6prorunofi nJI@Kocrll, npa!.
22
)
(rb
BEOJIbI
6oxoaof, Y fluaverp Bepuilnbl aJrbBeoJrbl Pamyc 6proruxofi Annna npunepruuHHofi qacTlr
t - -
t ' l l
1 6 , s (8 2 x l O O) L 2 , 2 ( 8 1 ) ( r O o )
14,2( 88 x 1OO)
1 s , 8 (7 8 ) (e s )
L4,O (86 x 99 )
L 2,O.(80)( 98 )
24
troJ roEr I 60( m;x nrl ryrI !rE [IE
ca) rtl :rTtlf l e r
37
35
34
t O,3 69u
;ars
Xoporuo BHpaxennas cy6KoHltqecKas $opva pocrpa oTrt.rqaer l.triscissrrs oT ApyTr|x Br{AoB Cr{6Hpcrxx Acrocoelites, cxoA}rbtx c onncbtBaeMbtM Br{aoM no cTe_ nelrH yAJrr{HeHHOCTU pOCTpa. Hg sanaAHoeBponeficKr{x Br{AoB 1aUArot K ;l.lrisci.s_ sus: A- banzensis (llolb.) u A. gracilif ormis Kolb, orJruqarournecs6dnsrunrra6oroBbIM c)KarileM pocrpoB, A.subtriscissus Kolb., pocrp KoToporo npn B3rJUrAe cBepxy cy6ulann[pmecKofi $opr'arr, A. persulcatus (Jinenscli) u A. cortoideus (Oppel), I{Merorrrue pocrpbr c 6oaee ArurHHofi 6prourHofi 6oposaoft. Hacronbxo Bce srr{ Bl{IlbI caMocTogreJlbHbrr Benbgg peuraTb 6es nepecMoTpa AocraroqHo rIF€ACTaBr?terrbltbDr l(oJrneruufi pocrpoB Ka)t(loro uo Hru(, Koropbrx B HarrreM pacnopgxeHun Her. p o c r p s t c o 6 p a n s l B M e J r K o B o A H bnr e oaunaJrbHas 'npuypoqeHHocrb. xcqaHo-rnHHr{cTbrx ocaAKax e BuarcftcKoM H AHa6apcrorra paftona>r; eAnH}rqHbr€pocT_ pct HafiAelrbl B 6acceftHe Ouonona B FJrltHltcrbrx ocaAxax BAanr{ or ropcKoro MaTepuKa. Moxgo npeArlonoxuTb, qro )Kr{BoTHbreflpeAno.tnraltu MeJIKoBoAHbIenpn6pexrrue o6cranoexn, xoTs, cyAs no BbrrsnyTocrr{ pocrpoB, 6unu cso6oAgorura-
t.
BAIOUIIIMI{.
22l,.
lo
t
ts
, )) X.
rc-
Boepacr u neoFpaQu.recKoe pacnpocrpaHeHrre. Hnxnnftroap (eoror Ilarpoceras falcifer u Dactylioceras commune) Ceeepuoft Cu6upn. onuH pocrp H3 Bepxos nanHc6axa (?). B 3anaasof, Enpone r{ Ha Kasxase roap, BosMoxHo, TorrbKo aepxHnft. MarepnaJr. 15 pocrpoB I{s Hr{xHero roapa 6accefina Bnapg H p. /lnnAe c6oprt T.H. HaauHgeeoft n T.H. KirpuHofi, 2 pocrpa us Hnxnero roapa H8 p. JIeaarfi Keqon - c6opu A.C..0,anxca, 1 pocrp rls Bepxog rurnnc6axa ( ? ) s 6accefiHe p. Ynxsi - c6opu 3.B. Ocnnonofi.
'c-
A croco e lit e s janenschi ( Ernst)
B) Bbt;
Ta6a. II,
Qnr. 5-7
B e l e m n i t e s t r i p a r t i t u s o x y c o n u s : Q u e n s t e d t , 1 8 4 9 , S . 225,T af. 26, ?-ig2L. B e l e m n i t e s t r i p a r t i t z s : J a n e n s c h , 1 9 0 2 , S . 1 1 g , T a f . 11' Fi g. 6-8. B e l e m n i t e s ( O d o n t o b e l u s )j a n e n s c h : E r n s t , IgZ4, S . 85, Taf. 12, Fi g.9,11. Acrocoelites rostriformis var. janenschi: Kolb, L 9 4 2 , S. 160, Taf. 10, F ig,5, 6. Mesoteuthis cf. lrl:scissc: Tyrxon, 1954, crp. L2d.. Mesoteuthis janenschi: Bopoueu, 1962, crp. 1oo, ra6a s2, $ur. 1, z. Mesoteuthis cf. subpyr'anidalis:.Krnacos, 1g68, crp. lg2, Ta6n. SS, QnnB. JIerroTr{u, esr6pannarfi B.H.CaKcoM: Ernst, L924, Ta6r. 12, $nr. 9.Bepx'Opanttrlu. xxfi roap CeaepeBocroqHofi Pocry cpeanufi, caa6o cy6xorn.recrorfi, oTnocureJrbno xoporrorfi IluanHos. (ila oroao 23O-32O1, cxatsrfi c 6oros (EE or SOuo gO), c otpyrflenno.-rpflMo)'noJIbHbIM no[eperrHblM CetreHl{€r"r. flpnsepuHHHble 6opCaAU 6prournan n cnunnO-6OroBble KopoTKle, Ho qeTxl{e.
))
Bueurnne npnsHaKr{. O6uac Oopr"rapocrpa Mensercg B rpeAeJlax oT cJra6o cy6ronnqecxofi no cy6rrnnnnApr{qecKofi. flpnaepurugHas qacrb san[Maer 6oaee noJIoBI{HbIfla' aepurllna cMeuleHa x cnnnnofi cropone, caa6o oaocrpega, Bq)ungrorf, ynon HaxoAI{Tcs B npeAeJrax 47-55" (ra6a. 6). Y aepunrcr HaquHarcrrcs npHBepufirHrrbre 6opoanu, 6prouHan n cnnnno-6oroBbre, He saxo1sune Aanee noJrb BHHbI llplrBepunnnoft rracrn pocrpa. Ep nrnaa 6opaaaa ysras u nay6oras y BeptuHHbI, 6eicrpo paculpsercs u BblnoJraxuBaercs, Ha npoAonxenun ee na6npnaeTb cs ynnoueHrre, npocrexqBaeMoe Ao arbBeonrpnofi qacTr,r p@Tpa. CnuHBac cropo_ lla Bbt[yxJras, 6oxossre saMerno yrutouenbr. flonepetnoe ceqeHue oxp]rnenuo-npsMo)ToJrbHoe, cxa?oe c 6oros. Crenegs 6orosono cr€Tr{s no Ar![He pocrpa saMerHo He
MeHgeTCg.
BHyrpeHHne npngHaxH u oHToneHes. AnbseoflasauilMasr oKoJIo O'35-Or4 lannu pocrpar BepuII{Ha ee snaq[TeJrbuo cMeulena r 6prournofi croponq 6ptotttttofi paAfiyc y BepurI{HbI aJrbBeoJrbl cocraBnser oroJro 1/B crnrngs-6proruuoro
23
t?
Ta6nuua 6 I4svepeHus pocrpoB
; l r - r
Ns87-24, p. Mapxa
flapaverPlt
[nr,r na fnpeanonanaeMas eHHas o6ruaq ly"tu"ottt AnuHa nocneaJlbBeo.urpHofi qacrll cnusHo-6p]oulfluaverp Hofi Y BePlxHHbI aJIb-
6oroeoft Y
BepullHbI
arlbBeoJr.bl
PaAHyc lnuHa
6prourHofi npueePuruHHofi
qACTH
Anavetp cnusHo-6prcuHofi B nPuoePurnHHofi
N 87-25, p. Mapxa
Ne 87-26, p. Ih.rHAe
6 5 , O ( S O Z)
73,O@tt) +8,7(406) 63,O(+OO) 44,O(284)(OOl 4t,O(342 )
2 L , 2 ( r O O)
1 5 , 5 ( 1 O O)
t +,ols igl
BEOJIbI
fluave'rp
ci
13,8(89)
L2,2( Z g )
B
H
.1 o
e1
6 H
1 2 , O (1 O O) 1 O , 3( 8 6 )
p p H JI
7,3(34') 3 5 , O( 1 6 5 )
4,I(261 2 5 , 3( 1 6 3 )
1 9 , O( r S e )
, (57)(rOo) L 8 , 4 ( 8 7 ) ( r o o )1 s , o ( g z ) ( r o o ) 1 o 4
fl
n
x H
qACTH
finaxaerp 6oroeofi e rTpr,rBepuuHHOft qacrH Ynon aJlbBeoJlnpuuft e c rnru Ho-6prcur Hofi IrJIocKocrn, Ynoa
17, 6( 83 )( 95 )
L L , 2 ( 7 2 ) ( Z S)
9 , O ( 7 5) ( 8 7 )
n lt f
29
FpaA. BepuI{HHbIft
B
48
DD
47
cnLIHHo-6prcuHofi nJIOCKOCTII, npaA.
Oceeag Ar{aMerpa. Aflr,eeoaapHsrft yroJr B cnnHno-6prcr:rHofi trnocKocru paB€H ?g'. rrtr1vtscaa6o r{sorHyra, npu6auxeHa K 6prournoft cropone (puc. 6). Ha Ha'{aabnblx 6oaee saocrpecraAnsx pocrpbr r{Meror 6oaee npaBunbHyro cy6xoHllqecKylo Softy, paBeu 22o, lrpu pocTpa vv AI'IaMeTpe 5 yrcn (eepurnunstft rrpll ALIaMeTpe Hbr 1l MM - 32o, npg AuaMerpe 21 r"ta - 460), no creneHl,I oTHocI{TenLHoftywtuHeHHocrr{ He oTJrgrrarorcrror B3pocnrrx (sHaqeHue lla rrpn AnaMerpe 5 vv - 32O, npn ALIaMerpe 21 r"rv - 3O7 ). 3 n a q e u u s f l a B p o c r p a x A a H H o r o B L I A ax o a e 6 n p t c R c n a 6 o , HsveHrruBocrs. B npeAenax or 22O ao 35O. Hecronsxo 6oasure MeHalorcs snarreHun BE. B KOJIneKupru ecrrr pocTpbr, cunr,Ho cxarble c 6oroa, c EE MeHee 8Or t't ecTb pocTpbl c EE orono 90. CpaBHeHue. Onucrlsaevrlft BI{A us roapa 3ana[uofi Enponur cuaqaJra SurlrpupoBan noA HasBaHr{euBelentnites triparlllus (Quenstedt, 1849, Janensch, 19O2), earev Spuctov (Ernst, 1924) tora 6rrn eurAeneu B Hosutfi e;ua janenschi, Kona6 ( Kolb, 1942) janenschi paccMarpuBar KaK Bapnerer Belemnites rostriforzzis QuenHenbsl cornacnTucn, nocKonbC eruwr nono)KeHr.reM stedt (1849). Ky pocTpat janenschi uvepr npr.rBepurHHylo6prcruuyr,o6opoeny H oTHocgrcs K poAy ,4r'rot'oelites, pocrpbl rct.striformis (r.aaaauruficrlHonuv
c
;
;
I P H c . 6 . f l p o a o n s u o e c e r { e H r n ep o c r p a A c r o c o e l i t e s j a n e n s c h i ( E r n s t ) d ]$ 87-24, Huxnufi roap, p.Mapxa ( x O,75)
24
Relentnites brettirostris dbrbigny, 18 42) se uMeror 6prcurHofi 6opoearr r{ oTHocnrcq K poAy lvlesoteutlis. He roapa Cu6upu A. janettschi 6rta onuca:a V.V.Tyrxonlrrur (1954) non HooBaHreM Ilesoteuthis cf. lrisr-issa u H.C.Boponeu (1962). Cu6upcpre pocrpbr HHqeM cyuecrBeHnblM or ganaAHoeBponeficrux He oTrurraroTcs. Or apyrnx Br{AoB ^lcrocoelites olTldcblBaevstfieHA AoBoJrbHocI{JtbHo oTJr}rtraercs ac}rMMeTpurrgofi 6auexoft x cy6rulallHApuqecroft @pvofi pocrpa n ero yKopotrenuocrbro. A.tramayensis (Lissajous, L92'7, crp. L4, ra6n 1, Sur.4, 15) ue Bepxnero rurrrHC6 a x a Q p a H u l t l {H A . s o l i d u s ( L i s s a j o u s , 1 9 2 7 , c r p . 2 0 , r a 6 a . 2 , Q u r . 1 3 , 1 4 ) H3 HHX(He|O Toapa cDpaHrUrra HMeror pocrpbl, cronb )Ke KopoTKHe, Ho c UeHTpanbHbIM
nOJIOXeHI,TeM
BepluHHbI.
QaullanbHan npldypoqeHHocrb. B c e p o c r p b l n H a u r e f iK o J r n e K u r {cuo 6 paHbl B MeJIKoBoAHbIxra npn6pe)KH.blxnecqaHo-rJrr{Hr{cTblx ocaArax Buarcficxofo parloHa. Yrasstearorca A.ianenschi ne Ana6apeXaranrcKoro n Oxorcrorg paftoHoe. CoefepuenHo oTcyrcrByer A.janenschi a $auuax oTtpbrrono Mop:r s OuonoHcKoM u Oaegercxorra pafioHaL Moxo{o Ayr"rarc, qro xuBort{ble xgJlr{ a npn6pexHofi soHer nPeA[oJroxs.ITeJIbHoB npruoHnbrx o6craHoexax. B rroJrbgy nocneAlreno r'oBopHT Maaag BbITCHyTOCTbpOCTpOB. Bospacr pacnpocrpaEeHue. u reot.paSnvecKoe B e p x uu n r g c 6 a xa (?) - uuxHnfi roap (no sonrr Dactylioceras commune BKJrrorrnTenbgo) Ceaepnofi Cn6upu, Toap (oco6eHrro eepxHufi) 3anatHofi Eaponu. MarepnaJl. J-5 pocrpoB r.rs BepxoB nnurrc6axa (?) - Hr{xnero roapa 6accefiHa peK Bnarcs, CuHefi n llnnAe - c6opsr - T.H.HanbHseBoft, T.V7.If,;r1pr{Hofttr H.l.foas6pafixa.
o)
A c r o c o e l i t e s k e d o n a n s i sS a c h s , s p . n o v . I Ta6a. l,
Qnr. +6
Gastrobelus sp. Caxc u HaauuReaa, 1967-2, fonotun MgPr
sag liblx rpetTl-
'J20, r5o, LCJIt
H
I l:-
4) 'i 'nf c.1bCrfL,}l'HM
..,t)
J
Ne 87-28.
Myeefi Hff
ta6. g, $nr. 8.
CO AH CCCP, Hoaocn6npcx. Pexa Keru-
Toapo
.Enaf H03. Poctp cpeanufi, currr,Ho yAJrr{Hennbrfi(IIa oKono 5OO-75O), cy6IlHnHHApl{.lecKIrfinan caa6o BepereHoeuAEbrfi, crraruft s cnuHHo-6ppuHoM Ea[paBJIeHI{H' c BbIpa)KeHHbIMI{B pagnr'rqnofi cteneul{ npr{Bepur{HHbtMu 6pnurnofi n cmrHHo-.6oxoesrvu 6opoeAaMr,r. B ne ur rr u e n p H g H a K lr. llpueeprunHHag .tacrb gaocrpeHa, cocraBJrger orotto L/ UL/ 7 nocneanbBeonspnoft .racru pocrpa. Bepurnna pacnonaraercg s 6Jtuerol,r K ueuTpanbHoMy noJIo)KeHHu,Bepuurrnsrfi ynon HaxoAurcs B rrpeAenax 29-360 (ta6n. 71. y BepuHHbI Haqnuarorcrr cnlrHHo-6oroesre 6oposAlt, nenay6oxne, Ho npoCnex[Baloquecs Ha [poTg)KeHItH noqTl,t aCefi npnBeprugHggfi 'raCr[ pOCTpa. Eprorunar 6opoeaa Herny6oran, 6rtcrpo nepexoqr{T B )mnouenue, Koropoe rsnercn BAoJIb ecefi nocJleaJrbBeonspgofi qacrn pocrpa. 6nssgo-SoKoBbre 6opcarct 6oaee nny6orcae lr IJII{HHbIe H AoxoAsr Ao cepeAlnbl [ocJreaJrbBeon*pnofi qacrr{. Cmlnna.s cropoHa cJIerKa BblrryKnas, 6oxoarte cropoHbl yfinouenat, 6oxoBble noJtocbl eADa saMerH-bl. flonepevHoe ceqeHlde olpyrneHHo-TpaneqneBrrAlroe, cr(aroe B cmrHHo6prorurrov HanpagneHru. CreneHb c)Karng 6oasue B nepetnefi qacrrr pocrpa (snatleHue EE 1O4-L26), y6sreaer B rpnBepurnnsofi ero qacru (66 94-106 ). BuyrpeHHI{e lrpI{gHaKr{ 14 oHTofeHeg. A r b a e o n a n p g M a . s ,o a H r { M a e r oKoJto Or25-Or3 AaI,IHsI pocrpa, BepuJr{HacMeulena x 6prorunofi cropone, aJrbBOoaaputtfi yroJr oKoao 3Oo, 6prournoft pa[xyc cocraB.[ser 30-45% CE. Ocesac nuHvs, cnerKa HsotrHyra n npr6an)t(ena r 6prournofi cropoue. Ha EaqanbHbrX craAHr.x pocrp l{Meer cna6o cy6ronu.recxyro Qopuy n xaparT€pr{syeTcs enaqennsMu fla, 6ausrc{vll K BspocJlbru oco6sr,a (puc. 71. HeveHquBocru. CyuecrBeHHo roae6ruorcs sHaqeHug fla (s [peaeaax oT 5OO ao 75O ), BHeurnsrr Qopr,ra pocrpa or cy6ruuruHAprtrecxofi H cJrerKa Bepereno* HageaHue Br.raa AaHo no p.Keaon.
25
Ta6anqa
7
Hevepeuus
pocrpoB A c r o c o e l i t e s
hedonensis Sachs sp. nov.
Ne 87-28, P. KennvsP Aauna fnPennonanaeMas o6utag IycranoanenHas Anuna [ocJteaJlbaeolnP uofi qacrlr llnar'aerp cnnruro-6prournofi y BepuI{HbI aJlbBeoJIbI Jluar.aerp 6oxosofi y BepuruHbt
73,6(88? ) 60,O n 231
]t 87-29, p. Kennuap
64,0 (eS e ) 54,8(731)
8 ,3 ( 1 OO)
7,5(1OO)
9 , 1 (11 0 )
7 , 8 ( 1 O 4) _
AJIbBEOJIbI
Paruryc 6prorunofi Anusa ttpl{Bepuuunon qacr}r cnnnno-.6Prcunofi a llnavetp npr{Bepunnnofi qactn Anar'aerp 6oxoeofi B nPI{BePunnnofi rlacrt{ Yron arbBeoJlsPnrtfi B cnI{EHo6p nrnofi rulocrocrnr rPAA; Ynoa Bepuruunbtfi s cnnnno6puunofl ruIocrocrur nPaA.
-
32,0(386 ) 1 0 , 5 ( L 2 7 ) ( r O O)
9 , e (1 r g ) (e a )
2,3(31) 2 2 , 5 ( 3 O O) 8 , 1 ( 1 O 8 ) ( 1 O O) 8,6( 1ls x 106 ) 30
29
36
,snAgofi Ao cnerKa cy6xounrrecxofi, crerleHb cmnnro{oKoBono cxaruc (EE 1O4-126)Hr.aerqunficg uarepuaJl He nosBoJlger ycraHoBnTb Kaxrre nn6o oco6eggocrn B t{3r,aenrubocrg pocrpoB no ux reolonnqecroMy lrnx reoFpaQuuecroMy pacnpocrpaneHrlo. A.hedonensis npnnaAJre)rcrrK o6oco6leHxofi npynne BnAoB CpaBHeHHe. KoTopaq pesKo oTJluqaeTcs oT 6oauuuHcTBa Apynux BHAoB poIA Acrocoelites, BI{AA sTOfO pOAA CrUrnge6OKoBblM C;€THeM pOCTpa. Or OnuCUBaeMOrO Hnrxe orHosHaql{TeJlssofi ornuqaercs A.omolonensis sp. nov. paccMarpnsaeMbrfi EnA 'r 3OO-4OO (sHa.renng coorBercrBenHo fla pocrpa ynaunexHocrbro cgTeJrbgofi A. supralt alati BIlrbI sananHoenponeficxue' sanrrrorcfl 6nus11rvn Son"" ).. SOO-ZSO 6oasuefi 3aocrpeHHocrt,lo orJrr{qapuuficg or A,hedonensis rrus (Kolb, Lg42l, cnl{HHo-6ptotuqacru saMerHoro or'cyrcrBl{eM I{ lr BblTsgyrocrbro npuBepurnHttofi poctpoMt HMeloult{tll c (Blainville, ) L827 A.ttnisulcatus n pocTpa, Boro cxaTl{s cna6o cy6rouu'recKylo QoPt"tY. E o n s u a q q a c T b p o c T p o ac o 6 p a u a OaUnanbHan npuypoqeHHocrs. B npeurvryI[ecraexuo rnlrHucrbrx Sauugx orKpblTorp Mops BAanI{ or topcKorg Marepr{Ka g O)@rcxo-OvonoxcKoM paftoue coBMecrHo c aMMonnraMu. OlaeJrbHble pocrpbt uaftaentt raKxe B fnl{Huctsx Sauucx orKpblToro Mops B gilgoBbcx Onenera u Ha npaeoSepe)Kbe llexu. B npu6pexHblx Mem(oBor pafionax AaHHblft sua nrx $auucx e Bunpftcxov u Ana6apo-Xarallncxov ( nafines nurrrb oallH poc-Tp e 6acceftHe noyru coBepueuno orcyrcrByer Bunros). Cneryet ryMaTb, qTo xlrB6rTgbte xunu B oTKpbIToM Mope n o6pae xrSull. Benr! cgo6oggolulaBalouufi BoepacT u 1.eorpaQuqecKoe pacilpocTpaHeHue. Bepxnen qac:rb HI{)KHerp T'oapa (goga Dactyliocetas commune h estue) - BePxnufi aanen Cenepnofi Cn6npu.
ct
p.
HE
Ke cy BE
tIIP Hbi
pa 3a
HO
Cr CJI
np cfl
KA
AN
o, Pnc. ?. flponoJrbHoeceqexue pocrpa Acrocoelites kedonensis Sachs, sp. nov. Ne 85-21, Toap, p. Ileatrfi KegoH ( x O,85)
26
i-
N g7-3 1, p. KenHMsp
Ne 85-2 L, p. Ilesbtfi Kenon
87-3O, Toxlp-lOpax
87,Oftltl 85,5(76Ll-
se,o6zt)(aa)
7 8 , O( 7 8 O) 6 8 , O( O e O) 57,O(57OxZS)
7 3 , O( 6 6 4 ) 7 2 , O ( O S S)
L L , 2 ( 1 O O)
10,O(rOO)
1 l - ,o ( 1 o o )
L4,L( 126 )
11,O(rrO)
1 1 , 8 ( 1 O 7)
3,4(341 29,9 (IZAZ )(S r ) 1 1 , 8 ( 1 O s ) ( r O O)
3 O , 6 (3 0 6 )
5,O(45 ) 32,O(Zgr)
r 2 , 2 (1 O 9x r O S)
9,o(goXroo)
e,s(es)(roo)
72,o(rogXroo) L z , O( r O S) ( r O O)
30 35
6). 0.
M atepuaJl . 15 pocrpoB I{s BepxoB HHxHeFo roapa-Hu)KHeno aaJIeHaOxorpafioxa - c6opst A.C..[anuca, 3 pocrpa us roapa-aaJreHa Ha cro-OvoaoHcKoto p. KeaulaRp, 1 pocrp Hs roapa HB p. Ilemrcxe (npanarft npHToK fleusr), J- pocrp Ho HHxHero roapa p. Mapxu - c6opsl T.H. I{nprHofi. A c r c . t c oI e i t e s < t n t o l o r t e r t sS i sa c h s ,
"p.
nou.1
Ta6a. \', Qnr'. 4-8 loaorun Ne 87-34, Myeeft l4lf CO AH CCCP, Hoaocu6upcr. Pexa lleeHfi KeAou, nuxHr,rftToap, ooHa Zug
Hasea'r.re BlrAa AaHo no p.or"ronon.
27
Ta 6auua I
Pu -rp.
HeruepeHun pocrpoB A c r o c o e l i t e s o m o l o n e n s i s S a c h s , s p . n o v .
Ne 87-32, p. Keauvnp
87*33, EYoP-3E*"t pa },{H
64,O( 3721 6 1 , O ( 3 5 5) 42,4(247r(661
62,0 @tOl 4 ' 7, 5 ( 3 6 0 )
flnar"rerp cnunno-6prcurnoftY
L 7 , 2 ( 1 O O)
1 3 , 2 ( 1 O O)
Beprur{Hbl aJlbBeoflbl Jlnar"retp 6oroeofi y BepuI{HbI
16,5(gO)
14,O(106 )
23,6( r37) 14,5(e+)(rOO)
5,1 (39 ) 22,7(L721 L2,O(er)(100)
L 4 , O( 8 1 ) ( g O)
r2,2( 94 ) (102 )
fianHa o6uras fiauna
f npennoaaraeMatt ty"t.no"JIeHHas nocneaJlbeeonnPHoft
qACTH
5o 3T Byl pol
AJIbBEOJbI.
Panr,ryc 6prcunoft AnnHa npnBepunHgofi rracru flnavrerp clluHHo-6Prcurnofi a [pnBepurnHHoft rracrl{ 6oxosoft B nPI{BePluaverp rulrHHofi qacrl{ Yroa aJlbBeoJlnpnrtft B crII{HrpaA. Ho-6proruHofi naocrocrll, Ynoa Bepul{HHbtfi n cnnHno6pmunofi nJlocKocrn, npaA.
KOI
ltn )ral
.fet HG \{e
24
cTi oTl
4L
56
5ar
3at irxi
Oxou'ranue ra6nuuu Ne 87-35, JLorcer.a6ypr
ilapaverprr
npeAnonanaeMaq Anrsa ycTaHoBneHHas o6uag Anuua nocneanbaeonnpHoft qactu
54,1(541 ) 5 4 , L (5 4 1 ) g z , L (3 2 1 x S g )
80,O (+ A e ) 7 L , O( + S g ) s2,o(317)(OS)
Auavetp cnl{HHo-6prcuruofi y Bepul{Hbr aJrbBeonbl Auaverp 6oxoeofi y BeprIIrHbI anbBeonbr Paqrryc 6protuuofi AnuHa npr{BepuuHsofi qacru
1O,O(1OO)
16,4(1OO)
10,7(107 )
1 6 , 5 ( 1 O 1)
3,5(35) 1 5 , 3 (1 5 3 ) 9 , 1 ( 9 1 ) ( 1 O O)
2 9 . , 3r(z e ) L 4 , 7 ( 9 O ) ( 1 O O)
10,o(roo)(rro)
L4,7(gO) (roo )
Anar',rerp cnrHuo-6prourUOfi e npuBepuuHnofi qacrl{ Auar.aerp 6orogofi B nplrBepruuxnofi qagrr,r Yron anbBeonspHrtfi B crIHHHo-6prourHofi nnocxocrl{, rpaa. Yron BepullHHbtft s cn[HHo6prouruoft nnocKogru, npaq.
EB
BEI
t9
s.
19 Har
JIH H
35
50
6anxeHa x 6proruHofi cropoue. Ha Ha'ranbHblx cTaAIgx pocrp HMeJI 6onee scHo 86Ipa)KeHHyro qeM y B3pocJrblx )KuBoTlrbrx cy6roHxuecKylo {opr'ry: sHaqeHl{s IIa noqrn He orJrr,rqaJrr.rcbor ceoftcrBeHnblx BspocJlbtM oco6su (pnc. 8 ). l l o t n e p x e u s r x o a e 6 a n n s M B e J t n q n n af l a ( o r 2 4 7 n o 3 6 0 ) , HsveHquBocrb. ), pasMepbl BepuHHHoFo yrJla creneub crmuso-6prcuHoro c>xaruq (8L96-1O7 (35-560). B xonaerur{H ecrb pocrpbr, y Koropbrx cnuHHo-6oxoarle 6opoealt nst-
28
cK
tut6 .'Ibtr
2O 6op qac oce pbr
t eu
Pnc. 8. sp. nov.
flpoqoarHoe Ne 87J3,
c e r r e H l r e p o c r p a A c r c t c o el i t e s o m o l o n e n s i s S a c h s , Toap, p. Eyop-3exlr (x O,8)
pa)KeHbI oqeub pe3Ko, I{ pocrpbr c noqrr{ HegaMerHbrMrl cn}rHHo-6oKoBbtMu 6oposAaMl{. CyutecraeHnbrx r{sMenexr{ft poctprr pa3noro r€onopueecKora Bo3pacra lr r{3 paogsrx o6nacreft ue o6napyxr,rBaroT. CpaBHeHue. Pocrpbl onucblBaeMono Br,ulapegKo orrrr{qarorcrr oT 6onsurnrcTBa BIIAoB poAa Acro coeli/es orcyrcTBueM 6oxosono c)Karurr. }ro.t npugtraK ornr{traer Br.rE A.ornolonensis gr 6nxgxono K HeMy no cna6ofi cTeneHH yAnuueuHocrr{ pocrpa A.janenschi (lirnst). OrcyrcrByerr 6oKoBoe cx(arr{e y pocTpoe A.kedoftensl s sp. nov., onucaHue Ko16 pono [ago Bbrue, y A. unisulcatus (Brainville,LB2T) u A.suprapalatinus (Kolb, ]-9,42). 3ru Br4ltbrr.rMerorpocTpbr, ouaquTenrso 6onee yntl[rHeHHbre H SaoclTpeHHbte. QaqlIaJrbHaa npHypoqeHHocrs. P o c r p u A . o m o l o n e n s l s e C u 6 u p r .H r afileHbl HcK[IoqureJIbHo B FJIHHHcrstx SaUasx oTr(pblTofo Mops o Or.aoaoHcKoMu IleHo-OaeHeKcKoM paftoHax. CoeepureHno oTcyrcrByror srr,! pocrpbr B npu6pexnrrx MeJrKoBoAHbIx+alrnsx BufiIoficKoro r.r IleHo-XaraHrcroro paftoHoB. CrasanHo€ 3acraBJIgeT cql,ITarb, qro )KHBoTtrbIe)Kr{nH e o6ctaHoBKax oTl(pblToro Mopg, xoTg oTHOCI,ITeIbHOMaraq yAJIIIHeHHoCTbpOCrpOB yKasblBaeT Ha rO, qro oHH BprrA nn 6gtan TaKI,IMI{ xopouIHMH IrJroBuaMHrKaK BlrAbI c cnJIbHo BbrrrrHyrbrMr.rpocrpaMu. Bospact u r e o t . p a 0 I 4 q e c K o e p a c r p o c r p a H e H r . r e . B e p x H u f ni a u u c 6 a x 3anaaHoft Eapoust, HuxHttfi toap (sona l)actylioceras commune H aolue) - Bepxunfi roap (eoua Collina mucronata) CeeepHoft Cu6upu. Marepnan. S pocrpoB H3 BepxoB HH)KHeror{ Hr.rsoBBepxHeno roapa OruoaoucKoro pafioua - c6opoI A.C.Aarnca, 2 pocrpa r{s BepxoB HH>xHenoToapa HHooBbee IIeHbr u Onenexa - c6oprr T.H.HaasHseaofi n C.B.MeJreAHHofi, 1 pocrp Ho BepxHero nangc6axa /lroxcer.a6ypra - c6oprr B.H.CaKca.
P o a M e s o t e u t h iLsi s s a j o u s ,l 9 l 5 M e g a t e u t h i(sp a r s ) : D a y l ee t Z e i l l e r , l B Z B ,p \ . 2 6 ; L i s s a j o u s ,1 9 0 6 ,p . 5 4 ; S r o l l e y , 1 9 1 9 ,S . 36; B f ilow -T ru m m e r,1 9 2 0 ,S. I l 1 ; F o g e r, 1952,p. 712. R h e n a n i :D e s l o n g c h a m p lsB, 7 B ,p . l l 2 , l \ l a y e r - E y m a rl,B B 3 ,S . 6 4 1 ; l \ e r n e r ,l g 1 2 , S . 1 3 6 ;S c h w e g l e r1, 9 6 5 ,S . 8 7 . , 914, crp. 13, 196G, crp. 108. M e g a b e l u(sp a r s ) : f l a a h o e 1 M e s o t e u t h i sL: i s s a j o u s , 1 9 1 s , p . ! 4 , 1 9 2 b , p . 2 2 i l ( p r r r " r n o a u( u pars) J-9 32, c t p. 9, 19 4 7 , c rp . 2 O 1 , 1 9 5 8 , c rp . 1S 8; K ol b, !942, S . f OZ; C axc, Haaruneoa, L967-2, crp. 19. : a e f , I g 2 2 , S . 2 3 8 ; R o g e r ,1 g S 2 , p . - 7 L 2 . O d o n t o b e l z(sp a r s ) N Tnn poA a. B e l e mn i te srh e n a n u sO p p e l , L 8 S 8, aaJrenl oxH ofiqacrn epl . OnucaHI{e. Pocrprr s 6onsunHcrBecpeAn[e, cy6xorn.recKr{e, pexe cy6ulnuEApuqecrile,
c SaocrpeHHofi aepuuHoft, saHlrMaroutefi ueHrpaJrbHoe 1rofloxeg1,1e, mr6o cMelleHuofi x cnnunof, ctopone, co cr(arblv c 6oxoB oBaJrIrHbrM, pe)Ke oripyfrbtM nonepeqnbrM ceqenueM, or cr{JrbHo yAtrHHeHHbrx Ao Koporxux (fla or 1SO_
BbIITH
o), bI-
2oo no 12oo). v aepuunu qerrc{e ADe cnnHuo-6pnurrcre 6opoeA.sr. EprouHas 6opoeaa oTcyrcrByer l{qn cna6o BblpaxeHa. V Hexoropbrx BHAoB B rrpuBepuugHoft ,-/ qacru pocrpa ga6atoAatorcs Mearne 6opoeaKil vnil urpr{xu. BepurnHa aJrbBeoJrbrH oceBas rruEns cJterKa cMerueHbl r 6pnruHofi cropoH€. Ha HarlaJrbHbrxcraA[sx pocr_ prt cy6xoHuqecKue, oTHocHTeJIbHo6oaee yKopoqeHHbre rreM BgpocJlble. Bunoeofi c o c r a B . O m l c a n r t 2 0 e u r o e , r { s H r D (u a C e n e p e C C C P B c T p e q e g u l _ 1 . CpaaHeHHe. Or [p]'r-Hx poaoa noAceMeficrBa L,legateuthinae pocrpbr ltleso_ teuthis oTJrr{qalorcn npn npeo6aaAaroueft cxarof, c 6oxoe cy6xonu.recrofi Oopve
29
6ptotu6opoea H orcyrcrBl,IeM saauqueM AByx xoporuo Bbrpa)KeHHbIx cmdHHo-6oxoecrx I{ oT cxoAllblx c HHMII uoft 6opo3!.bt. 3r'or [pr{gHaK orJrnrraeT pocrpbr rllc.so/rrrllli.s no SopMe pocrpoB Ortltobelus u C.atatrutltis Ho floAceveficrsa Pa.ssalotcuthinae,
I \*
y Koropblx LIJII{ nnub eABa HaMeqeHbI. PoctnplrBepuuHHbre 6opoegrr orcyrcrBylor ,\lega teutltis orJruqaprcrr oT llr,.so/ t'ulhis paoBurneM y Bepuuubl rBtIpa)KeHHbIx orno6opoaa. a raKx(e 6oauuruv e pasuofi creneHr{ 6proruHori u 6proruuo-6oxosttx Poctprt Parantegateutltis cr{TeJIbtrbIM yrJrHHeHHeM pocrpoB B npouecce oHToreHesa. prr
sI lt N '
1. ,.'' oTJrHqarorcq ' i J
or
pocrpoB
Llesotettthis
npHcyrcrBHeM'
xoTrr r,r caa6o
npogB,nspueftcs,
\
3ar.aeqaHr,rq. Hauu B cocraB poAa Me.sotcuthis BK[IorreHbIHeoAHoKI]arHooru: cblBaBuuecs r.rs nnuHc6axa Espofibr pocrpbr c qerrt{Mn rrpHBepuI{HHbIMHcnuHHo-6o275, crp. K o B b r M H6 o p o e g a v u I B e l e m n i t e s v u l g a r i s ( \ ' o u n g e t I ] i r d 1 8 2 8 , r a 6 n . 1 5 , Q n r . L ) , B e l e m n i t e s e l o n g a t u s( P h i l l i p s , n o n l \ l i l l e r ) , 1 8 6 5 - 1 8 7 L , L962, crp. 138), crp. 50, ra6n. 7, {nr. 17 ), E. paxillosus var. ( (Schvtegler, B.paxillosus amalthei (Quenstedt, 1849, ctp. 401, ra6r. 24, $ur. 415;1859, 3nr pocrpbl o6sr.rHo ornocnrcs K rpylme pacrp. L76, ra6fl. 21, Qur. 15).|. xillosa (-poa Passaloteuthis) H TorrbKo KllI. Hyuy6nase noAo6Hblft poc-rp tre aoMepa Kaexaea c AocraroqHblM ocHoBaHr.reMorHecJra K poAy Mesoteuthis (Masoteuthis poxillosa Nutsubidze (non Schlotheim) (Hyuy6Nnser 1966, crp. 163' ta6a. 3 8, Snr. L ). HHonAa Ha pocrpax Mesoteuthis sa6npqatorcg BTopI+rHbIeysKlre 6opoeast cnr,rHHasH pe)Ke 6prcurnan (cv. ra6a. Vlll, fiur. 6). Tarue 6opoorct o6paeyrctca BcneAcrBue cnoco6Hocrn pocrpoB l\tegateuthinae pacKaJrbrBarbcs B crunHo-6prourHOM HarrpaB,neHHuH He AoJrxHbI rlnTblBaTbcs rrpn AlanHocTuKe, p a c n p o c r p a H e H r , r e . [ I a m H c 6 a-x a a J I € H Boepacr n reorpa$uuecKoe 3anaaHoft Fgportbr, Bepxr naxsc6axa - 6aftoc Cn6rpn n AaasHero Bocroxa. Onpe4e"nrreJtbHafi Ta6JtilqaBn4oB pola 1.
Mesoteuthis
Pocrprt c xoporuo pasBr.rrblMu npHBepur{HHbrMHcrmHHo-6oKoBbIMH 6opoeAaMH, JrriueHHble 6prourHofi 6opoelbt, cy6roHu'recrofi nau cy6uaJII,IHApI{qecxoft $opvsr, c)Karbre c 6oxoe.
2 (7l. 3 ( 6 ). 4 ( 5 ).
5 (6).
6 ( 7 ).
P o a M e s ot e u t h , i sL i s s a j o u s , 1 9 1 5 cy6ronurrecxre, c oBaJrbHbIMnonepeqHblM Pocrprr cHJrbHoyAJrr{HeHHbte, ceqenueM, lla oKono 600-800 u Ao 1OOO . . . . . . . . . . . . 3 Pocrpsr cpeAHr,re, Bepunna saHuMaer ueHTpaJIbHoe [oJro]r(eHr{e o . . 4 Pocrp c KoporKoft npnaepurr{Huoft qacrbro r.r xopouo BbrpaxenHbrMn npuBepur.runbrvn 6opo3AaMr.r,fla orono 600- 65Or M.raui (Werner, L972, S. 13O, Taf. 11, Abb.7) - rcap Pocrp c crrnbHo BblTrruyrofi npraepur{Huori r{acrbro n caa6o Bbrpa)<eHHbrMr{ rpnBepurHHbrMr.r 6opooaavn, fla 600-105O. - roap - Hr{xHuft aaJreH M. Iongiroslris Sachs, sp. nov. Poctp KpynHblfi c Bepur{Hoft, cueutenHofi K crruHnofi cropoHe, IIa oxoao
600-goo. Itl. pergrondis Sachs, sp. nov. - B€pxHr{ft roap - Hr{xur{fi aaJreH Poctpst yAnuHeHHbIe, fla oroao 400-600 8 . . . . . . . . . . o . . Pocrpst cyfxourr.lecKofi Oopvsr o . . . . . . . . . o . . . . . . . . . I I-loneperrnoe ceqeHne pocrpa oBanbHo€ . . . . . . . . . . . o . . . .1O Pocrp cpernnft, npnBepurnHbre 6opoealr pasBuTbr caa6o, IIa 3I-0-600. M. striolata ( Phillips, L865-18'T L, p. 51, pl. 1O, fig. 25 ) - HHXnnft roap 1 1 ( 1 2 ) . Pocrp r
7 (18). 8 (13). e(12). l-o (11 ).
30
L 3 ( 1 8 ) . Pocrpu cy6uunuuApnqecKori $opr"rsr . . . . . . . . . . o . . . . o . L4 1 4 ( L 7 l . flonepeunoe ceqeHHe pocrpa oBaJrbHoe . . . . o . . . . . . . . . . 15 1 5 ( 1 6 ) . flpnaepruuHHbre 6opoeAu pasBuTbr caa6o 16 (17 ).
17 (18). 18 (32 ). 19 (20).
I r l . . l a e u i s( S i m p s o n , 1 8 s s , p . 2 s , p h i l l i p s , 1 g 6 s - 1 g 2 1 , p. s7, pl. 10, fig. 23) - cuneMpp - roap flpnaepulllHHbre 6oposnei BbrpaxeHbl qeTKo M. rhenangJopp".l, 18S8, S. 489, S. 42g, Quenstedr, 1g46_1g49, I'at.27, I.'ig. 1) - nepxnnft Toap - Hr{)KHr{fiaaJreH llonepeunoe ceqenr{e pocrpa 6nusxo K oxpyrnoMy, |Ia Bbo-4so. Itl.pseudoelongataSachs, sp. nov. _ nanHc6ax _ xuxsnfi Toap P o c t p s t c n a 6 o y A a r . r H e H I { bI II ae ,o r o n o 2 O O - 4 O O o . . . . o.:..1g Pocrp oTrrerJrr{Bo cy6xoHu.recrofi o0pr"rsr, cpeAnnft, c KopoTKHMH rrptb Bepur{HHbrvu 60poaAaMH H OBaJrbHbrM[onepeqHbrM ceqeHHeM, [Ia
2 40-39o. 2 0 ( 2 5) .
hl.pyranidalis (Zieten, l8go, S. 30, Taf. 24, F-ig. S ) _ roap _ Huxsnfi aaJreH Pocrpsl caa6o Bblpaxenl{ort cy6xorrnqecrofi &opr,rsr c oB€urrrnbrMnon$ peqHbIM
CeqeHI{gM
2 r (22).
l. t-
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
.
r
.
.
r
.
.
2L
Bepuruna pocrpa saHuMaer ueHTpanbuoe nonoxerr{e, [pr{Bepu[nnge 6opq3lbr BbrpaxeHbl caa6o, fla oxoao gSO. I r l . s u b r h e n a n aL i s s a j o u s ( L 9 2 7 , p.22, pl. 2, fig. lg_2o) _ BepxHsrr qacTb Hr.rxHero Toapa 2 2 ( 2 5 1 ..Bepuruua pocrpa cMeIIIeHa K cnl{HHofi cropou€ . . . . . . o . . . 2g . 2 3 ( 2 4 1 . Poctp cpeAHufir c rlerK[Mtr, Ho KOpOTKTTMT{ rrpr{BepuuHtrblMH 60poaaavu, Beprur{Ha cJrerKa cMerrreHa K cnr{HHof, Cropone, fla oxoao 23o_39o. M.-inornata(Phillips, 1865-187L, p. gO, pl. 1g, fig. 46) _ roap_ oauoc 2 4 ( 2 5 r . Pocrp ge60asuroft, npunepuuHHble 60poerur BHpaxeHbI cna60, Bepur{Ha cHJrbHo cMeueHa K cnr{HHofi cropoue, fla oxono 200-290, M.subrostriformis vonrnez (Boponerr, Lg62, crp. 1oo, ra6n. s4 Snr. 3-11- ) - roap 2 5 ( 3 2 1 . Pocrprt cy6unaunrpFlecKofi @pr'6r . . . . o . . . . . . . . . . . . 24 2 6 ( 2 9 1 . lloneperrHoe ceqeHl{e pocrpa oBaJIbHoe . . . . o . . . . . . . . . . 2 7 ( 2 8 1 . Bepruuna pocrpa saEhMaer uenTpaabnoe [oJroxerr{e, pocrp r*relrrrrfi, IIa 320-46C, Hnoraa ua6apAaercs npuBeprunnnas 6prorunas 6opoena. M.tiungensis Sachs, sp. nov. - Hnxnrrft Toap 2 8 ( 2 9 1 . Bepruuna pocrpa cMeurena K cm{Enoft croponer pocrp cpeAmfi, fla oro_ no 34O-560. M.aequalis voronez (Boponeq 1962, crp. g6, ra6r. sg,$rr.2r7lnnxnnft Toap 2 e $ 2 1 . Ilonepe.rHoe cetreHlre pocrpa 6rurgro K ote)rrnoM)r . . . . . . . . . SO 3 0 ( 3 1 ) . Pocrp cpeanufi, fla oxoao 2SO. M'vulgaris (Young et Bird, 1828, p. 275' pl. ls, fig. 1) - roap 3 1 ( 3 2 ) . Pocrp vealcrf,, npr{BepunHHbre 60poealr pasBHTbr Cna60r- na oxoao 3 00-400. l + l . m a t i s c o n e n s i s( L i s s a j o u s , L927, p. 1g, pl. 2, fig. g_12)_roap 32. Pocrpsr KopoTKr{e, cy6roxn.recxofi @pr.,or, fira r.,reuee 2bo . . . . gg 3 3 ( 3 6 ) . Ilonepe.rHoe ceqeHl{e pocrpa oBaJIbHO€.. . . . . . . . . . . ... 84 3 4 ( 3 5 ) . Bepurnna pocrpa oa'r.rMaeT ueHTpaJrbnoe nonoxe'r{e, fla oxono . 1 75-2 00. ll.subpyramidalis(Lissajous, Lg27, p. 17, pl. 2,fig. S_g) _ BepxHsc rracTb Hr{xHeI.o Toapa
3 5 ( 3 6 ) . Bepuuna pocrpa cMeulena r cnnunoft cropone, M . br ev iro s tri s(d b rb i g n y ,
l.
HuxHnft
36.
aaJreH
fla oroao
L g 4 2 , p . 96, pl. 10, fig.
L4O-24O. 1-6) - roap-
flonepeuuoe cerreH'e pocrpa 6mrexoe K oreyraoMy, Ila orono lso. l + l ' s u b b r e a i s( K o l b , t g 4 2 , p . 1 s 4 , pl.10,fig.7,1s, 16) - aepxrurfiroap
3l
M e s o t e u t h i sp e r g r a n d i sS a c h s , , p . n o u . l Ta6a. Vl,
$ur.
1-3
Myeefi Hff CO AH CCCP, Hoeocu6upcx. Pexa llcBbtfi N g?J8. loaorun KeAou (6accefiu Or',roaona),nepxnuft Toap, sona Pseudolioceras rosenkrantzi. cntpocrp KpynHbrft,cnJlbHo yAnHHeHHblft(fla oKoJro 600-800), AnarHos. nonepeqoBaJIbHbIM c ( ), 85-90 oxono EE 6oxoe c 6o cy6xoHu.recxnft, cxarsrfi HbIM ceqegr{eM. Cnungo-6oKoBbre lrpnBepuuHHble 6oposgst pesKl'Ier llpor9rllBalorcg tro Ll 4 lla. Bneru Hue nplISHa Kn. fpueepruragHanr{acTb pOCTpa yAJrI{HeHatCOcTaBJItleT BepoKoro L/2 IIa. BeprunHa 3a-N4erHocMeureHa K CnI,IHHOfiCtOpOner saocrpeHar 6oposalt crI{HHG6oKoBble (ra6a. 9). IIpueepuHHHbIe unHHbrfi yron paBeu 21-260 Ho KopoTKile. Hapnny c HI{MH y BepuHHbI c 6puuHofi cTopoHbl ysrore u rny6orxe, Ho AopocTpa Ha6atoga1gTcg Tpu 6oposnrcl' y3KI'Ie' oqeHb roporrure (lo 10 vv), cropocrarotrHo rrerKge. Eoroette cropogbl MaJIo ynnoueHblr cnI{HHas u 6proruHaJl HbI NOTITU OAIIHAKOBO BbINYKJIbIE. AabseoJla saHI{MaeT oT t/3 u OHTOneHes. npr{sHaKrr BuyrpeHHr{e cMes aJrbBeOtIbI H oceBafl nvKtrg' saMerHo BepuNna no L/ 4 o6utefi AJurHbr pocrpa. y BepuIHHbI 34*35o/o AHaMerpa paAlryc oKoJIo (6prouruofi uegbr r 6prcuHofi cropoxe Ocesas JIHTII{I norrrl{ rlpgMasr caa6o ynon oKoao 25o. aJrbBeoJrbti. Arr"r"oaapnrtft (prac. 9). Ha na'IaabHblx r 6prouHofi cropoHe r{ HecKoJluro npu6au)Kega uoorHyTa Hecy6xoHlfqecKylo oopvy' cTaAlrgx pasBuTHrr pocTpbr r{MeJrr{ n)rque BblpaiKeHHyro flpt't rIeM xg{BorHblx. pocrpbl repocJrblx BbrrrrHyrbl cKoJrbKo MeHee orHocHTenbHo Ar{aMerpe ( CE ) 8,8 rurv sHaqeHue fla paeHo 5OO.
T a 6 r r uu a I Hsvepenus pocrpoB M e s o t e u t h i sp e r g r a n d i sS a c h s , s p . n o v . N e 8 7 -3 8, p. Ileestft KeaoH
Jlauna o6urag Aanna qacrl{
fnpeanoaaraeMan \ycrauoaneHHatl nocJleaJrbeeoarPHofi
flnaverp BepuHHbI
cnuuso-6PrcurHofi Y aJlbBeoJIbI y BePUI{HsI 6oxosofi
finaverp AJIbBEOJIbI 6prourHoti Paauyc lanua
aJrbBeoJlflpnblfi
1 6 , 8 ( 1 O O)
1 8 , O (1 O O)
14,5(88)
15,5(86)
60,0(gsz ) 1 3 , s ( 8 2 ) ( 1 O O) L 2 , 3 (7 3 ) ( 9 1 )
ffffiUar.) 32
nJlocKocrur
25
B crIHH-
naocrocrl{, rpaA. uo-6puuuofi Yroa BepuluHnbtfi e cnHHHo6ploruxoft
1 6 0 , 0 ( S A g) 160,O(889) 1-r2,O(622)(ZO)
6,1(34)
lrpHBepuHHHoIi qacrx
e cnnune6Prcutuoft [uaverp IIpHBepuHHHOfi .racrH 6oxosoft B nPHBePlluar'aerp uruHHofi rracrl{ Vroa
189,O(1099) 1 8 9 , O( 1 0 9 9 ) 123,5(73s x 65 )
N e 87-39, p. Ileestfi Kenon
rpaA.
- oeeHb6oaguroft.
26
Puc. 9. [poAoJrbHoe ceqeHr.rep o c r p a Mesoteuthis pergrandis Sachs. sp. nov. Ne 87-39, eepxnnri T o a p , o o H a P s e u d o l i o c e r a s r o s e n k r a n i z i , p. Ilenslfi Ketou ( x O,gS) tA
I'l e NaeH q r{B o cr r. B Br{Ay [email protected],r Marepr,raJraMoxuo orM€_ Tr'rrb nr{rub xone6aHrs B OTHOcl'TenbHoft yqnmreHHOCTH pocrpoB (lla or 622 ao 735)' srene'r' Bbrpa)KeHHOCTT{ npr{Bepu[HHbrx 60poea r{ creneHLr BbrpaxexHocrr{ cy6ronuvecxoft {oprraul poc,Tpa. CpaBHeHne. OnucbrBaeMbrepocrpbr nau6onee cxoAHbr c ToapcKr{M Il.raui (werner) fie pxHofi qacTr.ropl, orn'qarrcr 6oaee KpylHbrMr.{
r qbr
PAsMEPAMI,I,CMEIIIEHI{CM
EbI F
b TTbI o
x
'r/
Ne 87-4O, p. Toxyp-IOpxx
1 O 5 , 5 (9 2 6 ) 105,5(gzo ) 7 6 , 3( 6 6 9 ) ( t z ) II,4( roo ) 9,6(84) 4,O(35) 34,7(3OO)
ro,2(89)(roo) e , 3 ( e z) (g r )
27
K CNrrrrrOft
CTOPOHE
BEPUI.IHbI
I,I
HA PA3BIMI{CM 6prourHoficropoHe 6opoeaor. or M. lapiinskala" Yoronez paccMarpr{BaeMbIe pocrpbr ornrqarorcs 6oasureft yArrlrHeHHocrbro r{ n}nrue etrpaxeuuofi cy6xourrecxoft sopvofi . M. rongirostris sp. nov. HMeer roH*r{e cy6xoHr{trecKr'repocrpbr co cna60 pasBurbrMl{ npr{Bepu[HHbrMu 60poeaavu. Oaur{aJrbHarr npHypoqeHHocrs. pocrpu co6panbr B nper{My_ ruecrBeHH' nnl{H}rcrbrx ocalu(ax 6accefina KoauMbr BMecre c aMMOHI,ITa_ MH' B[aJrr{ or topcKono MaTepr{Ka. cynn no y4n'HeHHocru pocrpoB, )Kr{_ BorHbre Benn ceo60aHonra-eaburEft o6pao xrrsHH. B veurcoeo*"o .rpn6pe>rurx Oauusx gror BuA He Bcrpeqaercq. B o s p a c r r'r n eo '- p a o nnq e c K o e p a c n p o c r p a H e H n e. BepxHuft TOap - HUxsrli aaJreH ceeepHofi cn6wpw. Marep'an. 5 poctpoB- Hs Bepx'eno roapa I,r Hr{xHero aaJreHa6acceftna Koasrsr - c6opsr A.b.Aanaca.
Al. aff. pergrandis
N e 8 7 -4 1 , p. Ileerrft Keaou 1 5 2 , 0( f S O O ) 752,0 (13 66 ) L26,s(1128 )(eg )
Ne 87-42, n-oa YproHn-Tyrvyc
9 1 , O( r S s e ) 84,7( 1235 ) 7s,s(111Oxes)
r r , 2 ( 1 O O)
6 , 8 ( 1 O O)
1 O , O ( 8 9)
4 , 8 ( 7r )
4,O(36) 4 3 , O( e e + )
8 , 5 (z o ) ( r o o ) 7,8(7o )(92)
25
2,2(32) 24,O(353) 5 , O( 7 4 ) ( 1 O O ) 4,2(62)(e4)
16
3 4 9 33
Mesoteuthis aff. pergrandis Sachs, sp' nov'
Ta6a. VIl, Sur. L-2 pocrpa, Koropble flot rmsBaEr{eM Mesoteuthis aff. pergrand,i's MbI onucaru rpl{ yAJrHHeHHocrbIo orltocureJrbHofi 6orsuefi M.pergrandis or rnnlltrHbrx or,''qaro,rcs [pI{MeEbull{xsllaqeHugxA[aMeTpa.BnoageBosMoxHo'tITogrnpocTpblloTIIlI{8oqeub 6onsuofi yAInEeHHocrbK], ,cb or pocrpoB Bcex Boo6ue BrrAoB Mesoteuthis IITO pOCTpbt TUIIftIIlbIx BOgMO)lQtOt TeM C Br.aeCre BUAy. HOBOMy IIpItHaA4e)KaT K 1tpeAc163.1groTJlulub tpaflnue qtregrt oAnoro lvl.pergrandis u M.aff. pergrandis pSAAr KoTopblfi nO Mepe BaKOmeEI{s MaTepuaJla OKaxeTCs Bq?MOxHbIM OTHeCTU l( oAHOMy BuAy. 11pr{sgaKr{. Pocrp oqenb cIUrI'Ho yAJrrHeEHbIfi(sxa'reEsf, na o'(ob Bueunne caa6o cy6ronnuecKr{fi, cxarbtfi c 6oroa (EE 4O-9O), c oBa;1EF no 11OO-11,5O): Ilpnsepulrggble C1HHHe6OKOB6Ie 6OpOorct pasBIITbI CeqeH14eM. r6tM nonepetlgblM oKoJIo Caa6O. IlpnaepuugHas rracTb pogrpa CI{JIbHOB6ITsHyDat OHa CocTaBrrseT Bepulrlttlblfi cropOnet CmIHHOfi K CMeuIeHa L/ 3 tla, BepuI[Ha saocTpepar CJIerra yroJr o*oJro 2So. flpnaeprunHnbre cmtnno-6oKonble 6opoeqbt EeF ly6oKr€1 rlpoGll€xlrBbInyKJIbIer 6oroBarcTcs oKoJro 1 cr"r or BepurrHbt. Cmrnnag n 6pnunaa cropottbl qagTrr pogTpao ylg1orr1ega1 B flepeAgefi Bbte'cTopOtttt Caa6o BbtnyKJIbIer HecKOJIbKO oBaJrl'Hyro pocrpa cot(paHser oopruy' 6ya]qu Bcefi AJrnHe no floneperrnoe ceqeHl|e EE-71-89t BecKoJrb6o 6oaee cxarbrM c 6oxos B nepeAgefiqacm pocrpa(gna'legns
(
6#+921.
\ Arbseoaa saHuMaer o1oJto 1/6 aauxu pocrpar npr{sHaKr. BnyrpeHHne cE). 32-36% palxyc cosraDnser Bepuu*a ee cMeuena K 6proruuofi cropone (6plqurrol r yrofl He saMepsncg. Oceaas nptr6mtxena ilrrntrlr cnerxa fuiuaeongpnrrfi B nOnes HO BbtScHeHt He paccMaTpHBaeMofi $opt"tst OntOreHeg 6pnrlnOfi CTOpOHe. paEp peqHblx ceqeHxsx pocTpoB BI{AIIO, qTO EapauluBaHle BeIlecTBa pocTpoB lu[o HoMepHo n o6pasoBaHne em{pocrpa He [pogcxoArrno' B BanueMpacrropsxenux [MeIoTcg ronbKo rprr pocrpal xaHgveHquBocrb. SHaqeHIrsMH paKTepHsyrouuecg, KaK BITAHOtI3 Ta6I. 9, 6nrsrorMH oTtl@HTeJIbHbrM[ Merqy I{ Atlrltlbt. napaMerpoD, Ho rrptr pasHblx a6coarmtorx BeJrrqnHax ArraMerpa yBeJlt{rrec BospagraeT Mesoteuthis pocTpoB yAJtrrHeHHOCTb T€M, oTH@nTeJrbHan o6beallHeHue sTIlX HUeM alraMeTpa. flostoL'ry ao BblccHeHgs BHyTpeHHero cTpoeHug ycJroBHblM. pocrpoB B OAHOMBilAe ocraercg Onu6lBaeMble poCTpbI peSKo oTaur1arcry6goT Bcex Apyrlx r€Cpan6eg[e. 6rr 6btts oTHeceHbI BecTHbtx e Cn6npu l[ no cBoefi cl|JlbHofi yAIHHeHHocT[ Mornl| s'alpocrp. Orerr tlTo oTcyTcTByeT nocKoJll'Ky oTnaAaeT, poAy Satpingoieuthis, K (Zieten, 183OtcTp.28' longissinzs cxoAeg no BHeuHefi 4opMe pocTp Belemnites .Iact11 y He BI{AHbt npnHefo OPfr Ho toxHoft lopbt ns nnxnef, ra6r. 21, Qur. 10 ) Bepnepor'r (Werner' BepultgHbte 6oposApt rr EooMoxHO oH rpaBl{nbno DTcIIKFI€H 1920, crp. 1O7) n Eronon-Tpynr"pon (Bfilow-Trummer, 1g12, crp. ltgl Salpingotcuthis tyb!s cuHoErMIrKy 2OG) crp. (Schwegler, 1969, tU"."n.pou (Miller, 1823t longissimus Belemnites young Hnxnerrauuc6a)rcxile et Bird. laris
crp. 60, t.6n."'d; Or;. i:ti-no^r"t"rorcs
Jlaurov (L1ng, 1928, crp. 211) e
a cyfUrarrgAplrqecrc{e pocTOu uvepT He cy6rorunrecxlet pOa Pscydohdstites. omtcannblfi Ouruuncor'r (Philpocrp B. longlssimus Miller, prr. HnxHenannc6axcgfi sgaqureJlbgo cy6rogrrecruftr 32), crp. 66, ra6n. 13, Snr. 1865-1871, fip", ("""*."re ). 17OO 6onee fla p ergrondis aff. if . 6oaee r"rrt"yr V ""n n p u y p o q e H H O C T b . A B a a s p o c J l b l x p o c r p a n a f i A e t l b tB I I p e QaunarrbHas MaTepUKa. MeI' xMyuecTBeEBO nJr[HlrcTbrx ocaAKax s 6accefige Korur*bl BAaJlU OT ocaAKax necqaHo]-rJll{}orcrbtx B Me[KoBoAHblx o6napyr€H (prorf,?) er(3er"{narp *i qTo xl{BoTHbIe BeJII' cyA's rro pafiOge. Moxgo AJ7114aTbr a Aga6apo-)GraUrcxor.r cHnbHOfi BbITEHyTOCT1{pocrpoB, CBofuA}IorulaBaroufufi O6pae rlcrsHtr' p a c n p O C T p a H e H x e . B e p x n s cq a c T b H reorpaSu.reCKoe BospacT Zugodactylites braunianus ) - HIrxHnfi aaneu Ceaepxof, C)tHHr(HeFo roapa ("o"t 6upn,
34
J I
I
T
3
f
t
r I
E
6
n
H H H H
6
n u u
B
T n
H
M B
1.
pr JII
T
MarepHan. 2 pocrpa Hs BepxoB Hr.r)KHeno H BepxHero 6accefiua Ouone Ha - c6opbr A.C.,[anuca, 1 pocrp Hs "oapa l{r.rxueDo aaJreHa n-oBa YprcHr-TyMyc _ c60_ PbI T.H,HaJrbHseBoft.
M e s o t e u t h i sl o n g i r o s t r j S s a c h s ,r p . n o u . l Ta6n. VII, Snr. +7 loaorun Huxnnfi
t
E).
, l-
Toap,
Ne 87-43, Bepxngs
Myeeft l4lr co AH cccp,
qacrb.
Honocn6upcn pexa Bnarcfi,
AnanHos. Pocrp cpeAHI'tIi, CHJIbHO yAJrr{HeHHblfi (enavesus [Ia Mexlly 600 n 1o5o), cy6rounvecrcnfi, crxarotfi c 6oxoe (EE 7b-loo), nonepeqtrblM CeqeHueMo ilpueepurrnHbre "--or"rr"orM 60poercr pasBr.rrbr caa60. BHeuHne np'sHaKH. flpuaepuruHH€rs qacrb pocrpa cHJrbHo BbITqHyTar Aocrr{raeT oKoJro.noaonuHH fla, BepuHHa saHr{MaeT [eHTpanbHoe noJror(egr{e, ocrp€rs, BepuI{HHbIft yroa HaxoAlrrcs y BspocJrbrx eKseMrrrrspoB B r{pe[eaax rs-rgdf rri"]of B6auen BepIuHHbI caa6o HaMeqel{bl npr{Bepur{Hnbre cm{HHo-6oKoBbre 6opoe4rr. 1o-. lepeqHoe ceqeHue a eaAHeft qacTH pocrpa MeHee cxaro c 6oxoe, rreM y Bepurr{Hbr arbBeoJbl, c[r{HHa-s u 6prcuxas cropoHbr oAHHaKoBo Bblnyrglbl, 6oxoasre cropoHbt TaK)Ke BbIrryKJIbIe, Ho B 6oaee caa6ofi crerreHu. BHyTpeEHI{e npI{3HaKI{ a oHToneHes. Argseona saHuMaeT oKoJIo L/ +L/ 5 lurrt{bl pocrpa, cJIetKa cMeueHa i 6prcruHori cropoHe ( 6prc'rxoft paAnyc cocraBJlser 3445o/o cE). Aagseonsprorfi yroJr 2Lo, ocesaq rilr'urr noqr' upcM€I.s, JII{tub eABa usonHyra r caa6o cMelueHa x 6prourHoft cropoue (pnc. 1o). H.,i"qaJIbHbIx cra'q'Hsx pa3BI'ITI{s pocrp cy6xoHu.lecrnfi, orHocr{Ten""o MeHee yA[uHeHHtlft 'rev y R3pOCJIbIx el(seM[JrspoB (eua.renxe IIa oKoJro 4oo npu cE 3r5 MM), B AaJrbtreftuev pocrp cuJrbHo ylnnlrseTcq r npr{ CE oxoao 6 vv fla qdcrnraeT eHa'telilft' 6rnSKItx teu, KoTopble ceofictseHHbl pocrpaM BgpocJlblx xuBol6rx (y
eKs. ]s 87-45 npu cE 5r8 r'ru fla 64g, npu cB grs MM 62g1. I4eveHqnBocrs. Mengprcs sHaqeHngna (or obo no ro-do--)r'ss (n npeg3 AeJIaX Or 7 4 no ), crelleHb BblpaxeHHocTr{ rrpHBepunHHbrxcnunseSorosrx 60pq3& y HeKoTopblx pocrpoB oHI{ orcyrcrBylor, y HeKoTopsrx (pocrpbr Ne gZ-44, ]s 87-48 ) nrtpaxe'bl oqe'b qeT*o H flporfln]rBarorcs no,qrr{ Ao oKo}rqa'[s npnBepuluHtloft 'racrH pocrpar Qopr.aa pocrpoB - oT yru:rr{HegHo.
(nar.)
- c AJTHHHbTM pocrpoMo
35
Ta6nttua
10
iortgirttstris Sac'lts,s[)' nov' i4suepeuuq PocrPoB , 1r1s o l c t t l l ; r s
N 87-43, p. Bunnft
I-lapaverPsl
AnnHa fnpennonanaeMas o6uraq I y.ranoaneHHan ,lau ua'rrocrlc&JlbIJ€onspHofi qacrH cnuuuo-6prcul!.uaverP Hofi y Beptullubl dJIbB€OJIbI
flnar'reTP 6oxoeofi Y BepulrHbl aJIbBeoJbI Paglryc 6Prourgofi Aar.r Ha rrPHBePul{HHoft qACTI,T flnav erP crna sHo-6p]oulHofi B nPueePurllHuofi qacru e 6oroeofi !.uavetp qacrl{ npl.IBepuusHoft e Yroa aflbBeonspHllfi [noccnnnrro-5prcuHoft Kocrll, rpaA. n Yroa aepuunuttft inprnHo-6prour Hofi
88,O(978) 8 0 , 2 (8 9 1 ) 69,O(ttt)(zg)
N 87-46, P. TroHr'
s +,oir o8o) 52,O(r o+o)
N, 87-45, p. Bra.rrrcft
90,o @+l I 74,2(ZAr) 68,7(623)(ZO)
9,o(roo)
5 , O( 1 0 0)
9,5(1OO)
7 , 7( 8 6 )
4,L(821
9,2(98)
1,7(34) 3 6 ,2 ( 4 O 2 \
L8,2( 3 64 )
4,O(42\ 28,-7(3 02 )
7 , o ( 7 8 ) ( 1 O O)
4 , 5 (e o ) ( r o o )
' 7 , 7 ( 8 )r ( r o o )
6 , ' t( 7 4 ) ( 9 6 )
4,o(80) (89)
6,9(73)(90)
t6
L2
19
IJIOCKOCTHTI'POAr
rIacToTbI BCTpeqaeMocTI',I Ha ypoBlle fla oKoJIc 600' pa3AeneHHbre pesKHM CnaAOM cTaAI{sx (npn ruraveTpe pocTpoB' Ha Hat{a'b'blx CyAn ,,o BHyTpeHHeMy cTpoeHHIo oT lpyna.' [pyT He oTIMqaIoTc't BuAa r.,rvr) o6a 2,5.3,5 oKollo pocTpa sHarIeHUs u yaJTEHglo/rctt 6onee Bce M longirostris pocTpbl B Aaru,HefirueV eoePacralor. lla MoryT BbITgHyTbIe pocTpbt tl'longirostrts flo c,renegl{ yAnHHeHHocTI{ Hau6oaee Toapa+aHHero nosAHeno I'I3 pergrandis 6trrs cpaBHr{BaeMbt c pocTpaMI.I Al. aff. BbITgHyTbl (fIa oKono 11OO)' aareHa cu6upra. Onnaxo noCneAHHe 6Onee oopvty' Cy6rOin'recKylo Kpyrrrrbre, r.rMeroT MeHee rrcHo Bbtpax(eHHylo Hs 6acpocrpbl elrllY orgoctrTcn tITo paccMarpl'IBaeMoMy K BoevoxHor noA Ha31 9 5 3 ) ( L 9 4 7 , K p u v r o n b u e M cefiua p. BunrcR, onHcaHHue l.fl. Ha OnHcblsaeN4rlfi stintula. Bueurue oHn OqeHb IIoxoXH BaHr{eM filesotettthis ecrs 6ptouHag KpurMnonbua l'f' sKseMIUIt{poB u3 y oAHOro HO EaMr.r Br{A, 1864( D u m o r t i er' s t i n t u / a 6opOetar qro c6nuxaeT ero c Acrocoelites B FIaueM pacBTopllqHaq' qro 6oposaa 6prcuman He HcKnloqenor L874). c BTopl{ttHofi 6prcumoft 6oposAofi' noprDKeHr{H LlMeefcg pocTp M. Iongirostris 6onee 1OOO e Hauleft KonneKul{I{r Hacq}ITrtealoueft xe 1. stimula Pocrpoe Her BoBce' C u 6 t ' t p l l ' T o a p a pocrpoB Nlegateuthinae'I{3
I
I I
(
t
ll
S
( PHc. nov.
36
longirostris Sachs, SP' I - l p o A o a u H o e c e q e H l { e p o c r p a L l es o t e u t h i s p. MotoPqYHa a a J I e H ' H u > r H u f t Ne 87-48, 10.
H
\1, 87-44, p. Buarcri.i
N 87-96, p. Trcur
97,A (A+g ) 7 9 , O( O e z )
93,O(93O) 8 7 , O( e z o ) 68,6(686)(76)
1 l - , 5( 1 O O)
1O,O(rOO)
N 87-47, p. Trcnr
Ne 87-48, p. Morop.ryHa
8 1 , O( r r s z )
s s,o-(oee)
7 7 , 8 ( l _1 1 1 ) 6 3 , 1 ( e O 1) ( z g )
49,O(Ors)
7 , O( 1 0 0 )
8 , O( 1 0 0 )
9,O(78)
7 ,g ( 7 g )
6,2(89)
6,l-(76 )
4 I , r ( s s z)
39,O(390)
29,5(42 L)
27,5(3 44')
9,2(AO)(rOo) 8,4(73)(91)
8,6(AOXTOO)
5,7(8t_)(100)
7,O(88xrOo)
7 , 6 ( 7 6 )( S 8 )
5 , 3 ( 7 6 )( 9 3 )
s,8 (73)(83) 2L
19
J. fe 4., lc
)tr I
19
19
cDaunaJrbHas npHypoqeHHocrg. pocrpu B octroBHov c o 6 p a n b rB c p a B _ HI'ITeJIbHoMeJIKoBoAHbIxnecqaHo-nrlHHucrbtx ocalKax Buarcficxoro pafiona, oqenb PeAro B rJrugr{crbrx ocaaKax oTKpbrrono Mops s 6accefixe Koasrvrt. Moxero EpeA_ noJlafaTb, qro )r(uBoTHbIeo6nraau B MeJrKoBoAHbIx npn6peruttx o6cranoBKa& Hor cyA.g no cr.rJrbHortylanneHnocru pocrpoB, BeJrH cBo6onrronaaBaroulnft o6pae xHsHI{. Boepacr n reonpaSu.recKoe pacnpocrpaHeBne. B e p x n s sq a c r b HH)KHeroToapa (eoHa Dactylioceras commune n nsrure) _ rrnxHxtr aaJreH Ceaep_ ttott Cn6npu. {-
lC-
l(otr-8o
t3-
tq .t2
ttfi ag,
tcrfi.
t0 I
oo
sp.
t7
Puc. 11. Hsven.rneocrb pocrpoB M es o t e u t h i s l o n g i r o s t r i s S a c h s , sp. nov. u Mesoteuthis striolata (Phillips) ro orHocureasnofi rvnt_ He nocJreanbBeorspnoft qacrn ( IIa)
6 ,
z q00
600
800
/000 lIa 37
6accefina p' Bunrcq -L 37 pocTpoB n3 BepxoB HH)KHenoToapa Marepra aaJIeHa Hu3oBbeB Huxnero I{3 poCrpa 2 C6Opot T.H. HaaggCesofi v T.V1.K4punofi, u BoctoqYpnnn-Twyc rF-oBa aareHa pocrpa u3 2 flenrr - c6opu T.l4l(rrpranofi, Toapa HI{xHeno BepxoB 4 pocrpa ns Bqrrg Tahrarrpa - C6Oprt C.B.MeneAnnofi, A.C.Ilaruca. c6Opct Httx(Heno aaJreHa 6accefina p. oruonoHa
Me s o te u thsi s tri o l a ta(Ph i l l i p s ) Ta6n. VI, Snr. 6-8 p.-59,pl ' .10, fi g' 25' I' s' b; l \erner' : i l l i p s , 1 8 6 5 -l B? -1, B elem nit ess t ri o l a tu sPh
1969's' 196'Abb'79' 191; i: i.za,Tuf.-ii, nig' r; sc'hweglet' 1908' crp, 46, ra6n. Belemnites aff. compressusi Bopncnr, Mesoteuthis /ongdsslma: I(prtr'anonur51932,
CTpo 17, ra6l.
18,
4' Ur. 1, $nr.
19.
l-4-16.
,ra6r' 10' au6panrotfi B.H.Carcou - Phillips, 1865-187L, JlerroTun Bnr6n. $nr, 25 l. ToaP Arnuu, cy6xonnyAJuHeHHbIfi(eHa'lenns fla 310-600)' cperurnfi Pocrp AnanHos. flpttceqeHueM' nonepeqgbtM (Sn osanbgblM c Z5-96), qecgfi, Oo** cxarsrfi " cna6o. Bepunnnble 6opoenu pasBlrrbl cocfpuaeprunggac qacTb pocTpa cuJIbHOBblTsnyTar npusBaKn. Bneunne ocTpas' noJloxeHuet TaBngeT 6onee [oJloBr'rHbt flar. eepul{Ha saHI{MaeT ueHTpaabHoe Bepurl{Hbt cna6o HaMeqeHbl Beplun'Hbrfi ynoa pa'e* 22-25-0 (ra6n. 11 ). B6nngu caa6o BblpaxeHHas na6npAaercs y Bepuugbl HnorAa 6op*aot. cmrnrro5oKoBbte qacTu HeCKOJIbI(oMeeno lrrTpr{xOBaToCTI'. flOnepevnoe CeqeHue pOCTpa e eaA}refi Cnnn80-95)' BE npr (66 85-95 qeM y Bepullrgbt Hee c)r(aro c 6oxoB "rrtt"t*, BbtnyKnble' 6oxosrle cTopoHbl BbIII]KoAHHaKOBO pocTpa Eag I| 6pnrunas cTopoHbr HCr1€346OxOArIe 11OJIOCbI' Jlbte B prenUuefi CTeneHIrr Ha HHx na6A16Aatgrgs napHbte qacrl{. toune B nplrnePunnuofi Ta6rIH ua
11
(Phill ips) I4svepenns PocrPoB M e s o t e u t h i ss t r i o l a r a
]t 87-50, p. BnnPft
Anusa fnPennonaraeMas o6ruag lY.t""ottteHHas Ilrurxa nocJlearbaeon*P Hofi
6 3 , 0v t e l
5 5 , O ( O Z g) 4 2 , 8 ( 5 2 8 ) ( O e)
qACTfl
crnrnno-6Prcuuofi Y llnavetp Bepult{Hbt aJIbBeoJrbI Ilnar"rerp 6oxosoft Y BePUI{HbI
38
7 O , O0 t + ) 6 8 , 6 ( ? O O) 42.6(438)(61)
8,1(1OO)
9 , 8 ( 1 O O)
6,8(84)
7,8(80 )
ATbBEOJIbI
Pannyc 6Prcrunofi qacrn Annna llpuBepuuttttofi a cruHno-5Ptounofi lluar'aerp rlpuBepuunnofi qacnr Jlnar'aetp 6orosofi B rrpuBepunHnofi qactrl Ynoa aJtbBeoJrsprurfi B cm{HHo6prcunofi ruIocKocrl{r rPaA. Yron Bepurl{HHbtfis cmnHo6proruttoft nJlocKocrnr rPaA.
N e 87-51, p. TrcHr
26,2$241 7,o(86)(1OO)
3,3(34) 3 O , O( S O O) 8,8(89)(1OO)
6,1(7sx87)
7,5(72)(AS)
23
22
9.
BHytpeHHHe npusHaKI4 U OHTOreHes. Ansseoaa s a H u M a e To r o a o l / 3 ltIuHbI pocrpat BepuEHa ee cMeIIIeIa K 6prorunoft -rr""n,cropone (6prorurrof, paAr{yc o*oJro t/ 3 cE ). Aaseeoaspnrrft yroa 3 10. oce;.r roqru rrpsMas, npu6auxena K 6prourHoft cropo'e (pnc. L2'). Ha Ha.raabHbrx craAusx pasBnrus pocrp cy6ro'uvecxufi, IIo crelre1u yIU:MHeHHocrt{ MaJro oTJltlgatounftcg oT BspocJlblx gr(oeMrurgpoB (l-la 5oo npn cE 2,g uvr 4go npn cE -7 i,"r*rr 47L npu cE grg r.ar.ay erc). Ne 87-51). Hs l',re Htr H B oc rb. MeHgroTcs B saMeTnbrx rrpeAeaax snaqems IIa (310_600), EE ( 75-96 )' floAaangputee 6onuu'HcrBo pocrpoB xapa*Tepngyercs s'arreHusMu rla vex
Ne 87- 52, P. Mapxa
6 3 , O( t z e ) 5 4 , O( o z s ) 46,7(sOg)(zs')
8 7-53, Mapxa
54,O(ssr) 46,2(47Ll
8 , 1 (1 O O )
9,8(1OO)
7,8(9o)
8,1(83)
26,6(328) 7 , 2 ( 8 9 ) (1 O O)
3,2(33 ) 28,4(2 90 ) 8,s(87)(1oo)
6 , 7 ( 8 3) ( e s )
7,2(73 ) (es )
51, O(ogg ) 51,O(Ogg) 3 2 , 3 ( 4 4 3 X O S) 7,3(lOO) 6,6( 90) 2,5(34) 2O,7(2841
6,5(89x100) 6,2(8Sxgs)
31 23
25
23
39
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tt
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L2,
(Phil-' fponoau*oe ceqe'He pocTpa Mesoteuthis:striolata
roap, p' Mapxa lips). Nes7-53, HuxHI{ft
ilu/il I llll I \:lf I I/l y I I \l l t tI
U/ V
ocaAKa( Ana6apo-Xana'r€Kono B MenKoBoAHbrx ilectra'o-rn'Hr{crblx pafioHa r4 sa Tafivutpe r B lrpeuMyluecrBeHgo 1.nngucrblx @aaKax pafio'a BAaJru or ropcKofo Marepura. cyAs no )rAnnbr-ro".*oro o6pae xu3pocrpoB, xl,IBor'bte BenH cso6oA'ornaBaloul'fi o6cranoeKaMrr. ""rro"t" 6unn cBssa'br c npu6pe)K*brMr{ MenKoBOAHbTMH ,r, ' r ^Hha^mnalrorfllCHII}("o u reorpaSuu-ecroe pacnpocrpaHeHl{e' Hnx'
Boepacr Cu6upn' uufi roap (c sonu Ha.poceias falcifer u euure) Ceaepxofi Aon6acca' Kasraeau Toap Eepontl, xnufi Toap 3anannofi
pocrpoB I{3 HU)KHeno Toapa 6a9cefina Bualog-c6Opbl 39 Marepuan. 3 poclpa I€ BepxHeft qaclu Efixuero roapa u T.V.lfrpnnofi, T.l4.HanrggesoR c6opu T.H.HaasHneaofi, 2 pocrpa ns eepxnefi no6epexua Aga6apcKol-o saJIHBa 3 pocrpa qacTu Hr{)KHenoToapa Bocto.Igoro Tafivtlpa - c6oprt C.B.MeneguHoft' A'C'.[arnca' c6oprt rrHlKHenoToapa 6accefirra ouonona H3 BepxHefi qacru M e s o te u t h is l a p t i n sh a j a e V o r o n e z Ta6n. VI, Sr'rn.b\i
ra6n. Vll,
Mesoteuthis taptinshaje: BopoHeu, Lg62,
finn. 3 crp.97,
ra6n' 58, Qnr' 5'
5. UlM, IeHnrnpaa' KOnTIeKIII{q I o n o r u r1: Bopoueu, 1962, ra6n. 58, $ru'. roap' N 9209. fl-oe Yprorn*Tyruyc' cna6o cy6xonrrecKrfi' (tla-350-650), D,uarH o3. Pocrp cpeAuld, yAnuHeHHtrfi c oKpyl-neHHo-npsMoyToJlbllbtMnonepeqHblM ceqecxarufi c 6oxoe (EB-75-95), HHeM.XoporuoBbIpaX(eHbIcnHHHo-6oxoerlenp[rBeput,IH}Ible6oposart. Ta6nr.rua L2 Y oronez I,lsuepeuuq pocrpoa Me soteuthis taptinshajae
N e 8 7 -5 5 p.Kenuvap
flapauerpu
Ilnr.rua (npeanonaraeMaq o6ruag I v.tuttou JIeHHas Anuna fl ocJleanbeeonaPuofi
1 1 5 , O( 7 6 7 ) 1 0 5 , 7( 7 O 5 ) 85,3(s39)Q4)
9 7 , O( 8 O 8 ) 8 3 , 7( 6 9 6 ) . 6 3 , 9 ( s 3 3 )( O O )
. 1 5 , O (t o o )
1 2 , O (1 O O )
qACTH
Il.naverP cnuuno-6ProutHoft y BepIII,IHbI anbBeoIIbI !.uar',rerP
6oroeofi
Y BePiuI'IHbI
N e 87-56 p.Kennvnp
L2,4(83)
e,e(8b)
ANbBEOJIbI
Paar.ryc 6Prouruoft qacru Annua nPHBePuuHHoft a Auavrerp r clluHHo-6Plotuuofi 'racrll npl,tBepuuHuoft 6oxoeofi B nPHBePAlraverp urnunoft qacru B cnHHVron anbBeonsPnuft rPaA' nnocxocrl{r no-6prcruuoft Yron BePuHHHblft e cnuuHoI'PaA. 6prouruoft flnocKocrH'
40
5,5(37) 3 5, 1 (234) 11,5(77)(1OO) 10,3(69)(?)
27
32,5(271) 1O,O(83)(1OO) 8 , 4 ( 7 O) ( 8 4 )
20
BneuHHe n p l ' I a H a K H . I - l p u e e p r u a H H a t rq a c r b p o c r p a B b r r r r H y r a , c o c r a B n s e r oKono I/2 fla, eeprul'IHa 3aMerHo cMeuleHa K cnussoft cropoHe, 3aocrpeHa, BepurnHnulft yron HaxoAI'ITcg B rpeAenax 2O-290 (ra6n. 12). flpunepurrHHbre cnr.rHHo6oxoarle 6opoeArl qerKlre' nporrrr-r.rBatorcrr .qo r/2 fra. I4norqa na6nroAaercr{ rxrpuy ocrpl{fl. EoxoBble cropoHbl ynnouleubr, Ha Hax npocne)KaBarorcs xoBarocrb napuue 6oroBble noJlocblr cnuHHafi n 6prouHaa cropoHbr oAHHaKoBo BbrnyKnbre. flonepeqHoe ceqeHHe B eaAHeft tlacrn pocrpa 6luxe K oBanbHoMy, MeHee cxaro c 6oxoe (00 go-1oo), B nepeAHefi qacru craHoBr,rrcrr oKpyrneuHo-npsMoyr-onbHbrM, npr'r groM Bo3pacraer creneHu 60Koeoro clKarurr. BuyrpeHHI'Ie np l'I3HaK I'M oH ToneHe3. A n a e e o n a 3 a H r , r M a e ro K o n o r / 4 pocrpar 1/3 Beput{Ha ee cMelueHa x 6pnuuofi cropoue, oceBarr nvr.vrs. cnefKa H3orHyra H TaKxe cMelueHa x 6ptolLHoft cropoHe. Eprouruofi pagr.ryc y BeputrHbr aJlbBeonbl cocraBtuler oKono 35-450/" CE. AnueeonapHulft ynon B cnuHuo-6ppuHoft nnocKocrl'I paBeH 27o. Ha Ha'ranbHbrx craqaqx pa3Bnrurr pocrpbr r.rMe,gr ,,oqTI{ TaKyIG'xe' Sopvy KaK y Bspocnbtx xl'IBorHbrx, HecKonbKo MeHee BbrrrrHyrbl. 3HaqeHr4rr fla, Aocrrl.alourue 47 S npw C6 g vrv, Bo3pacraor Ao b 17 npu CE 12 vr,,a u qo 532 n p n C B 1 7 M M ( p o c r p J V sg 7 - S g , pHC. 13 ) . CreneHs 3aocrpeHHocru pocrpoB uao6opor - Bepur.rHHblft yron B cnr,rHHo-6prourHofi coKpauaercl rrnocKocr' MeHqercs or 22o npu A'aMerpe pocrpa g vv go 2so npr' o"lr"rp" 12 MM. Velz.eHr{r,rBocrs. 3Ha.{eHns [Ia MeHrrrorcrr B npeAerax or BSO qo 650, y 6onuuruHcrBa pocrpoe 4oo-sso. Taxxe H 3 M e H r { r { B b r3 H a r r e H r . r s E E - o r z b a o g 5 , 6oru'rueft qacrbro 8o-9o. Pocrpnt us oN,ronoHcKoro pafioHa xapaKTepHsyrorcrr 6dnulxI{MI'I 3HarreHuqun EE, veHauteft yAnl{HeHHocrbro (fla 35O-SOO), MeHee ocrpbrMrr BepuI{HHbIMLT JmnaMr{. BslAengercs pocrp NJbg7-S7, y Koropofo, cyAs no nono_ xeHHIo np?rBepurHHbrx cnnHuo_6oKoBbrx 6oposq, BepruuHa cMeureHa He K cnr,rHHofr, a x 6prouruofi cropoHe (.ra6n.VI,Sun.4). CpaBHeHHe.
OflHcrleaeurrfi
enq
sau6onee
cxoAeH no Sopve
( W e r n e r ,1 9 1 2 ) a e r o a p a H c M .c f . b e n e c h e( tS c h v r e s l e r , 1oos,
I{3
BepxHero
aaneHa
rcxuoft
qacrll
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Pocrpu
o6orx
85,2(6ss)(7s)
1 2 8 , O( 7 4 9 - ) 1 0 8 , O( 6 3 6 ) 91,O(SSZ)(71)
"rp
c M.raui
. 9 4 , r a 6 n .s g )
HasBaHgbrx
M 87-58 1 1 3 , O( 8 6 9 ) 94,9(730)
pocrpa
BHA'B
N e 87-59 p.Ileesrfi Kenou 1 0 8 , O( 6 8 8 ) 97,8(623) 70,7(+so)(66)
1 3 , O (1 O O )
L7,t( 1OO)
1 5 , 7 ( r O O)
1 1 , O( 8 5 )
1 5 , 3( 8 9 )
1 4 , B( 9 4 )
43,8 (337 ) 11,5(88)(1Oo)
7,7(42) 5 5 , O( S Z Z ) L4,O(AZ1(1OO)
5,5(35 ) 3 6 , ' 7( 2 3 4 ) 12,o(76)(100)
L3,4(78)(96)
12,O(76)(100)
s , ' 7( 7 s ) ( 8 4 )
o/r!
27 2L
29
41
P n c . 1 3 . l l p o g o n s u o e c e q e H H ep o c T p a M e s o t r t r l l t i s l a p t i n s k a j a e uttxHuft Toap' soga Dactylioceras commune' Voronez N 87-58, (x O,8) eaana AHa6apcrorii
BbIpa)KeHHyIo cy66Onee yAnI{HeHHble, I{MeIoT llyque BepuUHa He cMelqega K CnI{HHoft clopone. KOHI{qecKI{ro fupw, pocTpbl I{MeIoT HeKoTopbI€ npeAno SopMe cy6KoHl{qecKue Enusrfie A . o x y contLs (Zieten), A. graci liforni s p o A a A c r o c o e l i t e s cTaBHTenI{ I pocrernx (Janensch) ] , Bcex Ho y ,4.triscissrrs Kolb, lrl.striolata 6oposAa. PocTpbI poB ecTb npHBepuIIHHas 6ppunan (Phillips) 6onee ToHKI{e I{ cTpoftHble, uMeIoT Iryque BbIpa)KeHqacTblo CvTHOcItTenbHo funee Hy1g cy6KogllqecKyro $Opr',ry, 6onsurefi sstlil.striolata y pocTpoB 6oposgu BbITsHyTbr. flpunepruuHgbte paanu3ru pa)KeHbr sgar{uTenbHo cra6ee r{eM y M.laptinskajae. qus $UKCrpyIoTCq U B OHT\OI'eEege, BCneAcTBUe qerp Henl'3g pa6KaK IoHbIe 0opr"6r M.lalttirtskaiae. cMaTpr{BaTb M.striolata Tb. PocTpbr BcTpeqalorcrr Q auHanbH as npHypoqeHHoc H OCaIIKaX Bnn1gfiCrOtg neCqaHO-nnHHIlCTbtX KaK B MS5KOBOAHbIX
HOclITenLno
r'/ / i
Ana6apCr9ro
pafiOuoB,
TaI( I{ B npelMyIUeCTBeHHo
r.nugllcTblx
oCa[-
Kg)( OMorroHcKoro u OneHeKcKoro pafioHoB BMeqte c aMMoHIrfaMr{ BAaJIH or ropcKoro Marepr{Ka. I-lpu eroM B OMonoHcKoM pafione npeo6JlaAalor MenKIn€ 3KgeMnnrrpbr. CyAc no Bbrrrmyrocrl{ pogrpoB, xuBorubte Benu cBo6oAxonnaBatounft o6pas xusgu KaK B OTKpbIToM Mope, ra11 u B npg6pexg9fi 3OHe. pacnpoc'TpaHeHre. Huxuufiroap (soua u neorpaQuvecKoe Boapacr Harpoceras falcifer r{ Bbnre) - nuxHnfi aaneH CeaepHofi Cu6uptl. 14an, 20 pocrpoB us HrDKEeroroapa 6accefrna Bunrcg -c6opst T.H.Hant. Marep ngeaofi lr T.H. Kupuuofi, 5 pocrpoB Hs HHxHeno aaneHa p. KenuuRp - c6opbl T.l,l. Kupnuofi u T.H. HaruHseBofi, 3 pocrpa I{3 HI{)KHeno roapa AHa6apcrofo 38rII{Ba - c6opbt T.H. Harunaenofi, 10 pocrpoB rr3 Toapa HIDKHeTPaafleHa Ovononcxono pafioHa - c6oput A.C. fiarnca.
Mesote u th i s p yro mid a l i s (Z i e te n ) Ta6n. IV, $un, 5-7 p y : i i n s t e irn Z i e t e n ,1 8 3 0 ,S . 3 0 , T a f . 2 4 , F i g . 5 ; D u m o r t i e r , B e l e m n i t e s r a m i d a l i sM 1 8 6 4 - 1874,v ol. 4, p .3 6 . : l b , 1 9 4 2 ,S . 1 5 9 , l ' af. 10, Fi g. 8,9. A c r oc oelit espyra mi d a l i ' sKo Mes ot eut hisox y c o n a : Bo p o H e u ,1 9 6 2 , c rp . 95, Ta6n.60, $r' r.8. Zieten, 1830, ra6n.24, $rn.5. HnxHsc ropa px(Hofi qacru OPl. f onorun. Pocrp cpeAHufi, yN,repeHuoyqnuHeruuft (fla oxono 24O-38O), AuarHos. c oeanbublM nollepeqxblM cy6Kogr.recKufi, cxarslft c 6oroe (EE oxono 78-90), 6OpOeceqeHueM, co cpaBHuTeJrbHoKopoTKuMr{ [pI{Beptut{HHbtMI{CnHHHO-6OKOBbIMH AAMH. BHe uuue
tl. llpuneptunnnas qacrb pocrpa BbITnHyra, cocraBJlser npr{3HaK 3aocrpeHa, 6onee flonoBlrnbl fla, Beprr.jnHa cnenKa cMeuleHa K cnl{HHofi cropoue, 13). llpunepurraHnble cnruH>6oBenr{trr{Ha Bepur{Hnono yrna oxono 32-350(ta6n. nuulb B sallueft nonoB[{He npHBepfipocnexl{Batorcs KoBbre 6opoeasr nerny6orue, pocrpa oqenb nocreor npr{Beprunnnofi K cpeAuefi qacru fiepexoa ruunsofi qacrr{. yflfloueHbl, cItHHHaa u 6prour 3aMerno croponbl Eoxoesle neHxulfi, eABa saMerurrfi. no AnHHe pocrpa cxaruq Haq cropoHbt oArrHaKoBo BbrnyKJrbre. Crenerrb 6oxoeoro 3aMeTHO He MeHrreTcrr -
42
3HaqeHug
EB
u 66 cyuecTtseHHo
He pa3nlrtlaprcg.
Puc. 14. flpoaorrbuoe ceqenne pocrpa Mesoteuthis pyramidalis (Zieten) Ne 87-64, uuxxufi Toap, p. Burnoft(xO,gS)
BryrpeHHr{e npr{sHaKu r{ oHToneHes. ArrBeorra saxuMae,,r lreMgonlrM Menee nonoBr{Hbr AJrBHbrpocrpa (O,41_O,4S qmrgrl), Bepruu_ HA ee cnenKa.cMerqesa r 6prcuxofi cropoHe - 6prouxoft pannyc 42_4a% cE. Aru'neonnpnrrfi ypofl oKoJro 2B-92o. ocenag Jrr{Hus notrr[ rrpsMas, m{ub He3Haqr{T€JrbHonp[6mxeHa r 6proruxoft cropoHe (puc. 14). Ha HaqaJrbl{brx craal{ex pasBltrus pocrpbr l{Mercn cxoAlryp c BspocJlbrMu oco6svu Qopr"ry, JTBlubHecKoJrI'Ko 6orree saosrpenbr (y eepocJrbD( pocrpoB BepurrgHrrfi yron paBeH 32-370, y ronbrx - 22-260). 3Ha.renug lla no r{epepocra pocrpa saMe?Ho He MeHsrc/rcs. V sxs. M g7-64 npr cE 4 vu fla paeno 325, npn cE 8,3 uv fla - 33O. HsveHqI{Bocru. P o c r p s l h l . p y r a m i d a l i s ,x o r R B K o r r n e K u r f ir u c H a c q r { T b r B a € T cg 40 aK3. r,r3 pa3nbrx paftouoa Cu6rpu, orrrrqaprcs 6orgunM rrocrosgcrBoM npn_ sHaKoB. floqaeflspuree 6onsullHcrBo pocrpoB ( 7 5y") nrrreror 3uaqegtrfl fla s np"q* tle.x JZO-ggO u EE 7g-gO. lononsno nocrosnrra u odurac Qopr,aa pocrpoa. He orJllrtlaloTc'I no fiopvre u napaMerpaM r{ pocrpbr M.pyramidalis us ON,ronoxcroro paftoHa. CpanHenr.re. Paccvarpr{BaeMbre pocrpbr cxoAxbr c pocrpoM us nefiaca px_ Hoft .{acru 6orbsHe 6opoeAu. EproruHaa 6opoeaa y cn6npcKl{x aKseMnnspoB, KaK r{ y pocrpa, ongcaHgono LlureHoM, orcyrcrByer, xors psA nccneAoBar€neft (Werner, 1912; Naef, Lg223 Schwegler, 1969) oraocnnn K BI{Ay pyramidalis pocrpu c nprrBepuuggbrM}r 6prcgr HbrMr{6opoeqaun, T.e. np[FaAnex(auue K poAy Acrocoelites. pocrp, reo6paxex_ sufi H.C. Boponeu ( f gOZ ) Ha ra6n. 60, $nn. 8 noA HaoBaHEeMA.oxycona 11g Toapa AHa6apcxofi ry6rr, Toxer cyAs no orcyrcrBuro 6poruuofi 6opoeArr, orrrocrncrl K onl{cblBaeMoMy BI{Ay. oau uanbHas npo'ypoqeHHocrs. Bonbun'crBo pocrpoB np,,]poqe'o K npn6pexHblM cpaBHI{T€nbHo MerIKoBoAHbrMnecqaxo.nrrartrcrbrM ocaAKaM Bnnofi_ cKono u Ana6apo-Xararu.cKoro pafiouon. Oqnaro oKorro rpertr EaxoAoK cBssarro c $auu*r'lH orKpblToro Mops IIeHo-OneHeKcKono u O*rononcgoro pafiogoB. Moxuo rpeAronararbr qro xors Mesoteuthis pyramid.alis npeAnoqrran npn6pexuo-MerlKoBoAHbre o6cranoaxu, oH HepeAKo aanJrbtBan E B orKpbrroe Mope. Bospacr tI neonpaOlaqecKoe pacnpocrpaHeHEe. Bepxnar qacrb HrDr(Heno roapa (c soHrl Dactylioceras commune r{ BbruIe) nrrcrufi aanen Ceaepnofi Cu6npu n 3anaqxoft Eaponu. Marepl{an.7 pocrpoB I{3 Bepxnefi qacrtr Hrr)KHeromapa 6accefina Bnnog c6oprr T'H. Knpusofi n T.H. HansHgesoft, 14 pocrpoB r{s Bepxnefi qacrg H11xgero Toapa no6epexsa Ana6apcxofi r;6u H n-oBa Yponr-Twyc - c6opu T.H. Hanbngeeoft, 8 pocrpoB I{3 Toapa p.Kennurep - c6opsl T.I4. Krpnnofi n T.H. Halrureaofi, 8 pocrpoB I{3 BepxHeno roapa - HtrxHeno aanega Or,rononcroro pafio'a c6oprr A.C. Aannca.
43
Ta6 nHua 13 ( Zieten) Hevrepenus PocrPoB M e s o t e u t h i s p y r a m id a l i s
87'6O, Kenr.Irunp
Ns 87-61, p. Kenuunp
66,0(550) 54,O(45O) 3g,4(SzO)(58)
7 6 , 0 ( 5 O 3) 64,0 G24) 4O,O(26s)(s3)
flapavterprt
Anuua InpeanonaraeMan o6ruag ly"tuno"neHHasI Annna nocJleanbneonnPnoft qACTI,I
finar"rerp cnuuno-6Prouuoft Bepu[IHbl
aflbBeonbl
firaaverp
6oxoaoft
Y
Y BeptullHbl
L 2 , O( 1 O O)
1 5 , 1 ( 1 O O)
9,O(75)
11,4(76)
ANbBEOIIbI
Paanyc AnuHa
6,5(43) 32,4(215) r4,4(95)(1OO)
6Proutuofi npI{BePIUHnHoft qacru
e fiuarvrerp cnuuno-6PrcruHoft qacru npl{Bepununofi Anar.aerp 6oxoeoft B nPHBePruungoft qacrll
2O,O(167) 9,6(80)(1OO)
1O,8(72)(76)
7,7(O+)(80)
Vron anbBeoJlsPutlfi B cnHHHonPaA. 6plouxofi nnocKocrl{r Vnon BepuuHHblft e cnllHHofPaA. 6prournoft' nnocKocru,
35
L ' l e s oet u t h i s p s e u d o el o n g a t aS a c h s r t p ' n o u '
37
I
Ta6n. VIII, Srr. 1-3 T-"j.24, Fig' +,5,8' B e l e m n i t e s p a x i l l o s u s a m a l t h e i : Q u e n s t e d t ,1 8 4 9 ,S . 4 0 1 , { 7 ' f i g ' p l ' 17 7 , p ' 50, B e l e m n i t e s e l o n g a t u s :P h i l l i p s , 1 8 6 5 - l B 7 l , Abb.29 S . l 3 B , 1 9 6 2 , u " . . S c h w e g l e r , C : p a * i i l o " u s i . e . S . Belemntte" r a 6 n . 39, {nr. 1. Misoteuthis paxillosus: Hyry6raee, 1966, crp. 163, M a7-66, Myseft Hlf CO AH CCCP, Hoeocu6apcx. Imcert6ypn, lonorun eepxur.rfinnnHc6axr. 3oHa Pleuroceras spinatum: Eerrev6ypr, Pocrp cpeguufi, cy6ronuvecruft, yMepeuHo yAn]rueuuuft (eua'reuug rla Iua.rHoo. c oxpyrnbrM r{Jrucna6o c)KaTbrMc 6oKoB nonepetrHblM cerreHl,IeM oKoJro 35O-.450), Ilprmepulrr{Hbre cmnnro-'6oKoBble 6opoensl qerKue, Ho KoporKI'Ie. (BE oxono 88-98). B H e g r H l r e n p H 3 H a K p r . f l p u n e p r u u H u a aq a c r b p o c r p a c o c r a B n 9 e r o K o I I o n o n o saullMaer ergrr fla (ra6n. i+). Bepur{Ha 3aocrpeHa (eepurunuslft yron 34-360), 6nueroe K ueHTpanbHoMy nonoxeHl{e. Y eepiuunbl qerKo BbtAenslorcg npl'IBepuHHHbre cnr{HHo-6oroesre 6opoeAul, Koropbre nporarllBatorcn nuub Ha t/ 3 npHBepuLIHHoft .{acru pocrpa. flonepeqHoe ceqeHne pocrpa oKpyl'noer cJIerKa cxeroe c 6oKoe. Bce qerbrpe cropoHbr qBJrnrorcn BbInyKJrbIMu,6oxoaue B HecKonbKo l,lenuLuefi creneHn qeM cn[HHaa u 6Ptouuaa. n p u 3 H a K H r r o H T o r e H e 3 . A n s e e o n ac o c r a B n s e r o K o n o BxyrpeHHlre pocrpa, Bepur{Ha ee F\depeHHo cMeueHa K 6ptotuHoft cropoHe^ Anr.rHbr 3340"/" (6prourHofipaquyc 32-4O7o CE). Anueeonspuuft ynon cocraBrlser oKorlo 31". Ocenas nv1fis npu6nuxeHa K 6prouruovy Kparo, cnerKa Ll3orHyra (puc.15). Ha HatranbHbrxcraAgfrx pocrp l,rMeer orqernuByro cy6rcoul{qecK}'IoSopt"ty, 6onee :aI
HageaHLIe
44
B l r . a au o c x o A c r B y C a t a t e u t h i s e l o n g a t d ( M i l l . )
N/-b87-63, Aua6apcxaa ry6a
Ne87-62, AHa6apcraa ry6a
N-b 8 7 - 6 4 , p . Banroft
6 7 , O( 5 8 3 ) 5 4 , O( 4 7 O ) 37,8(329)(s6)
'7 2,o (497 ) 6 3 , O( 4 3 2 ) 42,8(295)(s6)
1 1 , 5 (1 O O )
14,5(1OO)
8,3 ( 100 )
IO,2 ( 89 )
1 1 , O( 7 6 )
7,7(93)
5,5 (48 ) 2 r,3 ( 185 ) 1 0 , 1 ( 8 8) ( 1 0 0 )
2 ' 7 , O( 1 8 6 ) 12,8(88)(1OO)
3,5(42) 1 8 , O( 2 1 7 ) 6,9(83)(100)
8,O(70)(79)
9,6(66)(7s)
6,3(76)(91)
2a 32
4 5 , O( 5 4 2 ) 3 3 , O( 3 9 8 ) 2 7 , 4 ( 3 3 O) ( 6 0 )
32 34
26
ocrpeH qeM y B3pocrlblx oco6efi, ornllqaercg MeHbuleft orsocnrengnoft yAnHHeHHGcrbr'o. Y axs. i\lb g7-69 np' quaverpe 7 vv fla paBHa 314, npr quaverpe u r ' r 12,5 I-la 35o, nppr Ar.raMerpe 16,g MM 3Har{eHue l-la gg4. cDopva pocrpa Hs cy6xourqecroft craHoBarcq c)6uHn'rqpn'{ecxoft, ceoftcreeuHoft espocJrbrM xHBorHbrM, npa AHaMerpax 1O_12 vv. H s N a e H q I ' I Bo c r b . P o c r p s l l v l . p s e t t d o e l o r t g a t a H 3 eBponeficroro nauuc6axa u cu6upcxofo roapa cyuecrBeHHo Apl.n or qpl.t.a He orJrr{rra,orcs. B o6eax npynnax pocrpoB ecrb 3KaeMnJlgpbl' clKarble c 6oros u c noqru oKpyfnbrM ceqeHgeM EE eapaapyer B npeqenax 94-1oo, y 6onsLuuHcrBa pocrpoB 90_96. pocrpsl c pa3nHr{Hofi creneubro yAnllHeHHocra (l-la 3oo-4ao) acrpevaprcs raKxe e o6eax Fpyfinax, npeo6naqapu]'IMl{ flBJrsrorcl 3uaqeunrr paeHslvu [Ia, g4O-44O. y HercoTopblx pocrpoB c MeHbult{ME qeM y qpyrHx quaMerpaMH, no-BHAHMOMy npr{HaAnts xaurnx eue roHbrM xHBorHbtM, o6uras $oprura ,IBJr.rrercfl nepexoAHofr or cy6xoHr,rec_ xoii x cy6unnapqpuuecxoii. cpae He H He. PaccvarpHBaeMbre pocrpbr HrrrreM cyqecrBeHHbrM He ornr.rqarorL c g o r p o c r p o B ' o n P I c a H H b I xn o q H a 3 B a H I ' I e M Bclentnites elortgattts @unnancov (Phillip's, 186s-1871) ue nnwuc6axa Arn nnn H KeeHUreAroM (euenstedt, 1g49) H lljeernepoM(Sch*egler, 1962) ae nnrHc6axa rcxnoft qacrrd opl. f l o q o 6 nuft )Ke pocrp r.r3 nlrr.rHc6axa CeeepHoro Kasxasa npuBoqarcq K.tlj. Hyuy6raqee (1g66) n o q H a 3 B a H H e M ' l l e ' s o lt u t l t i s p a x i l l o s u ' s . cne,ryer yKasarb, r{To H BHrrpeHHee crp}e H H e p o c r p o B t ' t c n a u e f t K o J I n e K u l ' I r ' r( p a c . 15) coorBercrByer nso6paxeHurrM KeeHureqra (Quensredr,
r a 6 n .2 4 , g v r .. g ) r . ro u n n H n c a( p l , i i l i ; r ; . i o . r r , A g a r p 1. I ) . P o . s s a l o t e r r l / er il.osn g a r a ( I I i l l e i i,e Z O , c r p . 6 0 , ra6n.7, $rlr,6_g) H3 nnutc6axa
AHrnan I'IMeer pocrpr naruegusliir cyArr no uso6pax{eHqrrM, npHBepur'HHbrx cnHHHo6oxoar,ix 6opoeq, a He Moxer 6srrs o6se.luner{ c onacsleaevoft Qopvofr B oArn BII! a taxe poA. 3ro :racraBwJro Hac BbrtenaTb HoBslft eu,1 _ ,\!esotautltis pse,_ doelortgata. Enusxair 3analHceeponeficxair n e l - r a c o e s r i i e n , f _ , \ l . i , u l . q a r i . s( \ oune e_t
45
Ta6fluua
L4
Hsr.,repeuus pocrpoB
M e s o t e u t h i s p s e u d o e l o n g a t aS a c h s , s p . n o v .
IS 87-66, JIrorccev6Ypr
flapauerpu
Annsa fnpeanonaraeMas o6urag fV"t"to"neHual AnnHa nocflearlbaeonePnoft qacTn fiuar"rerp cnunno-6Ptoutttofi Y Bepu[HbI
Y Bepur-
AIIbBEOJIbI
Paqryc 6proutnoft Anusa npl{BePutlnnofi qacrtr a finar'rerp cnrnse6Pounofi npr{BepuuHxofi 'racru fiuaverp 6oKoeoft B nPHBePuunuofi qacrl{ Yron aJlbBeoJlspuulfi B cnI{HHo-6prcuuofi nnocrocrl{r nPaA. Vnon Bepul{Hublfi e cn[HHo6poruHoft nrrocKocrur nPaA.
12O,O(695) 99,5(575) ?o,5(4O8)(59)
L 1 , 6 ( 1 O O)
17,3 ( tOO)
L6,1( 95 )
1 4 1 9( 8 7 )
anbBeoJlbl
fluar'aerp 6oroaoft HbI
t22,5(696 ) 122,5(696 ) 17,4(440)(62)
Ir 87-67, Kapnanrt
-
3 9 , Oe 2 2 ) ls,o(8s)(too) 14,8(84) (99 )
36
6,8(39) 34,3( 198) 13,9(AO)(100) 12,O(69)86)
3,4
Oxontranue ra6nnurt }ite 87-68' p. Ieaufi Keton'
fiapauerpst
Anmra InpeanonanaeMas o6uac f v*.to"neHHas Anuua nocneanbaeonnPnofi qacru Auar',rerp cnunno-6protunofi Y aepftuxur aJrbBeonbl Auar"rerp 6oxosofi Y BepuuHbr anbBeonbl Paaryc 6pnunoft nnmra npuBepul{Hnofi qaclu Auar'recp cnxnno-6ptouuofi a flpr{BepuuHnofi qactu fl,uaverp 6oxoeofi B nPHBePurusxofi qacru YnOn
anbBeOnsPHbIH
1 O 9 , O( 6 0 6 ) 1 0 1 , O( 5 6 1 ) 13,2(+OO)(67)
1O5,O(625) 9 5 , O( 5 6 6 ) 64,6(384)(62)
t8,o ( lOO)
t 6 , d (t o o )
16,0 (89 )
L4,1 (88 )
7,O(39) 31,6( 176) 15,2(84)(100)
5,4(32 ) 27,O(161) 13,1(78)(too)
14,O(78)(92)
L2,Lfi 2\ (92 )
B CIIHH-
uo-6prourHoft nnocxocrl{, rpaa. Yron BepuJI{HHbtfie cnuBHo6prcurnofi nnocKocrl{, rPaa.
l $ 87-69, p. fleaufi Kenot
!)I
34
uueer MeHee yAn[Herrxslfi cnenra npr{Tyrnexxuff pocrp. Pacnpoctpallind, 1828) (cr,r. suxe) rarxe ornlrraercs no vensurefi HerHrrfi e Cu6upu M.inornata (Phill.) yAnHHeHHocrH, cxarulo c 6oxoa n cy6roxrnrecrofi Qopue pocrpa. B C r 6 u p r 6 o r u u l } r n c r B op o c r p o B H a f i A e n o npHypor{eHHocrs. OauuanbHas B npeuMyrlecrBeHno rnnHucrbrx ocaAKax orKpblToro Mops e Ol,ronoxcKoM paftoue. 46
PHc.
l-5'
tlpogoJrbHoe ceqeHr{e pocrpa
sachssp. nov.
/.ilesotetttrtis psettdoelongata
t$ 8z-69, Hnxnrftr@pr so'a zug;u";iiri"l"filunia_
nus, p. Ileaorfi KeAos ( x Org)
K neocnHrnr{HaJrbnbrM nrr{HrrcTbrM ocalKaM npnyporleHbr u HaxoAKu M'pseudoelongatan Kapnararc. pexe pocrpbr M.pseud"oerongata BcTpe_ qalorcs n npn6pexnbrx MenroBo[Hbrx @aaxa:< Bnnpficrono pafioua. Craeaulroe aaer ocHoBaHHe AyMarIr, r{To xl{BorHbre B ocHoBHoM B€nH cso6qa'onraB€uoulufi o6pas xr{g'r B cnKpbrroM Mope, xorrl H oarrnbr_ B{rnu B npu6pexnrle pafioHu. BoepacT H feonpasnuecKoe pacnpocrpaHeHHe. flnuuc6a:r 3anaanoft Enponu, Kapnar r,r Kaaraea, nuxuufi Toap (eona llar_ poceras falci'fer n arrue) Cn6upr.r. M arep[a[. B pocrpa r{s Bepx'ero nnr{Hc6axa .trorcev6ypra c6opnr B.H. Carca, 6 poctpoB us Hr{xueno nnnrc6a;ra Kapnar _ c6opu A.c. Aanng"r_ 1o pocrpoB Hs Hr.lxHero Toapa or.aonoscrono pafioHa - c60pu A.c. Aarnca, 6_ pocrpoB r{3 Hr{xgeno roapa Bm'oficK.n. ,c6opu T.H. HalrrHsesofi u T.V. Knpuuofi.
paftoua -
Me s oteu tl ti s i n o rn a ta(p h i l l i p s ) Ta6n. Xlll, Qun.S-7 Belenmites compressus: Blainville, lg25, p. 84, pl. 2, fig. 9; Quenstedr, t}46_l}4g, 5.424, Tab. 27, fig. 4. Belemnites inornatus:Phillips,]s^6sr-l8il, ?. -s-0,!1.lg, fig. 46; Denecke, lB9B, s.42,Taf.3,Fig.6; Janensch, ig02,
Fig.6.
s. ll2, i^!r. n,' b"";;l;;"i905, s. 290,Taf.26,
Be lem nit esr hena n u s W : e rn e r, \? 1 2 , S. l 3 Z , T a f. 13, Fi g. l . Mesoteuthis cf. inornata:.Kolb, 1942, s. 163; r"r. o,i,g.'ir,'rz. Be lem nit esr hena n u si .e .S .: S c h w e g l e r, 1 9 6 5 ,S. gi , ,IU U .-S S . Mesoteuthisrhenana: Hyrry6naee,1966, crp. 1Sg, Ta6n.Bg, Qrn.7. Mesoteuthisinornata: caxc u Harunaeaa, rg,6-7-2, ra6n. 1, 0rr. g, Mocxaneu_ Ko r 1 9 6 8, c r p. 30, T a 6 n .S, S rr.2 . Jl e r r or nn, ne6p a H n u ftB.H . c l x c o v :p h i l l i p s , 1g6s_1g21, ra6n. go, Q*' 46 l" flef,ac Anrn*r (Bnroerx). Mecron.-*l*o.r"e ne*rorr{ra rrer{sBecr'o. Aral.Iros' Pocrp cpeqxuft, yN{epeHHO yAn[HeHHbrfi, cna6o c]6KoHr.rqecK[fi, cxarufi c 6oros (BE oxono 7o-9o), c osanbHbrM noneperrHbrM ceqeHr{eM.Bepunna crlenKa cMeuleHaK cIIl{HHoftcropone. I-lpuaepult{Hubre cnngxo-6o6oBbre6oposnrr pee-
KI{e, HO KOpoTI(ne. BxeuHHe IIpI{3HaKI{. flpuaepruuHuafl qacrb sanr{Maer Meuee nonoBr{nu lla, flepexoA or Hee K cpeAHefi qacrE pocrpa oqenb rocrenernuft. BepunHa cna6o cMeueHa K cn''H'oft cropone, y 6onsuuHcrBa pocrpoB npr.rryflne'a, Ber.rtrrrr{HaBeplulrlr'ono ]tn''a xone6nercs B nPeAenax (r.6rr. rs)." npn""p*rrHHbre 34-45D cnEHHo-6oroerte 6oposqbl qerKEer ysK[er rrpocnextrBatorcs ua 2/3 flpnBepu'snoft qacL Tlr pocrpa. Eorosbte cropoHbr cJrerra yfinouenbl, Ha HEx BqAHbr Irgpfrbre 6oro_. BbIe nonocu' cnrs'as ,n 6poruxae cropo'br pocTpa oo"r.rioil"I",rry**le. cTenens 6oxosoro cr€TEs [osrrr He MeHsercs no acef, AnEHe pocrpa (e'aqeHas BE n 66 nparcrnqecr[ oA[narosrle ). BnyrpeHH,'e ''pr{gHaK,n u oHToreHe3. A t t b a e o J u sr a ' , , M a e r o K o n o , , o noBHIrbI ArInHbl pocrpa, BepurHa ee saMerHo cMelqena x 6poruHofi cropoge ( 6prour Hoft paqryc 30-38% cE). Ansseonapnufi yrorr usMepsercs, 2s-2g": o.;;';;ngg npr6'Irxe'a r 6poruxofi cropoae E saMerHo rrsonHJrra. Ha Ha.ra[b'brx cra.n,nsx pocrpbr nMelor 60rce scHo BbrpaxeHEyo cy6roHr+recK)rc o0prry (prc. t6), ;"_
47
Ta6nr.rua 15 I4suepeuul pocrpoB ltle sotc til lti.s irxtnrata ( Phi lli ps) Ns 87-7 1, p.Ilenurft Keqon
flapaverpu npeAnonaraeMag
Innua o6uras Anuua
VCTAHOBNEHHA'I
nocneanbeeonaPHoft
8 9 , O( 6 0 1 ) 8g,o (60 1) 45,5(SOZx51)
Nb 85-17, p. Ileesrfi Keaon
8 0 , o( o o z) 77,O(579) 37,5(282) (47 )
qACTH
firaaverp cnuuHo-6Prou:Hoft Y
14,8(1OO)
13,3 ( 1OO)
anbBeonF,I
BepruHHbl
ftuar'aerp 6oxoeoft Y BePuwHL'l
1 2 , O( 9 0 )
L2,7(86 )
anbBeonbl
Paauyc Anuna
6prouruoft npl{BepuwuHofi
qacrPr
a cnusno-6PtoulHofi firaaverp nprrBepuuunoft qacru Auar'aerp 6oxoeofi B nPI4BePuruuHofi t{acrl,l B cnHHrPaA. uo-6prcr-uHofi nnocrocrl'Ir Yron BepuunHblfi e cnuHHo6promttoft nnocKocrur fPaA.
4,4(3O) 2 1 , 5( 1 4 5) 11,6(78)(100)
5,1(38) 1 9 , 5( r 4 7 ) 1L,7(88)(1OO)
1O,O(68)(86)
1O,4(78)(89) 2a
anbBeorsPnulft
Yron
Oxon.{ague
36
-
3
4
'ta6nuuu
flapauerptr
Ne 87-7 2, p. Kenuvap
Ne87-73, Ana6apcxan ry6a
9 8 , O( S S S) 93,5 (557 ) s 7 , 7 ( S + g) ( 5 9 )
8 8 , O( 5 6 1 ) 68,O(433) 4s,o (287 ) (51 )
Araaverp cnllHHo-6prcunrofi y Bepul{Hbr arrbBeonbr Auaverp 6oxoeoli y BepuI{HbI aflbBeonbI Paqrryc 6prouruofi Annra npr{Bepuunsoft qacru fluaverp cil]ruHo-6prcurHoft n npl{Bepuusnoft qasrl{
16,8(1OO)
1 5 , 7 ( 1 O O)
L 4 , 4 (8 7 )
t3,4( 85 )
26,stioo I
6,2(33 ) 2 L , 7( 1 3 8)
L4,O(87)(100)
1-2,9(ez1(1oo)
6oxoeoft B npl{Bep[uauetp urungofi qacru Yron anbBeonspnufi B cIItrHno-6prcuruofi nnocrocrll, rpaa. Yron Bepul{HHbtft s cnuHHo6prcurHofi nnocKocrll, rpaA.
1 l - , 7( 7 0 ) ( 8 4 )
1 1 , 3 ( 8 1 )( 8 7 )
Annra I npeanonafaeMal o6uaq fv.tunouneHHaa AnuHa nocneanbeeonnpuoft qACTI,I
25 45
43
flpu yArII{HeHbr rreM B3poctlbte. Y axs. Ne 85-17 oruocuTenbHo nplt r"ta 35O, al{afla 8 npn ALIaMerpe Ar.raMerpe 5 eHa.reHue fla paBHo 37O, p r u 2 a 2 . f l a Merpe 1313 Pocrpst rlf.inornata ornl{qalorcn cyuecreeHHoft HsMeHrrl'IBoHa1,aeHqHBocru. npnqeM npeo6nagator Bentaql'Iubl nopsAKa ao 38O), crbro eHaqeHr.rfi fla (or 220 p a s 6 p o c n a 6 r n o A a e r c n I{ B 3Harrenusx EE (or 7O no 92). 28O-32O. Boncruofi cKonbKo
48
6onee
f ".?' 16. llpoaoJrb'oeceqe'ue pocTpa Mesoteu,tis inornata(pl rilIrps/. Ns gs_17, ,"po"n ;;;p, p. /learrft KesoH
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3aecs MaKcr'rMvM (e npeaenax76-84) BbrpaxeH Me'ee qerxo. Me-
Ut /ll
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::liff= .':ffi,,i",1#i.:Hi:"; lluIII "n.".;?:t.l#;;::i:*:lg_"1
o*ono 50 poclpoB aa'Hoto BuAa' He yraercs noAMe?r{T6 6o eaxouoMepHocTl'I Ka*r.r" B r'rsMeHeHr.rfl pocrpoB, cBffsaHHbrec "]"rx ""*o_-
"^:T:Y .H,:.:Tffff#I' ;:,:i*i::*t*;. H H cM ; ;;
a TaKXe no o6ruefi *opve He ornuqarorcs on cu6upcrru( *opr"r.
tllf llt Illllll
Ilwl I
W / tvt.rhenana(opper), ,.!t
:T::"::,JTI :k::'x:^.:iy:*,'*'
,i, c KoropbrM eno qacro "L?_ o6reAuusror. M. rhenana';J:";";:;:JT:1"" scHo BbrpaxeuHrrfi cy6ronuqecxufir rrp"Onn*arunrficg x cy6qnuHApr{_ 'recrorr 00pve, oTHOCU?eJrbHo 6qnee yAnHHeEHbrft (euaveune 4oo)' na o*oflo Te x(e npxo'ar(H o'nl{qarcrr M.inornata ' i " " i n ootr r o trvl.pseudoerongata a lueernep (schwegler, sp. nov. rg6s)'oor"onrr"r . BW Beremnites vortzii
nosopvepocrpa. Hl:ii:;,.t3,.,?,",l.r:!lii::*:r{re,rb'o cxoAHun oqxaxo pocrp nprBepu'''*)4o
6poruHyo 6opoe,qy n xeH orHocr.rrbcs K poAy Arroro'ftrfr.t"t AonPocrp fil'inornat.' onrcaHxuft 3.4. MocxaneHxo ( 1g6g ca? ) flpnavlpan, ) uo cpq[Hef, ropu (6afioHMeer npn6nuxapu)rocs K cytuan[HApr{qecxofi Qopr,,rJr.Iloao6;_
;;-;*F-.'Ji,ffir' ":'#"ff JiHHr*Tffii oi"'or* tiz-24' roaror,av ""rrwr.nJ
3.A. MocxaneHKo.
Cpearn ca6npcxux
LrrrucblBaeMoro
BI{Aa Qopr"ry, ollncagg]rlo
BaAoe 6nt,Isxrfc rrahor,^hh
:",:*j j;;;;:;;t'r;,:,h:,$#;-":?:H;:ilH#T."#n'i.ilftr,"IH OaunanbHas irpuypoqeHHocrs. pocrpbr ofirrcbrBaeMoro rroAaBlnrtoul€M 6onsurlrHcrBe B [ r A ac o 6 p a x u a B nnr{Hr{-crbrx oca.q*r* or*pbrrono Mopg Toap€FaarleEa oraonoxcxono r rle'o'one""*.*o.o paftoHoa E e 6af,oce paftona. EqaHrrrrote pocrpbl Axa6apo-Xaranr.Kono HaftAenu B cpaBHrrr€Jlbno MemcoBoAHbrx necqago_tJr[_ HI{crbIx ocaAKax Toapa nnaaJreHa Anaoapo-iato"*o"o pafiona. cosepue'Ho cyrcrBy'T HaxoAKa M'inornata s orL Bnnoncrot-o"oor"r r-A€ pacnoflararcs Meru(oBoAHbrri sanr{B 'opc*ono Mops. HcxoAs ;;;J; ""
ff ,lt"T.XJ,i.il];;I llff[Tffi il:rifii"H-'"oooo'o.,*"'o*o*?"J.:ff
Bospacr r ' r n. e o n p a { n v e c K o e pac,,pocrpaHeHne. Hr{x'ero roapa (eoHa B e p x H s sq a c r b zugodactyrites ur*"i"",irr _ ;;; ToKa, aepxHafi Toap _ narr
yprom-rn ;5:?,;f:n::-:"9:fr:t*fl'ffi.",il;ilffi ." r-oBa i. _;;;;;' Me soteuthi s ae qualis Voronez Ta6n. VIII, gar.4_7 Me so t eut hisaequal i s : Bo p o u e u , 1 9 6 2 , c rp . g 6 , ra6aS g, $rn.217. IIe x r o r r.rrr, eu6paHnrrfi B.H.Caxcou: Boporreu, 1g62, .IleHaru-paq, KoJrn. ]S g2og.
ll-os
4 4 9
Ypronr-Tlnvryc, T@p.
ra6[. Sg, Srn. 7. UI.M,
49
Ta6nsira t6 Her'repenus PocrPoB M e sot euthis ae qualis Voronez I$ 8 7 -7 5 , p. Bunoft
flapauetpu
Annna fnPeanonaraeMas o6ruas lV"t""otIIeHHae Annna noc[eanbaeonsPnoft {,acTu llnar.rerp cnurnro-6Prouuofi BepUlllHbt
AJII'BeOJIbI
Anar*erp
6oroaofi
IIbI
8 2 , O( 6 4 6 ) ?1,5(563) 58,8(463) fi2\ Y
Y BePurH-
AJIbBEO'IbT
PaAryc 6Potuxoft qacru llnrna rrprrBeputrrnofi !,narvrerp cnnnxo-6Pouruofi a rrptrBepurrnnofi qacrlr 6orosofi B nPuBePllrauerp urrorofi qacrl{ Vron arlltBeonsPnufi B cnI{Hno-6protuxofi nnocrocrHr rPen. Vron Bepurl{HHbtfte cntrrgo6prcunofi nnocKocrl{' rpafl.
L2,7 ( lOO ) 10,5 (84 ) 4 , O( 3 1 ) 31,8(250) L z , O (9 4 ) ( 1 0 0 ) 1O,O(79)(83)
2A
l t 87-76, p. Bunofi 81,On L7') 78,8(697) 60,8(Sge)(75)
1 r , 3 (r o o ) 8,5(75)
26,0(23O)
1 1 , 3 (r o o ) ( 1 o o ) 9 , 1 (8 1 ) ( 8 1 )
32
Pocrp cpeannfi yMepeuxo yAnEHeHHuft, npu6nuxaloueftcs x cy6uuAnarHos. ngnAp1aqecrofi {opMbt, cxaTufi c 6oxoe (eentrrruHa 6E oKono 1+9O ) c 6nugxul"I K oBanbgoMy rronepeqHbrMceqeHueM, c KoporKt{Mu (orono L/4 lla) npnnepurunnbrMl{ cnunno{oKoBblMl{ 6oposnauu. npn3HaK r{. IlpueepurnHHastracrb pocrpa BbITsHyra, cocraBnger BHe uuue 26HecKoJrbKoMeHee L/2 f|a, BepuHHa 3aocTpeHa' BepuHHHblft.yt.on B npeAenax g2o (ra6n, 16). BepruuHa 3a6r{Maer 6nnsroe K ueHTparbHoMy ronoxeHl{e u Kaqactu nepxunfi xeTcs cnerKa BsAepnyToft 6naroAaps ToMyr r{To B npuBepuuHHofi flpueutnyrcnslft' Hao6opor xpafi Kpafi pocrpa eABa 3aMeTHo BorHyr' a 6pouuofi [pe3a He BbD(qrsr Ho Koporruer ray6orue, Eepuugnbte cmnrso-6oKoBbre 6opoensl qacrlr. HHorga e5n}ren BepuIHHbl ua6rnoqagaAgefi npuBepuurnrofi nonoBr{Hbr Aerlbt eTCs rxTpru(OBaTocTb.EoxOssle cTopogbt ynnoqeHblr Ha HHX BblAe[spTcn napHble 6orosule nonocbt. Cnnngas u 6pourHarr cropoHbr oAHHaKoBo Bbln)'KJlbre. flonepevpocrpa. Hoe ceqexr{e cxaro c 6oroe 6onee unil MeHee paBHoMepuo no acefi AnI{He o K o n oL/33 a H H M a e r rrpH3HaKu H oHToreHe3. Anseeona BnyrpeguHe (6prourctopoHe r 6protuHofi cMeueHa 3aMerHo t/S qngxbr pocrpa, BepuuHa ee yron oKorto 3Oo. Oceeag m{Hutl npuxofi pamryc ogoJro O,3 CE). Aruneonnpxrfi pa3: 6mrxega r 5prorunofi cropoxe, cfienKa usorrryTa. Ha HaqaJrbHblx crarl{flx cy6xoHuIaMeIoT pocTpbl (puc. t7 | MM BuTtrs rtpu Aua]vreTpe oKoJIo 4-5 qecK]ro 6opr"ry r{ Hr{qeM c}rulecrBexHbrM He orrl}rtratorcs or Apyl'ru( Bt{rloB ltlesotexapaxrepuofi aJIlI pacg uthis.
50
Ne 87-77, p. Bnnrcft
N e 8 7 -7 8 , p. Brnofi
60,0 (674) 56,0(629) 47,5(s37)(79)
N s 87-79, p. Trorn
er,oJozz ) 65,O(5OO)
61,5(4O2) 6L,5(4C21
8,9 ( lOO )
1 3 , O( r o o )
12,8( lOO)
7,1(80)
1 O , O( 7 7 )
1 1 , 8 (S e I
22,3(251) 8,4(94)(100)
7,r(80)(8s)
13,O(roo)(roo)
3,5(28) 2 7 , 2 ( 2 1 3) 1 2 , o ( 9 - 4 )( 1 O O)
10,1(78)(78)
1O,7(84)(89)
36,2(27 g',)
30
26
25
27
Aanr B Hatul'IJ(c6opax pocrpbl cy6uununap*ecKr{er. t{To H orpaxeHo B npeAnanaeMOM Ar{ar*o3e Br'rAa. B u[po*r,rx npeAenax BapbHpFOT 3HaqeHug [Ia (or 320 ao 650) u EE (or 70 qo g6). Ecnu HcKnroqnrb peAxue orKnoHeHas, ocHoBH€rs Macca pocrpoB HMeer sHaqeHurr [Ia a rpeqenax 940_560, E; _ 74_gO, llpu sTOM' KAK BI,IAHO HA PHC. 18,A,
KPI,TBA9 H3MEHvI,TBOCTA
I-IA,
11OCTPOETTHAA
HA MArepaane no J.2o pocrparr,r, HMeer ABa MaKcr.rMyrva(34O_42o r.r soo_56o) c 3aMerHbrM cnanoM qacrorbr BcrpeqaeMocr' sopv c f'a, paBHbrM420-500. [pu_ t{HHbr raxoio xoAa xpaeoft ocrarorcff HerrcHbrMa.Eonee yArrHHeHHbre n MeHee yA_
l(on-60
60 70 80 90 /00 300 56 P u c. 17. Ne 87-79,
500
700 lla
flpogo.flbHoe ceqeHHe pocrpa trlesotettthis aeqttalis Voronez nNxHr,ri Toap, soHa Dactilioceras commune,p.Trour (x orTs)
Puc. 18. HgveH.{uaocrb pocrpoB ,lles ot e ut h i.s ae q uali.s Voronez A - no orHocr.rrensuofi anr{He nocreanbBeorrrrpHoft qactu (l-la); E - no ornocl{Tenrrnoft Belraquue 6oxosono AnaMerpa y BepuuHbr anrBeorrbr (SS)
5l
nI{HeH}Ibte
aKaeMnnspbl
ilo-tsr{Ar{MoMy
nepeAl(o
nOnaAaIoTCs
B o{turry( rronyflJlUtfix.
Kpraae
B OABID( n nex-xe H3MeHeHI{fi
FE
Aaer
o6paeUaX,
T.9.
oAHoBeput|lrHblfi
rpaOru( (pnc. 186). I{ Ap.) npncyrcrB}tlor y3Ha Heroropbrx pocrpax M.aequalis (pocrp l.fe 8?-?8 KHe rpeur{Hbr rrma 6opoaasl Ha cnr{Hxoft n pexe 6ptotuuoft croponer K€lK npaBl{rlo' B tlcrresaroutae npu npn6nuxeulrl{ K BepuI{He ur cneAyer Ay}rarb, o6paeyrcuhecs pocrpa cnttlrHo{ptoutHovy no BeulecrBa BbrrqenaquBaHus pesynbrar€ nosArreftueno cHcr€MarIF A'aMerpy. Mbr He crrr{TaeM BogMoxHbIM 3TI{M 6opoeAar',r nptml{cblBarb qecKoe SHaqeHue. no 6nuerofi r cy6uunuHApl{.lecroft $optue pocrpa I{ CMeIIIeHI{K) cpaBHeH[e. ero Bepur{Hbr K cnunnoft cropoue onncblBaeuufi nnq orrrtrqaercs or 6oruuruEcrsa (Oppel, 1856-1858t eHAoB Mesoteuthis. CxoA,rrbreno Sopr.re pocrpbl M.rhenana S.363)uM.pseudoelongatasp'nov'orHocurenbHo6oneeyAnrHeHbl'6oaeeMac\cI{BtIbl,HMeIoTMeHeeBbtTgHyr}4onpuBepuHIIH}'IoqacTb. OaunarrbHarr np uyp orreHHoc ru. Pocrpw M.aequalis s 6oruurx Korlrqec!tolBax, 3arracryp coBMecTHo c npeAcTaBI{TeJrflMI{Nannobelinae n Possaloteuthis a Bunoficrov necqaxo-nnl{Hlrcrblx MerrKoBoAubrx $aunruc B (Pavl. Bcrpeqatorcs /i ), pafione. Qaunnx a Ana6apo-Xaraxrexoru paftoHe r{ HecKoJrbKo pexe B Tarnx-xe pafiOnenercKoM a Mops orKpblToro Orer15 peAKr{ pocrpbr aroro Br{Aa B ocaAxax ttTo npu6pexllbtMl{ C casslgbl 6blnu xt{BoTgbte npeAnofloxl{Tb! one. flpuxoAl{Tcs C npruloHHblMtr yc[oBEgMI{ XIlSHtr. O6CranosKaMr{, BG3MOX(HO MeIKOBOAHbTMH p a c npocTpaHeHUe. Toap (eonaHapou reorpaQrsecroe Bospacr Cn6rpn. Ceaepuof, ceras falciferH Bblrue) per Bnruon, Cnnefi u 1O1 pocrp t{3 HExHeno Toapa 6accefina MarepHan. aepxnefi qacTt| pocrpoB 15 KUprHofi, T.H. 13 rlnHae - c6opul T.H. Hansnsesofi H - c6opu T'H' Hanuugeeofi' 2 pV HHxHeno Toapa no6epexua Ana6apcKoPo 3aIII{Ba c6opu T'H' Knpuuofi' crpa [I3 Toapa HIr3oBbeB p. OaeueKa
I
lvlesoteuthis tiungensis Sachs, sp' nov' Ta6n. lll,
$vtr, 4'7
BnMyaefi Hfr co AH CCCP, Honocu6rpcr' Pexa Ne 87-81, lonornn. commune' Dactylioceras nrcft, u[xunfi roapr 3oHa cy6Pocrp rrenxufi, yAnrseuHrrfi (eenu'IuHa IIa oKoIIo 320-460)' AnarHo3. Ueutpanr 3aHl{Maptqefi Bepuruofi, c UunlrllApl{qecxufi unw cna6o BepeTegosttAxulfit cerleHue cTOpOHe. Ilonepevuoe HOe nofioxeHr{e r{nH cnerKa cMeueHnofi x cnnnnof, cnunxo-6oKoBble flpunepuuggble (EE 8O-96)' oxono c 6oroe oBanbHoe, cx(aToe 6oposnur BblpaxeHbl B pa3nofi crenenu' fla' f l p u e e p u r r ' r H H a qq a c T b s a H I { M a e T M e H e e n o J I o B I { U t l npr{3HaKn. Bueunne gnU Qna6O Cy6Ur4nnrUp11rfeCfOfi cpenuefi OT OTAenseTcq OHa AOBonbHo oTrIeTJIltBO qaCTI{ pogTpa. BepurnHa saOCTpeHa, Cna6O CMerUeHa X CnnnnOfi BepeTeHgagAuofi yBep(ta6n" L7l' B npeAeflax 2g-4go cTopoHe, BepuHHHbrft yron rone6nercq npnllonoBuHbl ao 6oposAbr, nporsrt|BalolUHecs cnr{HHO-6orosure ruHHbt ua6ruoAaprcs 6opoe6prorusag npuBepugggas u npHcyTcTByeT Hnorqa pocTpa. BepuJHHHoft qacru napnue 6Oylurguregblt HHOfAa Ha HtD( HAMeqaprcg Aa. EOrOBbte cTOpOHbl CnefKA 6pnutecTb KorAa CTOPOHaT B TeX cflyqas( KOBbte noJr@bt. Epuuuan K ne6nRXe TOXe JTTUIOUeHa, B 3aAHefi nonoslrHe PocTpa uas 6opoeaa, h 4 | | t tl It I tr
l\i/l
Irl \ il \l/ Y 52
-r1
1 Hassa'ue no p' r,'*' BuAa tiungensis Sachs' pxc. 1g. flpoaonruoe ceqeuuepocrpa Mesoteuthis commune, Dactylioceras soua .sp. nov. \fe BZ-Aa, uroruufi roap, P. Bumofi
Ta6nuua L7 l4svrepeunfi pocrpoB
ltlesoteuthis tiungensis Sachs, sp. nov.
flapauerpur 1.[nuHa 06ruaff
I nPeanonanaeMas I ycTaHoBneHHaq
1]nnHa nocneanbaeonspnoft
IJ b8 7 -8 1 , p. Buntoft
4 s , ; ( s s 8) 35,O(438)
TIACTH
Irb 87-82, p. Bunrcft 37,o (597 ) 3 4 , O( 5 4 8 ) 26,0(+rg) (70)
fiuar"lerp cnuHgo-6prouruoft y BepuuHbt gnbBeoJlbr
I,O ( 100 )
6,2( 100 )
finauerp
6,5(81)
6,0(97 )
6oxoeoft y Bepuanbl
AJIbBEOJIbI
Paqnyc 6prouHoft flnr.lua npnBeputlHnoft .{acru flnaverp cnunno-6prcu:uoft e npuBepufiHHoft qacrn Auaverp 6oxosofi B rpr{Beprunuuori qacru Ynon anbBeonfiprufi B cnr{HHo-6pror:ruofi nnocxocrt{, npaA. Vnon BepuHHHHft e cnuHHo6pronuofi finocKocru, rpaA.
15,5(194) 8,4( ros )( 100 ) 7,O(86)(83)
1 J - , 2( 1 8 1 ) 6,3(102)(1OO) 5,7(e2)(go)
43
32
Orou.ranue ra6nuqu
[Iapauerpu Anuna npeArronanaeMarr o6uac ysraHOBneHHAg Anuna nocneanbeeoanpnofi qacTI{ Auartaerp cnunso-6prournoft y Bepurt{HbranbB@nbr .0,uarr,rerp6oxoaofi y Bepun{Hbr anbEeonbr Faaryc 6prcunofi Annna npuBepurnnoft .racru A,uaverp clr{Huo-6prcurrofi e flprlBepurnnofi qacr[ Anar'aetp 6oroeoft B rrpuBeprulluHofi qacru Vnon aJrbBeonspxrtfi B cIIlrHHo-6prcurHofi nnocroerrr, npaA. Yron BepuulrHbrft e cnurHo6prorunofi rtnocKosru, rpaA.
l& 8 7 - 8 3 , p . Bnruofi
s o , ; ( s o o)
]s
87-84, p" Baruo*
24,8(4O7 )
3 3 , O( 5 6 7 ) 29,3 ( 5O5 )
6 , 1 ( 1 O O)
5,8 ( 100 )
5,3(87)
4 , 3 ( 74 )
L2,7(208) s,7(93)(100) s,o(82)(88)
2,4(4Ll 13,5(233 ) 7 , O (1 2 1 ) ( 1 O O) 5,7 (98)(81) 23
29
37
pealreMy KoHUy pocTpa cTaHOBTTCs BHnJnsrofi. Cnrngas cropofla Bbrn]rrrras no eceft AnuHe pocTpa. flonepevu@ ceqeur,e [MeeT 6opuy cxanorp c 6orog oBalra, cTene'b 60rosono cxarus sAMemo Ee Me'Cercs B nepeanefi n s*Eef, qar* poc?pa. BsyrpeHHI{e npI{3HaKI{ n oHToneHes. AlbBeotra sarrrMaer orono or3 AJrr'rHbrpocrpa' BepuaHa ee cJrer*a npn6nuxe'a r 6pounof, cropoxe (6prourxofi PaAayc oKono or4 cE). Arseeonepnuft )Torr 2go. b."""" ,r""-ii saHgMaer 6rnrs.Koe K ueHTparlbHoMy [onoxeHl{er fio.trl{ coBepIIIeI{Ho npgMa.g. Ha naqarlbgbrx cra-
53
6oll.tittngensi.s upretor cna6o BbtpaxeHHyro cy6xoH[{qecKylo $opuy, Ar{flx pocrpbr qeM pocrpbl B3pocJrblx nee 3aocTpeHbr, HecKonbKo MeHee oTHOCtITenbHo yAnI,IHeHbl npr,r CE, paBHoM 3,5 r.av slra'{eHl'Ie fla Y pocrpa Ne 87-84 oco6eft (puc.19). f l p r ' r v r v 5 O 5 . auavrerpax oKono 4 vv pocrp yx1e npnpaBHo 37O, npu CE 5,8 eLIAa cy6uunullApt{qecKylo Sopr"ry. Y espocnrtx AaHHoro Arnr xapaKrepHyro o6peraer nepexoA or cy6uunullApmecxofi SopMbI pocrpa 3KaeMnrspoB HepeAro ua6npAaercn x cna6o BepereHoel'IAHoft. H s v e H q r . r Bo c r u . O 6 u r a q S o p v r a p o c r p o B n o A B e p x e H a 3 a M e r H b t M I ' I 3 M e H e H H S M . u cna6o cy6xocna6o BepereHonuAuoft Bcrpe.raroTcrr pocrpur cy6ur.rnululpnrecroft, 3Har{euus fla (or 26O npeAelulx rone6ruorca B cylecrBeHnbrx H1aqecKoft {opvru. 1OO). ilpeo6na.q,aor s Haueft KorIrl€KIlHt{r Hacr{I'ITblllo SgO) u EB (or 70,qo Y neu EE - 80-96. fla B npeAerlax 320-460 p o c r p o B , 3 H a q e H u s 9 0 Baroueft (V u:lrporas I'I Henny6oxan IIpuHs 86) t2 Koropbrx pocrpoB r.rs 6acceftHa Burnos I{ Ap.). EopoeAa 3ra KMeer uHofi BepuLIHHan 6prcuruaq 6opogga (pocrp Ne 87-82 Y 6onsy p o c r p oB po'qa 'lcrocoelites' t { e M y 3 K a n 6 o p o 3 A a u r n y 6 o r a a xapaKrep flpu 3ToM pocrueft .{acrr.r pocrpoe 6prcu:uan 6opoeaa coBepueHHo orcyrcrByer. pbr c .rerKofi 6proruuoft 6opoeaofi u pocTpbt, nHueHHbIe ee' nonaAaloTcq B oAHUx pocTY ueroropblx B oAHlIx nonynqqut{x. o6paeUaXr T.€. no cyqecTBy r,I Tex-xe pa3pon cna6o BbtpaxeHbl LI npI{BepuHHHbIe cnUnno-6oKoBbte 6opoenul. Bce sru HoBI{AHoCTHcBg3bIBaIoTc'IApyTcApyroMnocTeneHHbIMlInepexoAaMl{t,Ipa3AenI,ITb ux npaKTHqecKI'I HeBO3MOXHO. l+lesoteutltis Cpae He H r,re. Or 3anaAHoeeponeficxux no3AHeroapcKllx Meru(ux (Lissajous, w A c r o c o e l i t e s c c y 6 u r . r n u l u t p w q e c K l r M r .pr o c r p a M u I t t l . n a t i s c o n e n s i s
L g 2 7 , c r p . 1 8 , r a 6 n . 2 , 0 r { r .g - 1 2 ) , A . d i t t b e r g e n s i(sK o l b , L 9 4 2 , c r p . 1 6 1 ,
OBanbHbtM' a He oKpyI'JIblM onr{cbrBaeNatrfi er-rA ornlrqaercq ra6n.5, fir,rr. 22-24) uopogrpbl M'tiungensis nonepeqHbtM ceqeHneM pgcTpa. Her Ocuoeaugfi Ct{uTaTb pocrpa' (cyqs no cy6ulrnnuApuqecroft {opt"le Br{Aa noAbro KaKoro-nrn6o npyrono Y poclpoz M.tiungensis aro Moryr 6srrs pHbre oco6u ronbKo Passaloteuthinae). Bblpaxeubl npllBepuuHHble 6opoeArt' caMH poCTpbI BCTpe6onsurefi rracTbro xopouo 6acceftsa Burnos s 6ottbtul'Ix Kon[qecrBax. ornoxenuqx qarorcrr B HrrxneroapcKr{x pocTpbt ill.tiungensis, KaK I{ Bcex MegateuKpgrnle Toro Ha HaqaJlbHblx cTaAldqx K pa3MepaM B3pognono 111t{B9r npra6naxeHul{ npu nuulb LI thinae, cy6Xonr+recKne y ApIrpI{x e6doe MesoteuHono pocrp cTaHoBHTcn cy6UunnHApwqecKllM. IIpu arov r cy6uunurqcy6ronE'recroft or repexoA pocrpaMu this c c$uunuHApr{qecKr{Mu qeM AEaMeTp B3poc66nuurax, npu AHaMeTpax puuecrofi fropvre ocyulecTBJulercn nono Pocrpa M.tiungensls.
Pocrpu
M.tiungensts oqeHs MHoFOHHC-
neHHbIBMeJIKoBoAHbIxnpu6pexrrblxnecqaHo-|nuHl{cTblxocaAl(axBuruoficxonosanlllub B eAUHI+IHbIX gK3eMnnspax. rr'Ba u sa npeAeJraMri 3Toro santlBa naftAenu B MeJF o6uranu cAertarb BbIBoAr r{To xl{Bortrble AaHHbre nosBoJrrror flpuneaengbre ort{ tls6enann o6crauosKax npl{aoxl{blx Bo3MoxHo Aaxe npn6pexnblx' ,.otoq"o'*
KpbITQnOMOps. HuxHufi roap (eoBoepacr v reonpa0uqecKoe pacnpocrpaHeHue' Cu6npu' Ha Harpoceras falcifer r Bblue) Ceeepuofi c6oPu p o c T p o B E 3 H I { X { H e n Or o a p a 6 a c c e f i u a Bnnoc Marepuan.86 no6epexua Aua6aPToapa poctp lr3 HI{xHero 1 Kupuuoft, T.V. u Hanunsesofi T.l,l. roapa Ouonoscxono' cr
Mesoteuthis s ubrostriformis Voncnez Ta 6 n . XIV ,O u n . 5 -7 subrostrif ormis: Boporreu,1962, crp. 1OO, Ta6n.53, S"f' 9, Me'soteuthrs ra 6n. 54, S r m .3 -1 1 . 57, { rr' 3' M es ot eut hi s u b c o n o i d e aB' o p o s e q , 1 9 6 2 , crp.99, ra6n' cl p.98, ra6n. 57, Qnn.617. L 9 6 2 , c o n o i d e a .B o p o n e u , a ff. M es ot eut hrs ra6n' 55, S rn' 5' M es ot eut hias ff. tri s c i s s a : Ku H a c o e ,1 9 6 8r ctp' 133, 54
Ta6nHua 18 Hsr'lepennrr pocrpoa Mesoteuthis sttbrostriformis Voronez IIapar'aerpu
Ne 87-86, Aua6apcxnft
N e 8 7-87, p. Bnnoft
N e 87-88, p. Kennunp
SAJII{B
Annxa fnpeanona,l naevan o6rqas ) )r"t""orL nenHas finnHa rrocJreanbBeoJrspnoftqacru [uar.aerp cnr{Hr{o6prounoft y ,Bep-
3 9 , O( 3 9 O ) 5 8 , O( 3 4 9 )
3 8 , O ( 3 S Z1
36,0(360)
4 9 ^ , 2( 2 9 6 )
2 9 , O( 2 6 9 )
24,2(242)(OZ1
1Or6 ( lOO )
1 O , 8 ( 1 O O)
1 O , O( 1 O O )
LS,z(80)
8,5(79)
7 , L ( 7L l
24,3( 146 )
L4,3(132 )
4,1(41) L 2 , 3 (1 2 3 )
IUT{HbI ANbBEOJIbI
flraverp 6orosoft y BepuEHbt aflbBEO'IbI Pa.quyc 6prouruoft Annna npr.rBepuugnofi qacrtl finar.,rerp cnr{Hno6pourxofi B npr{BepurunHofi qacru flrar'aerp 6oroeoft B flpr{Bepurunuoft qacTI{ {."on anbBeonspHslft g crlltHgo6poruuoft nnoc\KOCTI{, npefl. Vron aepurnxufi e cnnmro-6plorunof, rrnocKocTl|, rpa.q.
1 3 , 8 ( 8 3 ) ( 1 O O)
8,O(74)(100)
7,s(75)(1OO)
1O,8(6s)(78)
7,2(67)(90)
6,4(64)(85)
34
39
36
32
Jlerrornn, asl6paxHurfiB.H.carcou: Bopone[, 1962, ra6n. s4, $nr.4. uFM, IleHnnpaq, KoJureKu[g JS g2og. FI-oe yponn-Tyluyc, Toap. AnarH o3. Pocrp ne6orguoft, cna6o cy6roxr.recxufi, )nvrepegHo yAn11HeHgbrfi (exaveuun IIa oxono 2OO-28O), cxarufi c 6oroe (EE orono 76-g2), c osanb_ HblM nollepeqHblM ceqeur{eM. Bepunna gaMerxo cMeruena K clr}rHHofi cropoxe. llpnaeprurHHble cnnruo-6oKoBble 6opoequ pa3Br{Tbr B pa3n6qgoft cre11egg. BnelIJHue nplrsHaKE. I-lpnaepuruHnasqacrb sanuMaer ot(ono florroBr{xu IIa, qerKo He orAensercs or cpeAHefi qacrn pocrpa, BeprrluHa cna6o oaocrpega, Bepurunnuft Jron orono 36-990 y Bspocrrblx oco6e.ft (ra6n. 1g). llpu""p**"oi. .n"r_ xo-6orosue 6oposAbl y HeKoropblx pocrpoB nporsrr{Batorcs Ha Bctcl AJprHy npnBepunnnoft tlacrur y Eeroropblx c rPYqor',r npocnex14Barorcs Ao cepeAr{Hbr 11p11Bepu6lrnof, qagrlr' rHonAa )Ke fioqrr{ coBepueuuo HegaMerurl. Eorosbre croponbr yn[oIleHbl' Ha HrDc MoxHo Blrqerb y orAeJIbHbIx pocrpQB napHble 6oroasle nottocbr. Cnrnsas n 6poluHas cropolrbl BblrryKnbE, y HeKoropbrx pocrpoB sBJrsercs 6oree aunytcnoft cnI{HHas cropoHa. Crenenc 6oxoaoro cx(arus pocrpa saMergo He MeHsercg no ero ArIEHer nonepeqHoe ceqeHHe l{Meer $opr"ry oBana, r{HonAa cflen16a Tpaneq[eBr{AHoe 6nanoaapa pacullrpeHrn 6prcruHofi cropoxu. B H y r p e H H t t e n p u 3 H a K u n o H T o n e H e g . A n s s e o r r as a H u M a e r 4 0 % annnrr pocrpar Bepulllla ee cMelueHa K 6prouxoft cropoxe. Epourxofi paAr{yc cocraBnser or 3O qo 40% CE. Anseeonaplufi yron paBeu 34o. Oceaas rrrnus floqrg npcMas,
55
cna6o r{3ongyra B cropoHy 6pou-rxono Kpas u npu6nux(eHa K HeMy. Ha Haqarnrolx g o6ueN{ Mano orrl+IaKtlq}4ocraAurur pocrpbr onr{cbrBaeMoro BIrAa uMetoT 0opt"ty, Bepuuunbrft )tronr nf{ule 3aocrpeu 6onee Hecronsro oco6eft. BspocJrbtx crr or qacrn pocrpa. 3naqenn:r fla Bbrpax(ega cy6rour,recKarr $opr'ra fiocJleanbBeorttlpnofi c pocroM xrrBorgo1.o florrrl{ He MeHgrorcs. V pocrpa, ueo6paxeuuoto H.C. Bopoueu (1962) $a ra6n.54, Srn.3, rpH AEaMeTpe 4rS r't'a fla paano 27O, npu r,,ru - 257, npr{ Ar{aMeTpe 1315 Vr..,r- 268 (pUC.2O). 618 Ar{aMeTpe p o c r p o a M . s u b r o s t r i f o r m i s n O K a s a H aH . C . B e HsN,reHrrr{BoCTb. HeN{e}FIusocTb (fgOZ), poHeu roroparr Aana Soronpa0un 13 pocrpos. CneAyer noAqepKHyrb' qTo oTHOCT{TenbHarr AtrHHa pocrpoB MeHqeTcs oqeHb MAIIO, B [peAe[ax or 2OO ItMeao SOO, ocgoBHarr Mscca pocrpon (8O7. ua 52 3K3. B naruefi ronnerunu) 2OO-28O. 3uaqureJrbllo fla sgaqerrns er 6omue pae6poc sna'renrfi EE (ot 66 go 98), tro Koropblt( 65%rprocqurcsBa Pocrpsr ue BunpfiBenr*tunbr 82-92. Puc. 2o, IlpoAonsnoe cKorp pafiorra HMeIC/tr KaK npaBunot ceqenue pocrpa MesoteuMenburne pasMepbl rIeM pocrpbl I{3 AHathis subrostriformis Voropafiona (npeo6naaalq 6apo-Xararr\cKono nez (BopoHerr. L962, lune gHaqenng CE cocnBercrBeHno 8-11 ra6n. 54, Qrlr. 3). Toap, r'rM). Han6onee MenKue' EloSrrr 10-14 (xO,7 ) r..oB VprolrsTwyc MoxHo roxble pocrpbl co6panu a Onenexr"r). cKoM paftone (ns o-ro C p a B H e H ue. lrl'.subrosttiformis BnepBbre Hs Ceaepnofi Cn6upr (c n-osa Yprorr'(fgOZ), nprveru ep 6slna noKasana E BHyrptrBt{T1nayc) onrcan H.C.Boponeu pocrpoe. B onucaHaer AaHHoe H.C. Boponeu, BKpar{sMeutrr{Bocru AoBaq SopMbr nncb Herotrnocru - 6rlno yKa3auo, qrp BepuJnga pocrpa cMeuleHa r 6pnurnofi croBepurl{Ha naoSopor cI{JIbHo cMeIIleHa K cnulrHofi poHe, e Aeficrs[Terlbnocru-xe BWIoB Passaloteuthidae. CNreuexre Beput{Hbl y MHonHx H Ap)'trHx KaK cropoge, pocrpa K cnuHgoft cropoHe xopouo BI{AHo na Qororpa$unx, npuroxeHHblx K pa6oTe H.C. BopoHeu. Quen6ngexul"r sEAoM rrBnsercrr M.breairostris (d'Orbigny) (rostrifotmis u 6onee trpaBDrJlbxblficy6roxEqecstedt), nNre19u11ftorgocuT,enuHo 6onee roporrufi xar M.subconoidea Vor.r rufi pocrp. Pocrpur, onncaHubre H.C.BopoHeu (fgOZ) xors oHl{ (Oppel), B M.subrostrilormis, ct{HoHHMtrKy exnoqHnH vul M. aff. conoidea BepuHryflble I{MePT 6onee 6oposa cnnnno-6oKoBblx npr{BepuugHbrx Hanrrir{u npr{ pocTp Hbr qeM y 6orUruuHcTBa pocTpoB paccMalpEBaeMoro BI{Aa. Hao6opor, touufi 6nus6onee Mbl ctt[Taer"r 12) (BopoHeu, ra6n.54, 1962, 0rr. M.subrostriformis l{cKnlosrnu. BnA subpyramidalis H us cHHoHt{Mr{KH Kr{M K M.breairostrrs(dbrb.) H s 3 o n b rl l i l d ' o c e r a s b i f r o n s H I D K (Lissajous, Lg27, crp. L7, ra6n.2, firn.5-8) M.subrostrilormis 6onee npaBurlbnofi cSxonero roapa Qparurnu ornrrraercs or cnuHxo-6oKoBblx 6opoea, pasBuTuto npnBepLr{Hnbrx no cytrs H, H'.recxoft {opvoft Kax cql{Taer poAY Acrocoelites, K a illesoteuthis, poAy He K orHocgTbcrr AorDKeH rluccaxy, Oauua nbHas n p Hyp oqeHH oc rs. IloqaBiullouee 6olgunncrBo pocrpoe co6ocaqKax Bnnroficrono [r paHo B npu6pexxbrx MerIKoBoAHbrxnecqano-nnunllcrblx paftoHos. EAuHI{ttHbIe MerIKHe pocrpbl' Bo3MoxHo toHblergaftAna6apo-Xararrcxoro no3AeHbr B rnuHrdcrbrx ocaAKax orKpbrrono Mopfl e paftone p. Oneuer. CKasaHHoe a MeflKoBoAHblMI{r qro npu6pexHblMl{ c cBflgaubl xt{BorHble 6rrnu Bonger cr{prrarb, Bo3MoxHo r{ npr{AoHHbtMuo6crauoeKaM[ u peAKo 3anJrblBanu e o6nacrb orKpblToro Mopfi. Toap (eona pacnpocrpaHeHne. u neorpasnvecKoe Bospacr Cn6upu. Ceaepuofi u euue) falcifer Harpoceras 37 pocrpoB r{3 Toapa 6acceftua per Bnnoc u IIHHAe - c6opu Marepuan. T.H. Kupgnoft n T.H. HanunReaoft, 10 pocrpoB H3 Toapa Ana6apo-XaraHrcroro paftoua r.r 5 pocrpoB H3 Toapa p.Kenr.lunp - c6opur T.V. Ha6Hsesofi. 56
M es ot e u th i sb re u i ro s tri s (d ' Orb i g n y ) T a 6 n . X VII, Su n .5 -7 Belem nit esbr eu i ro s rri sd: ' o rb i g n y , 1 8 4 2 ,p . 9 6, pr. r0, fi g. r_6; Quenstedt, l B5 B ,5. 287,T ab. 41 , F i g . 1 2 . B e l e m n i t e sc u r t u sd , C r b i g n y ,1 8 4 5 ,p . 2 7 5 , p | . 4 2 , f i g . l_0. Belemnitesacutus: Quenstedr,18.46_l}4g, S. qZq';fZt.-ZZ, FiS. 13_lZ. B er em nit esr os t ri fo rmi s :Q u e n s te d t,1 8 4 6 -1 8 4%q: 42s,i af.2V , Fi g.- 1g,20. A c r oc oelit esr os tri fo rm i s :K o ]b , 1 9 4 2 ,S . 1 5 9 , Taf. 10,' Fi g. 2_+ . A c r oc oelit esc u rtu s : Ko l b , Ig 4 2 , S. 1 5 9 , i " f. fO, ni g. tZ1f a. Nannobeluspau l o u r: Bo p o H e u ,1 g 6 2 , T a 6 n . S S , S nr, g. M es ot eut hiss ub ro s tri fo rmi sBo : p o n e u , 1 9 6 2 , ra6n. s4, Qnr.12. M es ot eut hisr os tri fo rmi s : Kn u a c o e , 1 9 6 9 , c rp. 132, ra6n. ss, {rm.2. rl e x r or Hn, ns6 p a n n u rft B .H .c a x c o v - d ' o rb i gny, 1g42, Ta6n.10, Qrn' .1. -lnanHo3. Pocrp ue6onsuoficy6xoHnqecx[fic orreHbroporxoft nocfleanbBeo-
napHofi qacrblo (IIa-14O-24O), c osanbHblM cxarbrM c 6oron nonepeqgbrM ceqeHI{9M (EE-7O-86), c pesKo BbIpaxeHHbIMI{npI{Beput{HHbIMucnuuuo-6oKoBbrMn 6opoeaaun. Bepu,nHa pocrpa cMeueHa K cnr{Hnofi cropone. BHe ulnue np lr3HaK I'I. Pocrp LIMeer cy6xounrecK)4o 6opr"ry, npaBr4nbHocrb xoropoft npI'I B3nnsAe c6oxy Hapyulaercg BcneAcrBue cMeuleHr.rs Bepuglrbr K cnr.rHHoft cropoHe. Bepurna saocrpeHa, Bepuunnuft ynon nopsAKa go_g60 (ra6n.1g). I-lpnaep':n'Ha' qacrb pocrpa pacnpocrpaHrrercrr Ha Bc,o nocJreanbBeonsp')no er.o qacrb. IlpuaepuuHHbte cnuHno-6oKoBbre 6oposqsr nny6oxue H ysKne, nporsfgBaror!cs. ua 2/3 l-la. Huorqa y Bepur{Hul ub6nroqarorcrr AonoJrur.rrenbHbre cKnaAKH u lurpr{_ xoBarocTr" CnuuHoft xpaft fioqru npauofi, 6nnxe K Bepur.rne uHorAa cnenKa BonHyruft, 6ptotuHoft rpafi cna6o nunyxnuft. EoxoBbre cropoHbr ynnoueHbr. flonepeqHoe ceqeHne r.rMeer $opr"ry cxarono c 6oxos oBaJra. BHyrpeHHHe npH3HaKI{ vr oHToneHee. Anseeona 3 a n r { M a e ro x o n o n o noBuHbr AnnHbr pocrpa, BepuHHa ee cMeueua r 6prcunoft cropoHe (6prouuoft pa_ AHyc 367o CE). Anaseonspuuft ynon paBeH g2o. Ocesas nulvs, noqrn [prrMarr, cnenKa npu6nuxeHa x 6prouuofi cropoHe. Ha Ha.{anbHbrxcraAunx pocrpbr 6onee saocrpeHbl' HMelor 6onee npaEHnbHyro cy6ronr+recKyro $opvy, creneub orgoclrrenbHoft yAn[HeHHocrH c Bo3pacroM He MeHqercn. Y pocrpa Ne 8Z-g1 npu Ar{aMerpe 3 r 3 . r ' a v [ I a p a e H o 2 3 6 , n p r { A r . r a M e r p eg r 7 u u - 2 5 4 ( p n c . 2 L ) . V l e v . e H q t , l B o c r s . M e u s r o r c s 3 H a r r e n u q[ I a e n p e A e n a x o r 14O ao 260, npu_ vevr npeo6naAapT BenlrqHHbr160-22O.3Ha.renns EE xone6nprcrr or 70 ao g6, y orqenbHbrx pocrpoa ao 90. He scerqa trOgraTOqHOoTqeTnUBO Bbrpax(eHbl npHBepur{HHbre 6oposgu. CpaBHeHue. Oqnn r{ Tor )Ke pocrp c xopotuo BbrpaxeHHbrMr{ npr{Bepur{HHbrMH Pnc. 21. llpoAonunoe cerreHxe pocTpa lvlesoteuthis cnnuuo-6oKoBbrMr{ 6opoeaauv u3 Toapa QpaHuuu 6sln onucan a'Op6r{Hbr.rB Lg42r, bretirostris (d'Orbigny ). n o A H a 3 B a H u e MB e l e n n i t e s b r e u i r o s t r i s Ne 87-90, Toap, p. .Ilnuae r { B 1 8 4 5 r . n o A H a s B a H r { e MB . c u r t u s (nocxonrxy eue B 1829 n. 6srn BbrAer n e H B e l e m n i t e . sp i . s t i l l i f o r m i s v a r . b r e a i ros/ri.s [laspail). B AanaHeftu:eu noAo6Hbre )Ke pocrpbr yKa3brBanr{cb KaeHUreAroN, [r3 BepxHero roapa I'I HHXHeI.O aaneua rcxsoft qacru OPf noq Ha3BaHr.rrrMr,tB.actt_ Itts w B'rostrif orntis. Kons6, npH3HaBarr caMocrorrrenbHocrb BHAOA rostrilormis vr curtus, BKnroqun r{x B poA Acrocoeliles. flpu sToM oH oTMerHJr, r{To y A.cultu.s 6prcnruan 6oposaa Ha pocrpe orcyrcrByer, y .l.rostrilorntis - nu6o orcyrcrByer, nr6o aulpaxeHa oqeHb cna6o. Paenn{tlg MexAy 9Tr{M}r BrulaMH Kons6 B}1AHT B Be_ nr{qHHe anbBeolnrpHono }Tna: 3Ta Ben[r{usa 300 y nepBono Brua, 26. 270 _ y BTO_ poro' cyaa no Soronpa$uarta, aMercTcs He3HaquTenbHbre pa3ntl.rH,t H B orHocgTenbnoft yqnnHeHllocrH nocJIeaJIbBeoJuIpHoft .racrn pocrpa fla paBHo coorBercrBeHHo
57
T a 6 n u u a 19
H
Her"repeHuapocrpoB M e s ot euthis br euirostrr's (d' Orbigny)
c
c
flapauerprr
Ne 87-90, p. IIEnqe
N e 87-91, p. Mapxa
r
r finnna [npeanonaraeMarr l ooulas LycTaHoBneHHaq finnna nocneanbeeonapHoft
44,O(427 ) 38,O(369) 2 4 , - 7( 2 4 C - )( 5 6 )
35,O{4O2) 2 9 , O( 3 3 3 ) 20,4(234) (s8 )
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10,3(1OO) 8,5(83)
8,7 (1OO) 7,O(80)
AJIbBEOIIbI
Paauyc 6prourHofi Vron aJrbBeonspnrrfi B crrrrHue6pounoft nnocrocr[, npafl . Vnon Bepur{HHblfi e cflHruro6prouuoft nnocKocru, npAA. Oronqanue
Anuna oorlae .d,nrua qagtt{
[uaverp
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cnurrno-6prcrunofty
BepurnHbr aJrbB@Jrbt flnavenp 6oxoeoft y BepuuHbI
21,O(265) 1 8 , 1( r 1 7 | 1 O , 2( l O O )
]$ g7-93, p. Jleaufi Keqon 3 0 , O( 3 8 4 ) 23,8( 3O5 ) ls,4(197)(51) 7 , 8 ( 1 O O)
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Paaryc 6prcunofi Yron anbBeonspnrrfi B crrnxHffiprorfiHoft nnocrocrr{, npaa. Vron Bepurnubrft e cnnnxo6prorunofi nflocKocrr{, rpaa.
167 u 2OO. Cu6npcrne pocrpbr no creneHr{ 3aocrpeHHocra 6nuxe Bcero n 0or MaM, OIIUCAHHbIM Q. KSeH|xteqroM noA HaSBaHHev-Belemnites acutus. Bce [epeqilcneHxHe orHec 3. Ele 0opr,rsr K oAHoMy Br{Ay ( brevirostrisl non-Tpyrvrr..rep (Bfilow-Trumrer, Msr [pncoeAnuse]rcs 1920). K oroMy saKJrrotIeHHIo I{ nocKonbKy npnBepu[HHac 6prcu:naa 6opoeaa Ha pocrpax orcyrcrByer, BKflroqaeMAanssrft Brul B poA [fesoteuthis. Her ocnosaHtlfi 3aMeHrrTb Br{AoBoe Ha3BaHHe Ha curtus, KaK ero cAenan A'Op6uHbt{, rar( KaK Pacnaftns (Raspail, 1829) BbtAerl[n nHub Bap[Ierer breairostris neoxoucKoro Br{Aa pistiltilornis, B Hacrorruee BpeMs orHocttMono K Hibolites. Otnnrns or 6nusrono Br{Aa fil.subrostriformis \oronez npl{BeAeHbtrIpH orrncaHuu nocneAHero. K M.breviroslrds AonxeH oruocr.rrbcrr roRrtfi pocrp M.subrostriformis, uso6paxeHxrrfi H.C. BopoHeu (Bopoueu, 1962, r-a61. 54, {rn. L2). CxoAsul raKxe pocrpbr BepxneroapcKono avga subbreais (Kolb, L842, crp. 154, ra6n. 1o, o r u e c e H H u r eK o n s 6 o M . K - p o A y Qnr.711s,i6), Salpingoteuthis, Ho orBeqapune BceM nplr3naKaM poAa ltlesoteuthis. Onu ornr{qalorcg 6nusrEM K oKpynnoMy nonepeqxbrM ceqeH[eM, 66ruurm.an 3Har{eHr,rflMgBepluI{HHOnO
58
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He ucxrnoqeHo, rrro pocrpbr, oruocl{Mbre K BI{Ay M.brevirostrr:s, Bce Hrr1tEacTKrrHor sBnlorcq MonoAblo ApITI{x BI{AoB Mesoteuthis, o6naAaBrrtInt(Bo BspoctroM cr crofiHlll{ 6onee KpWHrIMI{ n 6olee yArrlrnenHbrMu pocrparraa. OAnaKo cBgsarb ollttcblBaeMble pocrpbl c KaKHM-nu6o AplrI{M onpeAeneHHbrM Br{AoM Mesoteuthis xe npeAcTaBJuIeTcrI BoSMoXHbtM. Q a u r a an b H a a n p u y p o q e H H o c r s . B o r E u J r { H c r B p oo c r p o B M . b r e u i r o s t r i s HaftAeHo B npl{6pex(Hblx MerIKoBoAHbrxnecqauo-nJrnrrr{cTbrx ocaArax Bnnoficfono B pafioHoB, eABEI{rrHbrepocrpbr Bcrpeqaprcs Ana6apo-Xararncroro r{ B nrruHucrbrx ocaAKax orKpblToto Mopq n OnenercKoM n OuononcKoM pafioxax. Cyae rro roporL KoMy cy6rouw'recKoMy PocrPYr xl{Borgbre He 6rrnn xoporur{Mr{ nrroBuaMn r[ BooMoxHo Benr{ npunonubrft o6pas xr{snu. B o e p a c . r a r e o n p a 0 t a t r e c K o e p a c n p o c r p a H e H u e . B e p x n s nq a c r b HHxHero roapa (eoHa Dactylioceras commune u Bbrule) - nuxnnfi aanen Ceaepnoft Cu6upu, Tosp 7 HuxHr.lfi aaneu 3ana.qHofi Eaponrr. Marepuan. 8 pocrpoB I{s aepxuefi tracr[ HHxHeno roapa - Hurter€ro aanena 6acceftHa pex Burnos n JIEnae - c6opu T.l4.Hansngesofr u T.Ll.Knpuxofi, 2 pocrpa Es roapa Ha p. Kenuuap, 2 pocrpa ne eepxnefi gacrr{ Hr{xuero roapa no6epexbs Ana6apcroro 3armBa I{ 4 pocrpa r{s roapa Hr{ooBbeB Onenexa - c6opu 2 pocrpa I{s roapa 6acceftga Or.,ronoxa - c6opu A.C. Aanuca. T.H.Hanrnseaoft, P o q P a r a m e g a t e u t h i s ,G u s t o m e s o v , 1 9 5 6 . . Megateuthis (Paramegateuthis):fycroMecoB, lgs6, Caxc n Hanunaeea, 196?2, crp.13. P aramegateuthis: Hanunaeaa, Lg7 4, crp. GO.
crp.Z,
1960,
5 crp.,tglt
Tnn poAa. Paramegateuthis timanensis - lycror"recoB, 196O, crp. lgl, ra6n. 44, fivr,4r5; nlrxnnfi rennoaeft, p. Hxua (6accefts p. flevopu). On ucsHtle; Pocrpu ne6oruruue KolrrfrecKr{e n cy6roxnrecK]re yArrtrxeurrbre I{nI{ KOpOTKIite.B npnBepurUHgOftTIaCTI{ Xopouro pasBr{Tbr CnrrXO-6OKOBbre 6OpOeAbI' rlpocnexl{Baloulltecs Ha L/3 nnu:abl pocrpa. Epounae 6opoeaa KopoTl(as, He BcerAa qerlco BbrpaxeHnafl. A[useoJla npsMag, HHonAa cflenra ugonHyrag, co cna6slpr cMeuIeEI{eM Bepurl{Hu r 6pounroft cropone. AnuHa atlbBeoJrbr cocraBJlger L/+L/2 nnwtbl pocrpa. Ocenas nwfiur. noqru fipsMas. Ha na.rar6gbrx craggsx pasBr{Tr{s pocrpbr no Qopr"re noqrH He orrrrItraKlrcg \ or aepocJrbrx. Buqoeofi cocraB: 6 enAos na Ceaepe CCCP n I rnrrrb rpeArronoxuTerrbrro orHocrfMbrfi r AaruroMy poAy suF-(bajosr'cus ) a floaoroxue. CpaBHeHue. Or poAa Megateuthis, K KoropoMy aBTopbr B cBoEr npeAbrAf,uurc pa6orax orgocnnl{ npeAcraBrrenefi onl{cbrBaeMono poAa, oH ornt+taercg Aope3KO, rpeTAe_ BOJTbHO _Bcero orcJrrcTBreu 6pourHo-6oxosrrx 6oposa, a raKxe no TI{ny oHTorenesa. Poa Megateuthis xaparcrepgs}rercs 6onee KopoTt(trMtr pocTpaM5 Ha HatranbHbrx craAl{gx pagBETI{g t{ cl{JrrrHo BbrtflHyrbrMH sleHb KpyrlHbrMtr pocrpa_ MI{ Ha RSpocJIbIx craAusr(, c oBanbHbrM Menee cxarbrM c 6oroa, qeM y Paramegateuthis fionepetlnblM cetlenueM. flosroMy T.H.HarcHseBa B L9'14 r. BbrAetrsBF uuficc B.A. IycroMecoBblM noApoA Paranegateuthis B poAe Megateuthissoaeena B paHr poAa. Eonsuee cxoAcrBo orMeqaercs y poAa Paramegateuthis c 6onee ApeBHr{M peAoM Mesoteuthis. Or nocneAueno paccMarpusaeLrbrf, pq orntrqaercs fupvof, poc\TpoB - &Pouo aHpaxennofi KoHIFIecKofi Qopuofi, qerKr{Mr{ AaJreKo r{Aytrlt{Mr{ BAorIb pocrpa npI{BepuBHHbrMIr cnumro-6oroBblMu 6opoeAarrau rr Haflr+rr{eM ropor rofi 6pronrnofi 6opoegrr. ., 3aMerraHI[s. B.A. fycrouecoB ( 1956, 19GO), aslaensr [oApoA paramegai , teuthis, Aan Heroqnrtfi eno AEarHos. Kar norasarr flpocMorp Korrnetcunr B.A. f)roToMecosa T.H. Hanuugeeoft napy nprBepurutrbrx cnmxo-6oroBbrx 6oposa Ha pocr,pax B.A. IlcroruecoB fipl{Hsn sa 6prcune6oxoBbre, xorc Ha runoBoM gK3eM[nspe P.timanenstls coxpaHl{rucb Hatla[bHr'Ie KaMepbr Qpanllorona lr B]utBo noroxen]ro cf,-
59
Soua Ha nporlrBononox
BI,IAOB
HET.
Boepacr n n e o r p a S u v e c K o e p a c n p o c r p a H e H u e . B a f t o c - H H 3 bH r trxHeno KennoBefl Ceeepa n Aansnero Bocroxa CCCP ( or 3er'anu Qparura HocuSa r.r 6acceftua fleuopu ao Ceaepo-Bocroxa CCCP r.l flpr.rar'aypbq). no rpe6yroruuM npoB e p K e A a H H b r MB . f I . K u s a c o B a ( f g O g l P a r a m e g a t e u t h i s Bcrpeqaercs r{ B BepxHeM aaneHe.
; Onpe4e"nnreilbHanra6rnqa BnAoBpoAa paramegateuthis 1.
Poctprr c aByMs npI,[BepuI{HHbrMr!cnnuno-6oKoBbrMn 6oposlavu r cna6ee paeeurofi, He Bcefaa rlplcyrcrayrcuefi rpuBepurHxofi 6pnurxofi 6oposaofi, cy6roxr.recrofi I{Jru npaBunssofi KoHlrqecKofi @pvur, cxarbre c 6oxos. Pog Paramegateuthis Gustomesov, 1960
Pocrp yqnuueunurft, fla oxono 5OO-7OO. P,nescia Nalnjaeva, sp. nov.- BepxHHfiaanex ? - 6aftoc - Hr{xHr.rftxennoeeft 3 (7) Pocrpul yldepeHHo yAnHHeHHbre, I-la orono 25O-24O 4 . . . . . 4(5) Pocrp ue6oruuroft, eepuuua ueuTpanbHas, fla orono 35O-4SO P . m a n i fe s t a N a l n j a e v a , s p . n o v .- 6 a r 5(6) Pocrp cpeqnuft, BepuI.IHa cMerqena K cnuHuofi cropoHe, fla oKono 35O-4OO P.ishmensrs Gustomesov (flcrovecon, 1960, crp. 191, ra6n. 44, fiwr. 415) - 6afioc-Huxnufi rennoeeft 6(7) 3naqenue IIa orono 22O-25O. P.timanensis Gustomesov (rlcror"recoe, 1960, crp.1g1, ra6n.44, 0p;6) Pocrpur Koporrne, fla venee 22O . 7 . . . . 8 8(9) 3uaqeHue fla oxono L7O-22O, 6oxoeoe cxarue pocrpa yN{epennoe (nS 7 5-80 ). P.parabajosicus Nalnjaeva, sp. nov. - 6afiorcrsrxarrfi 6ar I 3Ha.renue ila orono 8O-13O, pocrp cr{rrbHo cr(ar c 6oroe (EE .orono 2(3)
50-7 s )
ol
A T
lu
A a P
Br
v 6
v
H
;
o
r L
u I
a
F
P.pressa Nalnjaeva, sp. nov. - cpeAnnft-BepxHr{fi6ar P a r a m e g a t e u t h i sn e s c i a N a l n j a e v a , s p . n o v . l )
E
) e
J
H
Ta6n. IX, Snr. 1-4 Belemnites borealis: EoAurnencxxfi, e CavofinoBrrr r{ Eoqurnencrufi, 1933, crp.33, ra6n. 1, pnc. 516. Megateuthis elliptica: Knnacoa, 1968, crp. L33, ra6n.B1, {rr. 1. lonorun Myeeft Hlul CO AH CCCP, Hoeocu6upcr. llonyocrpoB Ne 87-L42. nepxuuft 6ar. YprotnTylvryc, Pocrp cpequufi, cl{nbuo nurauyrufi (fla 45O-7OO), y3KoKogr{qeAuarHos. cKwit, B npreepurnnnoft qacrl{ rrerxue cnuHuo-6oKoBbte 6oposau. llonepe.ruoe ceqeHne oBanbHoe, cxaroe c 6oKoB (68 83-95). BneuHue n p I { a H a K H . K o n r . r e c r a a Q o p r " r ap o c r p a x o p o r u o B b r p a x e n a K a K B 6oxoeoft, TaK I{ B cnlrrlxo-6prouxofi nflocKocrsx. 3aocrpeHne K BepuI{H€ nocrerr€Huoe. BepuI{Ha ueHTpaJIbHas I{nH cna6o cMerrleHa K cnunnoMy Kparo, ocrpas.. Bepunnnrtft ynor orono 2Oo (ra6n.2O). B nonepetrHoM ceqeHur cnr{HHas u 6prcurHae croponbr Bbrnyl(Jlbre, 6oKogrle-ynfiouenH. Ilogepxuocrb pocrpa nnnni{as, Jrr{txb B npHBepuunnofi .racrn xa6noAarorcrr qeTK|{e cuurno-6oKoBble 6oposAul, nporflruBato^l ')N e s c i a
60
I
I I
I
I
(
1
(nar.)
-
nesnaKoMaq.
I
I
Ta6nnua 2O HeuepeHaa pocrpoB P a r a m e g a t e u t h i sr t e s c i aN a l n j a e v a , sp. nov.
llapavrerpr,r
-l.urHa I npeanonaraeMarr c6uras I v"tu"orneHHarr ^lnr{Ha nocneaJlbBeonepuofi qacrr.r .-lwauerp cnnuxo-6ppuuofi y BeptIII{HbI
]$ 87-142, n-oa YpnHrTyr'ayc 1 0 9 , O( 7 9 5 ) 9 7 . O( 7 O B ) 7 1 , O (s 1 8 ) ( 6 5 ) 1 2 , 5 (l O O )
AJIbBEOJIbI
luaruerp 6oroaoft y Bepurubr
BEOJIbI
?,6 (95)
6,5( 47 )
4,5(41)
3,5(47)
23
]s 8 7 -1 4 5 ,
Annra npeanonanaeMas 'lI o6uas ycrauoaneHHarr Arnrua noclreanu"eonnpuofi qacru flnar.letp cnuxuo-6prcunoft y BepIUI{HbI
n-oa Ypnnr* Tyr'ayc
9 7 , O (7 7 6 ) 8 7 , O( 6 9 7 ) 66,0(528)(67) 1 2 , 5 (l O O )
AJIbBEONbI
6oxosofi
y BepuHHbr
to,o(81)
AJIbBEONbI
Paq[yc
22
19
ra6nuuu
flapauerpsr
flNaverp
8 5 , O( 7 8 7 ) 6 7 , 5 ( 9 1 2) 80,5(7 45) 6 7 , 5 ( 9 1 2) 51,5(491)(61) 42.O(567)(62) 10,8(lOO) 7,4( 1oo) 9 , O( 8 3 )
Vron anuseonrpnlrfi B cnHHHo6poruHoft nnocxocrr,rr rpaA. Ynon aepurrusuft B cn[rrHo-6ploruHoft nflocKocrr{, npaA. Orouqasue
) , f e8 7 - 1 4 4 , n-oe YprcurTwyc
1O,O(81)
anbBeoJlbl
Pagnyc 6prcunoft y Bepur{Hbr anb-
).lb87-143, n-oe YpoHnTyr"ryc
6prourHoft y BeprIlr{Hbr arrb-
l,fe87-146 Aua6apcran ry6a 9 8 , O ( g B O) 86,0 (860 ) 6s,2(6s2 )(62) 1 O , O (1 O O ) 9,5(95)
6,1(40)
22 20
1 0 s ,( 7 7 1 ) 94,O( 67 1) 6 6 , 0( 4 1L ) ( 6 1 ) 1 4 , O( 1 0 0 ) 12,5(89 ) 5,6 (40 )
B@NbI
Yron ansseonnpuufi B cnfiHHo6pnunoft nnocKocru, Fpaa. Vron eeputttrHsrft B cnr.rHxo-6prcrunoft nnocxocrl{, npan.
ls 87-147, n-oe YprcnnTyr"lyc
27 20
24
urtrecs Ha L/ 4 gnanbl pocrpa. flonepeq'oe ceqeH'e y Bepur,rHbraJrbBeoJrbroBarrbBoe, cxaroe c 6oxos (EE gg_g5). BxyrpeHHI{e I I p t t 3 H a K B U O H T O r e H e g . A n b a e o n a3 a H B M a e r oKono L/g AtlIrHbI poclpat Bepu[Ira ee noqrE ueHTpa[bHasr 6ptouJnos paAlryc y Bepur{Hbr arrb'o".;" Beotrbl paBeH 40-47% cE. Arreeonrpnrrft yron zz-zlo, nu*us npoxoAr{r rloqrtr IIo ueHTpy pocrpa. Ha naqallblrblx craAtlcx pocrpbr cy6ron[qecKr{e, cunbHo BbrrsHyrble' fipn AHaMerpe 612 r.ar'aanAavulfi Ha nplrunn$oexe pocrp r{Meer fla, paBuoe 600 (puc.2Zl. HaveHqHBocrb. B Hsrre'Hofi xonneru[E pocrpbr B ocHoBHoM oAHorrrn*br, HaMen'rlteocrb HegHaqIITeJlbHagr Bbrpaxaercs B Kone6auligx orHocr{Terrbgofi AnI{HbI pocrpa (IIa45O-OSO), creneHr 6orosono cxarrs tnS Ag_g;), crererrr pasBrF TI{g cnr{Hxo-6orosrlx 6oposA. cpaBHeHEe' or Bcex ApyrEx BItAoe poAa ParanegateuthisorrrrcbrBaeusrfi anA orm{qaercs 60ree s'aqr{TerbRoft ygnnHe'Hocrbro pocrpa (lla 470_700, y p, ma_ nife'sta sp' nov' lt APyrux BEAos ne 6onee 4so). fro Qopve pocrpa r pasMepaM K P'nescio olleHb 6nlteox Megateuthis elliptica, olrncanHbrft B.fI. KgHacossll,,r (KaHacoB, 1968, crp. 133, ra6r.8l, Srr. 1) rs cpeAHefi opul Ceaepo_BocroKa
61
Pnc. 22. IlpoaoJrbHoe cetleHue pocrpa Paramegateuthis nescia Naln., sp. nov. It 87-145, ( x Or8) nnxtnrfi xertrtosefi, rr-oB Ypnnr-Tyv]rc
CCCP. Pocrp, neo6paxernrrfi B.fl. KusaconbrM, 6onee cxaT c 6oros (EE 7O), onrocnrerrbno 6onee yannHen (IIa orcono 7OO). TeM HeMF Hee Mbl coqnn BogMoxubrM BKntotrnTb $opr,ry, orrucantryrc B.fI. KrnacoB6rM, B cr{HoxltMurry P.nescia. (3ervrns
P a r a m eg a t e u t h i s m a n i f e s t a N a l n j a e v a , nou.l "p. Ta6n. IX, $rar.5-8 lonorun M 8 7 - L 4 8 , M y s e f t Hfl CO AH CCCR Hosocu6rpcx. fl-on YpromTyrayc, eepxHnft 6ar. nurraxyrufi (l-la 35O-45O), ronuD,u aF H o 3. Pocrp ue6onuurofi, yN,IepeHHo Ta6nnua 21 H s v e p e H u a p o c r p o B P a r a m e g a t e u t h i sm a n i fe s t a N a l n j a e v a , s p . n o v .
l s g 7 -1 4 9 , n-oa Vprcrr*Tyrvryc
rIapaverpu llnuua f npeanonanaeMarr o6ruas I y.""norneHHarr finuua nocfleanbBeonspnoft .{acru fluaverp cnnHHo-6prcruHofi y BepuHHbr aflbBeonbt filraverp 6oxosoft y BepuuHbr
6 O , O ( 6 6 6) 47,O$ZZ7 4L,2(457)(68) 9 , O( l O O ) 7,7(a6)
l t g7-149, n-oB Ypronr,-Tyr'ryc 62,0 (552 ) 5 O , O( 5 2 6 ) 36,5 (Sea) (58 ) 9 , 5 ( 1 O O) 8 , O (8 4 )
AJIbBEONbI
Paanyc 6prou:nofi y Bepurr{Hbr anbBeonbl Yroa anbBeoJnrpnuft B cnHHHo6prcunoft nnocKocru, npaq. Yron Bepur,rHHblfie cnr.ruHo6prouruoft nnocKocrr{, npeA. L
i t l a n i fe s t a
62
( nar. ) -
qsHas.
3,5(36) 24 2 7 i
qecKl{fi. BepruuHa ocrpag, IreHTparlbna"g, CnunHo-6oroarle 6opogAbr xopolrro Bbrp€txenbr, 6poruuan crnaxena. pocrp cxar c 6oxoa (BE g1_g-g). BHeuJHHe npI{3HaKI'I. Kontfiecxaa f,opva pocrpa nrrue Bbrpaxega a 6oroaoft nnocKocrlrr a cnnHno-6pnruHoft rrnocKocrr{ - pocrp ysKoKounrec1nfi. 3aocrpeHr{e K BepuI{He nocrerreHHoe, BepunHa ocrpas uelrrparrbHas unu cna6o cMerrlexa K cnl{HHoft cropoue. Beputnnnuft yron paBeH 25-2go ( ra6n. 21 ). B nonepeu'ou ceqeutlu cilI{HHas a 6pouuafl croponbt Bbrflyl(Jrble, 6.oroabre AoBorIbHo cnnblro yrt_ nouleHbt. flonepxHocrb pocrpa rJraAKas. B npl{Bepu[nnofi .racrg xopotxo Bbrpaxe_ HbI c[I{HHo-6oxoerre 6opoaabl' npocnexgBatouuecs Ha 1./3 anaxbr pocrpa. Epounan 6opo3Ara KoporKasr 3aMerHa He Ha Bcex 9K3eM[Jurpax. I-lonepeqgoe ceqegl{e oBanbHoe, 6oroaoe cxarue sgaqr{rerrbHoe (nS gl_gg). BuyTpeHEue I r p H 3 H a K I { t 4 o H T o n e H e s . A n s e e o n a H e n n y 6 o K a r rc,o c r a B nser oKono L/ 4 anurtbl pocrpar BepuI{Ha ee cnenKa cMeuleHa K 6prounofi cropoH€. BppurHofi pa.q[yc y Bepu[IHbI aJlbBeoJrbr'cocraBnser 36-41o/o CE. ArnaeonnpHrlft ynon paBeg 24-250, Ocega.s nwluet npoxoAr{T rroqrr{ rto uenTpy. Ha Haqanb_ HbIX CTFAI{gX pocrpbl KoHI{tIeCKneBbrrsHyrble. flpn Ar{aMerpe 216 vu pocry r{Meer [Ia 315r npl{ AarrbHeftutev pocre pocrpbr BbrrsrtrBarorcs. flpu ararr,rerpe 4r'S r.,t,r I-la paana 4OO (pnc.29), H g l't e H tI tt B o c r g. y onucaHHono Blt_ Aa r{SMeEgI{BacTeneHb BbrpaxeHHOCTH npnaepur{Hnrx 6opo3Ar y orAenr,Hbrx oKPuc. 23. flpoaoJrbuoe 3eMnncpoB cnr{HHO-6OXOAsre6oposarr crfla_ ceqeHue pocrpa parame_ )Kenbr u Ha cnnxnofi cropoHe outy[Iaercs g a t e u t h i s m a n i fe s t a N a l n . , nttulb cKoc B BI{Ae Kung. Y runu.{Hrx sp. nov. M 87-149, rlpeAcraBurenefi cnmrHo-6prcuHbre 6opos_ nepxnxft 6ar, n-oa VproHn_ AbI paSBl{"rbr xopouo u npoTnnr.rBaroTcrr TWyc ( x O,8) I{HOnAa nOqTU HA nonoBr{rry An}rHbr poCTpa. B ocranbHoM pocrpbr oAHcyrl{[Hbrl HOMeEeHI{g [apaMeTpOB - H€3H8r{UTenbHbIe. v p"a" pocrpoB na6nrcAaercs He3Haqr{TeJrbrroecMeuleHrre BepulrHbr x cnuunofi snopoHe. C p a a H e H H e . f l o $ o p r ' r ep o c r p 6 n n s o r x p . n e s c i a s p . n o v . , HO OTnHqaeTcg veuuuiefi yAnaHeHHocrbro(l-la y p.nescia 47O_7OO), nr{ue BbrpaxeHHhrMI{ cIIt{HHo-6oxoeuvru 6opo3AaMH. flo eHaqeHnro napaMerpoB K onucaHHoMy Br{rty 6nnsox
N 87-150, N e8 7 - 1 5 1 , Ne87-152, N 87-1 53, n-oa Ypnnr.-Tyrrayc [-oB Yprcnr.-Tyruryc n-oB VpronrsTyMyc Aua6apcran ry6a 5 5 , O( 5 9 2 ) 43,4(466 ) 32,5(S+g)(60)
66,O(694) 5 2 , O( s + t ) 3 9 , 5( 4 L 6) ( 6 0 )
47,O(540) 4 O , O( 4 6 0 ) 34,O(390)(tZ)
65,0 (541) 5 7 , O( 4 6 3 ) 4 6 , 0 ( S t+ ) ( 7 1 )
9,3 ( 100 )
9 , 5 (1 o o )
8,7 ( 100 )
L 2 , O( 1 0 0 )
8,2(88)
7,8(82)
7,3(83)
10,o(81)
3,6(41) 25 25
29
63
P.ishmensis Gust., roroparh cFrrut.tasrcr or P.manif esta sp. nov. y3KoKour{trecxofi Sopr"rofi n 6onee npl{Tyntrenuofi Bepur[Hoft. Qaq Ha nbHas nptl yp oqeHHoc rr. Pocrprr eroro B[rAaBcrpeqenbr rrcKJrrorrETelbHo B MerKoBoAHbrx rfilrHrfcro-arreBpuroaux $auusx ceBepa cpeanefi cu6rpn. p a c n p o c r p a H e H r { e . E a r C e e e p H o fCi r a Boepacr u reorpaQtlqecKoe 6npu. 2 pocrpa lrs HExHero 6ara u 9 pocrpoB us Bepxnero 6ara [-oBa Marep[an. Vprcr+-Tyruyc, 2 pocrpa Es HI{xHero 6ara n 3 pocrpa ns BepxHero 6ara Axa6apcrofi ry6u, I pocrp r{s Bepxnero 6ara p. Onerlex - c6opu T.H. Hansnnenoft.
P a r a m eg a t e u t h i s i s h m e n s i s G u s t o m e s o v , 1 9 6 0 Ta6n. XI,$ur. 1-4 Megateuthis (Paramegateuthis) ishmensis,'flcror'recoB, 1960, crp. 191, ra6n. 44, Sm. 4,5. Mesoteuthis aff. rhenana Mocxanenro, 1968, crp.31, ra6&VI, $rm. 1-6. Ne 254N1 -L26. feononrrecrufi ruysefi M|PH, r.Mocxsa. Pexa f onoran I4ro,ra, Hnxxuft rennoseft. Pocrp cpeAnuft, y3KoKonr{vecKnft, yruepenHo nurranyrufi (IIa 35OD,uarHo3. 418), cxar c 6oKos. Beprunua cflefKa cMeueHa K cflunnofi cropone. Crnuuo-6oKoRlre 6opoenrr BbrpaxeHbr xopoluo, 6pounras Bbtpaxeua cna6o. BueuHHe n p H s H a K I { . 3 a o c r p e u u e K B e p u r { H en o c r e n e H H o e . E o r o B b r e r { c n r H Hag cropoubr npgMonngeftnsl, 6prounas K BepuHHe cKoureHa, or gToro BepuI{Ha cna6o cMerrleHa K cnr{HnoMy Kpap. Cnunn>6oKoBbre 6oposau KoporKue, } orfl€JlbHbrx eKseMnJrspoB oqeHb qerKue. Bp6ursag doposaa qacro cnrraxeHa. Bepuuua (ra6n.22), llonepevnoe cerreHue saocrpexHan. Bepu-ruHnsrfiy:.on paBen 28-350 pocrpa no ecefi Anune pocrpa - oBanbHoe, cxaroe c 6oroe, EE 82-89. n p H g H a K H u o H T o r e H e s . A [ b B e o n a H e r f l y 6 o x a n ,c o c r a B BHyrpeHHne nser oKono L/ 4 anunbr pocrpa, cna6o HsonHyraR. BepuruHa crrenKa cMeueHa K 6prournofi cropoHe. EproruHoft paAfiyc y BepunHbr anbBeoJrbrcocraBJrser 35-42V" CB. Ocesas nr{Hua npoxoAr{r noqrl{ no ueHTpy. Pocrpbr Ha Haqanbxbrx craAuqx
Ta6 nuua 22 t uthis ishnensis Gustomesov H s v e p e u u s p o c r p o a P a r a m e g ae
)s 87-118, flapar"rerput
Inuna o6uas flnnxa
npeanonaraeMarr { ycraHoBneHHas I nocneanbeeonnpHoft
Ana6apcran ry6a
l $ 87- 12 0, Ana6apcxaa ry6a
1 O 3 , O( 6 6 4 ) 8 4 , O( s 3 6 ) 63,5 (4O9 )
8 3 , O( 5 9 3 ) 70,o (500 ) 51,O(364)
15,5( lOO)
1 4 , O( 1 O O)
1 3 , 8 ( 8 9)
1 2 , O( 8 5 )
qACTH
fluauerp cnrnno-6pr,ournofi y Bepur{Hbr anbBeoJrbr finauerp 6orosofi y BepuHHbt AIIbBEOJIbI
Pa.qnyc 6pnuruoft y Bepuuubr
6,0(42)
antrBeOJIbI
Vron anbBeonspHrrft B cnr{Hno6prouHoft nnocKocru, npAA. Yron BepuuHnbrfi e cnuuHo6proruHofi flttocKocrtl, npaA. 64
24
j I
1 J
l)a3BHTI'Iq cy6KoHprqecKHer
BbtraHyrbre.
[IpH Ar,raMerpe 4
ru'"r
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P n c. 27. flpoaonbnoe ce_ qeHue pocrpa parante gateuthis - pressa Naln., sp. nov. N 87-140, aepxHnfi 6ar, AHa6apcxllfi sanxe - 3 a r"re q a H r{ fl. f'o cna6of, cTene'u paoBnTus nocfleaJrbaeonnpuofi qacrg pocTpa P'pressa MO)|(er 6ura conocraBJleH C IIpeAcraBHTerurMa ceMefictga Belemnoteuthidae. OaHaro pocrp p.pressa Bce xe r{Meer ceoftcreenn}rro BceM Megateuthinae nocneanbBeonspH)4o qacrbt xorff ll KoporK]'Ior !t Hbrx ocHosanufi He BKrloqarb paccMarpaBaeNrslft no3ToMy Mbr ne BlrAlrM cepbegsEq B poA paramegateuthis. Oaur.ranbHas nptlypoqeHHocrb. Bce HaxoAru p.pressa cAelanbr B MerF KOBOAHbT*aneBp'Toaux saunsx AHa6apo-XarauncKoro pafioxa. BospacT w neot.pa0nqecKoe p p na sHuer H r -avc" ln' vp vorcl r, a e 't r e . u Clp e q g r f i _ a e p x n r f i 6ar ceaepa Cn6npu. Marephan' 4 pocrpa ,ns cpeA'eno 6ara u s pocrpoB n3 Bepx'ero 6ara n-oBa YproHr*-Tyrayc n Aua6apcxono 3anr{Ba _ c6oprl T.H. HansHgeBofi.
Nb87-136, n-oB YproHrTyr"ryc 25,O(400)
r4,o(2221 5,O(79).(L7l 6,3 ( 100 ) 5 , O (7 8 )
ts g7- 137,
l tte8 7 -138, rr-oB YponnTyr'ryc
]V e87-139, g-OB Ypnnr.Tyr,ryc
4 5 , O( 4 5 0 ) 4 5 , O( 4 5 O ) 8,2(82)(18)
4 5 , O( 3 4 6 ) 4t,o(315) 2L.,O(rgo)Gt)
3 2 , o(4 5 6 ) 2 9 , O( 4 L 4 1 6 , 0 ( 8 6 ) (1 9 )
1 O , O (1 O O )
1 3 , O (l O O )
7, o ( t o o )
7,O(7Ol
9 , 4 ( 74 )
3 , 7( 5 3 )
47
45
AHa6apcxnft 3AnI{B
2,5(40) 35
40
7l
P o a I l o l c o b r l u sS t o l l e Y1, 9 2 7 ( , a r t a l i . u l a f(i p 1 r s ): d ' O r b i g n y ,1 8 4 2 ,p . 7 : l ; Q u e n s t e d t 1 , 8 5 1 ,S . . ] t ] ( rl;) e s l o n g Z i o t e l , 1 8 9 5 ,S . 5 0 5 . S . 6 4 0 ; l 8 8 . i , , . b O ;M a y e r - l ' . i n r a r , r . l r a m p s1,8 7 7 O ( ) t , l i t t d r o r e i t l t(i ps a r . s )[:] a y l ee r Z e i l l e r ,l B ? 8 ,p l . 2 9 ; S t o l l e y ,1 9 1 1 , S . 1 7 5 ; L932, S r o l l e y ,1 9 1 9 ,s . 5 l ; l l i i l o r v - T r u m m e r1, 9 2 0 ,s . 1 9 6 ; K p u v r o n u r , crp.39. g r u p p eL: i s . s a j o u s1, 9 2 5 ,p ' 1 0 , 1 9 1 5 ,p ' l 7 ' .lcrot'oelitcs,2 .lt t lac ot c t r llt isN: a e f, 1 9 2 2 'S . 2 { 5 . l l g l t . o b e l r r .Ss t: o l l e y , 7 9 2 7s,. I l ? ; K p u r v n o n b u 1' 9 4 9 , c r p . 2 O 5 ; H o g e r , 1 9 5 2 , , 958' crp. 159. p . 7 1 7 ;K p u r t l r o n u u 1 n i t i n t e r m e d i a r eVre n t r a l f i i r t ' l r eS; c h r v e g l e r , 7 9 4 9 , D i e G r u p p ed e r [ - l e l e m n i t em s . 3 0 4 , 1 9 6 1 ,S . 6 1 , l 9 Tl , S . 1 2 3 . , L877 ' eepxuufiaaneHQpaH r p oa a. B e l c rrtrti tensttttti e ril )e s l o n g c h amps T r 'n UHI{. cprrbHo yAnuHeirHsre cy6xoHl{qecKl'Ie, cy6uunuHApt{qecKlae O n n c a H ?re. Pocrpsl noneK OKpyfnOMy cxaTble C 6oXOs unw c 6nUSfttM pexe BepeTeHoBl,IAHbIe, Hnlr BepuHHbl' AOXOlqIIIeii or peqHbtM CeqeHHeM, C 6ptoLuuOir 6OpOeAOft, Har{l'IHalolUefics cnafixoit y Haqana anbBeonbl. qacTH H riacTo conpoBoxAapulefics Ao anbBeotapnoft H pexe 6prounocnpluHo-6oKoBble KoporKue y Ha6npAarorcrr Bepuuusl He scerAa B npl{3aryxapqrle 6oroerre 6oposaul. Ha 6oxoeblx crop6Hax ABe napannenbHble K OpOTKI{e' o r H o c H T e n b H o p o c r p b l c r a A H t l x BepuaHHoii .racru nonocbr. Ila saqanbHbrx cy6xouw.tecKIte. B,qoeofr
cocran.
Oxono
10
e 3anaAuofi Eepone'
ellloe
Ha Kaaxa3e,
nTyprveflrruuHHAonesuu'HaCeeepeCCCPBbI[eJuIIorcs3BIIAa, MegateuthiCpa nHe Hue. Pog Ilolcobelus BbrAeJrsercfl cpeAu lloAceveftcrea oT Bepn p o T f l r u B a o u t e f tcs 6 o p o e A u l , 6 p n u H o f i p o c T p a x A n u s H o f i H a nae HanwqueM y anb(He BepuHHbl scenAa) r'rrHbr Ao anbBeonepHofi qacTLI n conpoBoxAaputeftcs (po('ylindroteuthidae p o c r p o B s H e u H e i i {ropvre Beonbr cnaftKoft. Or cxoqgblx no ornl'I.{alorcq pa3BI'ITI'IeM y pocrpbl Holt'obelus Aut I.agortiheltt.s a c1, lindroteuthis) 6opoaa H Hanl4qHeM Ha Hat{albHbJx CTaAHSIX OTHOCUTenbBeput{Hbt cnunno-6oKoBr,rx pocrpa. Ho KoporKoro cy6t
H o l c o b e l t rs w n a r e r r s i sT u c h k o v Ta6n. Xll, Onr. 1-2 I l o l c o b e l t t s t t n t a r a e n s i sT y i l t o e ,
1954;
crp. 123,
ra6n.4,
$rar'8'
\
ra6n' 4' Slrr' 8' Tayficxas ry6a' crp' 123, 1954, Tyrroe, f onorl'In. He[3BecTHo' ronorllna MectogaxoxAeHl'{e vt. Vruap, aaneH. yAnuueHuulft ([la orono 1OOO), cnaSo c r l n b H o r p y n H r r f i P o c r p AuarHo3. '{aCru ngqTI'I Cy6UunUnApU'reCruft, cxaTulfi C 6oXOo' Cy6fOHW.teCKufi, B CpeAHefi ua 2/3or BeprxplHbl I'I npoTgnHearcueilcs HarlrlHaplUeftcs c 6proruHoft 6oposaoft, 4/ 5 lla. f l p u o e p u r ' r H H a n q a c T b C I ' I S b H Oy A n I ' I H e H a ' 3 a H I ' , I M a e T nptr3HaKr,r. Bgerunue HecKonbKo cMelgeHa K cnllHuofi cTopoue' BepuHHocTpan' l-la, BeputlHa Or4-Or5 xopouo BblpaxeHHble cnI'IHY eepultgsl ua6npAaloTc' (ra6n.26). Hun ynon Zio qacTl't pocrpa' npunepuruuuofi Ha 3/4 HO-6oxoeule 6oposAbr, npocnexl{BapuHecg 3aTyxarcruan' He Io6oposAa, rny6oXas y3Kan H AoBonbHo Ha 6prourHofi cropoHe cTopoHa BblnyKB nepeAnefi qacrg pocrpa'6prcurHan xoArr Ao Bepu1arro, un""ao*,. p o c r p a ' B oxoeble cTopoa c e f t A n H H e nas. cnuHHarr cTopoHa cunbHo BblnyKnas no llapHble napafinenbHbje BblpaxeHbl cna6o OqeHb HLIX Irbr 3aMeTHO ynnoueHHble, Ha
72
Ta6avr.ua 26 Hsvepenas pocrpoB I I o l c o b el u s u n re r e t r s i . sT u c . h k o v flapauerpsl p.Tarr,lnnrau flnr.nra /npeanonanaeMarr o6ruas IycraHoafleHHarr {naHa fiocneanbeeonnpnoft IIACTU
124,; (88o ) > L24,5(Sgo )
L7, p. MyHynyqxax
1 7 8 , O( 1 3 3 8 ) 1 6 8 , 0 (1 2 6 3 ) -\-135,4(1O19)(76)
.(uaverp cnauno-6prourHofi y BepuEHbr anbBeoJrbr
L 4 , o ( 1 O O)
13,3(1OO)
Auar.aerp 6or
1 O , 2( 7 3 )
L 2 , o( g o )
{naua [prdBepuuHHo[ .{acrrl fluaverp cnnnuo-6prcurHofi e npaBepu[HHofi qacrra Aaalaerp 6oxoeofi B npr.rBep_ uruHHofi qacrr.r Ynon anbBeonspHslft B cnHH_ uo-6pourxoft nnocxocrn, npaA. Vnon BepuJHHHblft e cnunuo_ 6prourHor?nlrocKocrtl, npaq.
5,3(38) 53,5(382)(43) 1 3 , 6 ( 9 7 ) ( 1 O O)
5,3(40) 67,5(so8)(+g) 12,8(96)(1OO)
11,6(83)(8s)
1 2 , o( 9 0 ) ( 9 4 ) 22
27
nonocbr' 3aryxarculre fipu nepexoAe K npuBepruuusofi qacrr.r pocrpa. rlo_ nepeq'oe ceqeHl,re pocrpa oBarrb'oe, c,traroe c 60ron, np"r"r-"T€ne'b cxarr'rs 60rsure B nepeA'efi qacru pocrpa (EE ?3-go, 66 g5-g4). B n y r p e H H U e n p u e H a K u V to H T O , . e H e s . A r n s e o J l a3 a ' L r M a e r oKoJro 7/5-7/4 AtrrHbI pocrpa, BepuuHa ee cnenKa cMerrleHa x 6p'uHofi cropoHe (6proruHofi jaArryc cocraBneer oKoJro o,4 cE). Ansaeo_ napHrrfi y.on paBeH 220. oceeaq m''"s, crer*a H3.nrry"a u npu6nuxe_ Ha K 6p'unort cropo'e. cnaftra y Bepu'lHbr anbBeonbr Ha pocTpax lr3 Harueft KOJTJT.KUI{U orcyrcTByer. Ha Haqarr'brx cra'asx pocrpbr, KaK BI{AI{O Ha puc. 29, orHocl{TeJrbHo yKopotre'Hbre I{ cna60 cy6xonuvec_
:H;r:3"t"""t"
T[r11
7oo' npucE 11,s fia oronorooo, .,prn
He Nre Hqr{ Boc r b. orpaHuve'Hocrb MarepuaJra He fiosBorrseT cy_ Eurb o6 usvextrllBocrlt' lloabepxesa roae6a'r{rrM orHocrdr€rlbHas AJrr{Ha 6prorugoft 6oposau - or rrpuMepHo 2/3 ao 4/5 f1a. Cnafira B Konue anbBeorrbr' KOTOpas Ha6ruoaanacb v.v.Tyrxo"o* Ha onuca'Hbrx r{M pocrpax, Ha gK3eMnllspax ue Hauefi KoJrrIeKuI{r{orcyrcrByer. CpaBHeHue. OnucbrBaeMbre pocrpbr or pocrpoB Ap)rnux cu6upcxux BUAO' Horcoberus pesK. OTrr'qarorc.s ceoefi cy6rouuvecrofi {oprraofi u cxarr{eM c 60Koe. or sanaA'OenponeficKl{X H KaBKa.CKLTXBAAoe erono xe poAa paccMarpunaelotfi Bua orJr'qaercs'vensruefi npoTrrxe'Hocrbro 6prourHofi 6oposarr Ha pocrper H€ Aoxoa.gruefi llo Bepuu'br arrbBeoJrbr, u 6onee 3HaqI{T€JIbnofi orsoarr=*"ofi yanuueHHocrbro no cpaBneHuro c 6rnasxuv no o6uefi fiopve II.ntunier; (D".r6,rg."-horp", rg77,crp.63, ra6n. 5, Slu.. 3-6; Ta6n. 6, dru.. S_11).
llr Pnc' 28' flporoasHoe ceqeH'e pocrp a Holcoberrts ttnarersrs 'l-uchko' Ne 87-2 17, suxuufi aarre4 p. MyHynyA)rex >rO,"l f
lii
t7 73
HaM pocrpbt H.unaren
H o l c o b el u s g r a u i s ( G u s t o m e s o v ) Ta6n. XVlll, {ur.5-6 I . e r t o h e / 1 sg r a u i s f y c r o u e c o e , Sr.rr.J.1.
1966,
crp. 6?,
T a 6 n .6 , S r r . 4 ,
5,
ra6n. 7,
Mysefi MIPH, Mocxaa. PeKa MonoAor aanex? NeVl -141l3. lonorHn Pocrp ue6onuuofi aepereHoenAnslfi, cxarrlft c 6oroa, c 6pouuoft AnarHo3. 6oposaoft, HaqnHapuefics y Bepur{Hbr H 3aKaHql{Bapueficq He AOXOA9IAO BepUt{HbI anbBeonbl. ' B H e u H l r e n p H 3 H a K u . I - l p u n e p u r n H H aqna c r b c o c r a B n s e r o x o n o O r 3 f l a , BepuHHa 3aHHMaer 6nusxoe K ueHTpanbHoMy nonoxeHue, 3aocrpeHa' BepuunHslfi (ra6n.27\. BepereuoBrulHocrb pocrpa Bblynon HaxoAr.rrcqB npbAenax 42-490 t60-L'76o,/" CE npu cocraBJultor c6 3uaqeulls orqernllBo, paxeHa Aocraroqgo Eprcunan 6opoeaa y3Kaar xors H uenn$oran noKa3areJyrx na oKoJro 5'7O-77O. B npuBepurunuofi qacTu pocTpa, Aanee paculHpqeTcq t{ npuMepHo B CepeAuHe nocneanbBeonapuofi qacTH pOCTpa BbrnonaxuBaeTcq H nepexoAt{T B ynnoueHHe' KoTopoe npocnexyBaercrr noqTu Ao anbBeonspuofi qacrll pocrpa. CnuHHas cropoua no acefi AnuHe pogrpa Bbln)tKilas n nn1IJeHA6OposA. BOXoaule CTopoHbl HeCKOnbKO MeHee BbrnyKJrbte,HA HHX eABa HaMeqapTcr{ nap}rble nollocbl' 3aTyxatou}Ie B npI'F BepuuHgoii .{acr14 pocrpa. fionepeuHoe ceqeHl{e no scefi qnuHe pocrpa HMeer {opMy cnerKa cxarono c 6oxoB oBana. Creneua 6oroeoro cxarus Ha BceM nporqxeHr.tHpocrpa ocraercn npt{MepHo oAuHaxoeoft (SS Sg-1OO, 66 88-97). n p H 3 H a K H H o H T o n e H e s . A n b B e o n a3 a H l l M a e ro K o n o t / 6 BHyrpeHrrue (6proruJ,nurrbr pocrpa, BepuprHa ee rre3HaqllTenbHo cMerueHa x 6poLlHoft cropoHe nt4wP.s' ynon Oceeas He 3aMepqncs. AnugeonqpHulfi CE). soii pa,quyc oKono Or4 TaKxe He3HaqprrenbHonpu6nuxeua r 6prouHofi cropoHe. IOstte pocrpbt HMenI{ cy6uunugllpl{qecKyro fiopvy (puc.29) u ronsxo npu npu6auxeHul{ K pasMepaM Bspocnblx e}IAY BepeTeHoBlraHocTb. oco6efi pocTp npuo6pera.n cnoficrBeHHylo OnHCblBaeMOMy Hverouteroctt B HBHev eHr{ uBocrb. urer"t KonneKum{ MaTepuana HeAocTaToqHo, qro6ul cyAurb o6 HsveHtIuBocTI{ paccMarpHBaeMoro Buaa. Pocrpsr, onucaHHbre 6onee KpynB.A. tVcroMecoBbrM (1966),
P N c. 29-. flpoAonuttoe cer{eHlle pocrpil Holcohelus gravt s (Grrstonresov). N 87-2 15, ltuxttttii aaJIeH' p.KcnnrtRP(xO,9)
uu vv nporl{B 7,2-812 Hbte (c6.9,6 y sKgeMnnrrpoB r{s Hauefi xonnexqun), HecKoJrbKo 6onee yKopor{eHHble ( fla r.aeHee no no 600, y Hauux oKono 7OO-77O), (c6 o6ureft ,bopve noqru He ornl{qardrcq L75% C.E, y Haur.rx 160-l-76V"). C pa B HeH I{e. flo ceoer'r BepereuoBttanofi rfropve pocTpbr
74
ll.gluvis
pesKo
BbI-
Ta6 nwua 27 H s v e p e H u r r p o c r p o s H o l c o b e / r r s g r a z , i . sG u s t o m e s o v
N e8 7 - 2 1 9 , Ana6apcxan ny6a
llapauerpul
J V e8 7 - 2 1 4 , p.6yop- 3exar
Ns 87-215, p. Kenuvrnp
finnHa fycraHoa55,5(1233) 5 4 , o ( 1 2 73 ) 4 7 , o ( 1 O O O) o6uras InenHan Annna nocneanbBe- 5 s , 2 ( t z z 7 ) ( 7 6 7 ) > 5 4 , O ( t z z 3 ) ( 6 6 7 ) 4 7 , o ( r o o o ) ( 5 7 3 ) onnpuoft qacra 4,5 ( 100 ) flllauerp cnHHHo4,6 ( 1oo ) 4 , 7( 1 o o ) 6prourHor?y BepulrHbI anbBeonbr 4,5(100) flr.raverp 6oxoeoft 3,8(83) 4,6 (98 ) y BepultHbt anbBeonbr Paqnyc 6pnuHoft y 2,o(43) 1 ; 8( 3 8 ) BepuHHbranbBeonbr Anlrna npunepu_lluHofi 1 6 , 6 ( g g o ) ( 3 0 ) 2 O , O ( 4 3 5 ) ( - 3 7 ) 1 8 , 9 ( 3 8 5 ) ( < 3 8 ) qacTu /i[uaverp cnnHHo6proruuoft B Mecre uan6onr,ulero yroJtueHnq flnaverp 6oxosoft B Mecre Han6onsuero yToJueHr{rr Vron oepuuuHurft e cnlmso-6or
7,2(160)(1oo) 8,1(L76)(100)
8 , 2 ( L 74 ) ( 1 0 0 )
7,O(1s6)(97)
7 , 8 ( 1 6 6 )( 9 s )
6 , 8 ( 1 4 8) ( 8 8 )
49
42
Aeneldrcfl
cpeAl{ ApyI'HX BI{AoB poqa Holcobqlus a npu6nuxaorcq K pocrpav Hastitidae u oco6enxo pseudodicoelitirrae. B.A.'fycroN{ecoB ( rgOO) u oruec anA gravts K po,Ty Lenobell9, pocrpbr Koropor.e raKxe HNretor qnuHrryto Har{uHa1guyocrr or BeprljHusl 6ptotuuyo 6opo3Ay. oAHaKo B orJu.rr{He or Lenolteltts H.grauis He o6naAaer 6oposqau' B flepeq'efi rtacrl't pocrpa - npH3HaK, HcKnor{aouprr-r or}rece'ue 3rono BIIAa K Haqce:rreiicrBy Duvaliaceae, B Koropoe Bxoqsr pseurlOrlic1,1elitinae q u c n e rr B r.tx pol, I-€nobclus. @auna JrL,Hafl nprypoqeHHDcra. Eonsr'arr qAcrb pocrpoo II.gravis (c rK3eMnnspoB, cnncaHHbrx B.A. lycrolrccoBr,lM f{erpM ) coOpaua B necr{arrrtnH_ HHCTLIx NlenKoBoAHslx {tauttqx n6nt't.}tt no6epexbq topcKoro MarepuKa rra rgBrrrX npHToKax Ileuul tI y ycrbn Aua6apa. OraeabHbte rraxoqKa orlterrarorcq B nnHHHcrstx tbauufl'\ orKpbfrcl'o Moptl n JIeHo-oneuexcKov pafroHe. Moxrro npeAnonararb, r{To xr{BorHoe rrpeAnolrHTaflC npu6pexnr,re MenKoBoAHbre o6craxogKl{, rAe, cyAn no yAnI{HeHHocrI{ tI BepereHoBilnHoerH pocrpa, Beno cao6oAHonnaBarcuHfr o6pas )xTI3HH. Bospacr cr6Hpu. caIItIbIr
-1
t' reorpa,t,tr,recKoe pacnpocrpaHe Hne. Aanex ceaepnoii Yxa,airxne B.A. l;'croMecoBa (1g66) g paHner.o,pcKgtr.rB.Jspacre en!r_ ,{t\1 c p.Mo'ncao pocrpoB ti.grqtis t s b r 3 b r B a e r c u . r r b H r , r ec o M H e H u r r p n o c K o f l g -
rpadrc 'Aruua HHe na. B
nocneatrbDeorrrpHol-r qacrxo
Bo Bropblx
cxo6xax
aaHo 3Ho9e_
75
cpasy creKy no [aHHbrM T.V. Kupnnoft Ha p. Monoao HaA nnugc6axov ayer aaneH. 1 pocrp Hs Bepxueno aalreua Aua6ancxofi ry6u - c6opu T.H.HansMarepuan. u KenHMsp - c6opul uReegft, 2 pocrpa H3 HuxHero aaneHa pp.6yop-3erplr T.H. HansHseeoft u T,V. KuPnuoft. i achs,.p. nou.l I l o l c o b e l r r sA i r t a s o t , S Ta6n. Xll, $un. 3-4 B e l e m r t i t e ss p . ( c f . b e y r i r : f t 1 ) : P o m p e c k j1, 9 0 0 , p . l 0 : 1 , p l . l , t i g . 2 2 , 2 3 ' D a c t y l o t e u t h i sc [ . i n c u r t t a t u s : B o p o u e u , 1 9 3 7 , c r p . 5 8 , r a 6 n . 2 , $ v r r , 2 0 , Holcobelrrs sp. : Caxc u HansHReaa, L967 2, ta6n. 3, {ur.3, KuHacoa,1968t ra6r.69,
crp. 134,
Srr.2.
M A7-218. Myeefi Hful CO AH CCCP. Hosoca6rpcr. llonyocrf onorun poB Ypronn-Tyrulyc, BepxHvrfi roap ( ? ). Pocrp cpeAnr.tii,cy6ur.rnr.rnapruecruii,yAnuHeuHufi (fla or
1 0 4 - 1 0 s) . u o H T o r ' e H e 3 . A n s a e o n a 3 a H L I N { a eor K o n o L / 5 npH3HaKH BnyrpeHHHe (6prouuol'r pa,quyc npsMarr, BepuuHa cMeueHa x 6ploLuucfi cropoHe Anr.rHbr pocrpa, 2 4 o . y n o n p a B e s O c e saq ntIHI{g cnerA n u e e o r u { p u s r i i C E ) . 35-4I7o cocraBnrrer Ha HaqanbHbtx cra,:ltlqx pocrpbl x 6prouHoir cropoHe. Ka r.I3orHyra u npu6nuxeHa orHocaTenbHo yKopoqenbl. flpu CE 4r5 vrt fla HMeror cy6xoHnrecKyro fiopvy, go 7OO, npuqeNt pccrp nplto6pepaBHo 4OO, npa CE - 7 12 r"ru fla Bo3pacraer yxe
raer
cy6UunuHaprqecKyro oopt,ty. flpu lanuHer"ruet\t pocre Bo3pacraHua 3Haqettlts l-la (puc. 3O).
xtlBoTHono
IIe
npoucxoAr.rr 3aMerHono HsveHr{HBocrb.
Pocrpsl, orHeceHHbre K onncbrBaeiuol\iy Bt.IAy, ornllqarorcg 3HaqeHHflMlI H 3 M e H q l t B o c r b r o . Hapn.ly c cplnbHo yAnHIIeIIIIbt\llI pocrpaMl{co 6onuu"roft (ronorun, rr ep.) pocrp, onucarrHbtir B.ll.Kttttacoentrt, BOO-1OOO fla'oxono Bcrper{a1grcrr pocrpbr NreHee yAnHHe}Hbre, c fla rrerree 7OO (pocrp Nr 85-19). Ecra pocrpbt cy6ur.rnr.{traprmecrofi $opvr'r, ,xaxe c ileKoropbt\t cyxeHlleNl Ba)nu:lll r r e c r b p o c r p F , rr c n e n K a c y 6 x o t t t ' n r e c K t l e , c y A n t l BepuuHbt anbBeonbl (ronoran) u e H x o f i n p l r B e p r l H H H o [ " rq a c r b r o ( p o c r p \ s 8 7 - 2 1 9 ) . Pocrpul l/./rirra.soi,i ornr.larcTcn CpaBHeHHe.
cu6ttpcKtlx RltAoB or.lpynux Ilolcoltelrrs ceoeli cy6uunprHfpr.rqecKcr'r $oprroii tt uantmtleM cntttttto-6proutttono Il.umurertsi.s n o c n e A H r , r f i n p u s u a K t { e r K o o r n l t q a e r l l . l ; i n u . s , r . - / io r cxarr,Iq. 3ror crp. 1 18, Tuclrk., a raKxe or 3anailrroeaponei'rcroro Il.trauthi \Stolley, 1927, poAa
ra6n. 24, fivrr, 7-8 ) Cne.lyer H.C. Boporreu 'l *
76
orrrecrH ( 1937
Ha3BaH Bua BIIAA.
K paccN{arpHBaeMo\Iy
BIIAy u lrenoru{r'lir pocrp, cnttcaxHr,ti'i cKopoee Bcero tI:} aaJtella Bcp-
) H s H H o u e p a N { o B F , r xc n o e B ,
B qecrL,
B.fl.
KlrHacoBa,
Brl€rpBbte ontrcaBn.rero
pocrpbt
AarrHotr.)
Ta6 n uua 28 Hevepeuus pocrpoaH olcobe lus h i n a s o v i
llapaverprr
f-nRtatto*Anusa J naevaa o6rqas )'"ru"ouJ [*neHuan finuHa nocneanbBeorspHoft qacrr.r AAar,,refp CnI{HHO6prouHofi y BeputlHbI
N b8 ? - 2 1 8 , rr-oB Vprorr.Twvc
Sachs, sp. nov.
N e 87-2 19, p. Kennvep
1 3 5 , O ( l _1 5 4 )
Nb85-19, p. Tarllntltau
1 4 O , O( 8 4 7 |
77,6(978 )
96,0(821)
77,6(987)
-110,(94O)(ef)
7,8(1oo )
tL ,7 ( 1OO)
L6,1 ( lOO)
8,1( 104 )
l3,o(r11)
L 7 , 2 ( 1 O 7)
92,5(575) -111,O(689)(zg)
ANbBEOJIbI
Auavrerp 6oxonofi y BepuuHbr anbBeoJrbr Paanyc 6pnurHofi finnna npnaepuruuHofi qacTI{
2,8(36) 32,2(arS)(4L)
4 , 8( + r ) - 5 2 , o( + + + l ( 4 7 )
5,7(3s) 34,5 (22c-1(31)
fiuar.aerp cnr{Huo6prcuruoft B npr.lBepungsoft qacrr{
7,7(9e)(100)
1 1 , 4 ( 9 7 ) ( t o o ) L4,4( 89 ) ( tOO )
luarvrerp 6oxoeofi e. npuBepur{HHoft qacra Vron arrbBeoJrspHuft B crruHHo-6proulrrofi nnocKocTl{, trpFA. Ynon aeprurnnrlft n cnuHHo-6pnruuofi IInOcKocTt{, npan.
8,1(1O4)(1Os)
1 1 , 9 ( l O e ) ( r o + )1 5 , O ( 9 3 ) ( 1 O 4 ) 24
24
xoBbeB p'Eyper norl HasBaHI{eM Dactyloteuthis cf. incuraatus Zieten, Oror pocrp LMeer o6uryro Sopr*ry, cr(oAnyrc c T14n1atrgbrMnoKseMrr,,spaMn, fla 6onee 53o, E6 (e6nnor{ nepeA'erp Kortrra pocrpa) orono 1ro. MoxHo rlpe[rro,roxl{TerlbHo orgecrr{ x H.hinasovi n o6nouarurbrf, POmp' cn,'caH'brfi na aane'a 3eunn opa*r6-l40cn@ H.fbMnerIKEM (Pompeckj, 19OO). O a u rr a n b H a s rr p lay p o rr e H H oc r b. pocrpu H. h,inasoai scrpeqa_ Iorcs B lnEHucrsx Saunsx orKpblt\orc Mops a IIeIroForreHeKcKoM paftoHe' B Beprornse r{ B o:rorcxo.Konsnacxov pafione rr pexe B necqano_ ntrl{Hllcrbx saunrx Ana6apoXaraxncxoro pafiona. cneay€r,r crrnTaTb, trro xl[BstHoe Beno cso6oaHonnaBablrlrfi o6pae xr{srrrr n lpearroqmaflo o6c:raxosKr{ orKpbrroro Mops, Boepacr u pacnpocrpaHeanne. Hrxnufi leorpa$n'recKoe Toap (eoua Dactylioceras commune) - ."rr.i., ,o o.*""* B.fI. Knnaco Ba TaKxe 6afioc ceeepnofi cn6npn, npeAtrono)Kr{TerrbHo aareg Aamnero BocroKa u 3evnu
Pnc. 30. llpogoasnoe cetreHr{e pocrpa Holcobelus hinasoul Sachs, sp. nov. lt 87-19, aepxrurfi roap, p. Tatatnnuan
l l 1
t,!'
l ' l
77
3 pocrpa us HuxHeT.o roapa-aSneua Oxoncxo-Ovononcxor\o Marepnarr. paftoHa - cSopsr A.C. Aaruca, 2 pocrpa H3 Bepxuero roapa-aaneHa pp. Kenuunp u Eyop-3eKHT - c6opu T.H. Harn usesoft u T.h. Krpunoft, 1 pocrp u3 BepxHeno roapa (?) n-oea Yponr*TyNryc, 1 prcrp u3 aaneua Axa6apcrofi ny6u - T.H.Kupuxofi.
CEME IiCTBO CYLINDROTIIUTHID SA TE O L L E YI,9 I 9 tt B ToM rtl.Ic[e cpeAHeppcKru( Onucauue Br{AoB ceNaeftcrsa Cylindroteuthidae BI{AoB poAoB Cylindroteuthis u Pachytettthis AaHo aBTopaMH a pa6otax 1964 u 1966 rn. Hg 6arcrux ornoxeHrrfi Ceeepnofi Cra6rpu onllcauu: Cy lindroteuthis (Caxc ( C y l i n d r o t e t t t h i s ) , O w e n i O u , e n i ( P r a t t ) , C . ( C . ) . s p a t h i . s a c h ^ se t N a l n j a e v . a u HanunReea, 1964), Pachyteuthis (Paclryteuthis) o|tima Sachs et Natnlaeva' r. ( P . ) p a r e n s S a c h s e t N a l n j a e v a , P . ( P . ) s u b r e c J i v i u a( L e m o i n e ) r P . 1 P . 1 b o d y l e u sAii Sachs et Nalnjaeva.(Carc n Haru,HqeBa,1966). B 6afioce KanqAu l&sBecrnbl p a c h y t e t t t h i s( P a c h y t e u l f t i s ) s p . n o y . i n d e n . ( C a r c n H a n s u R e a a , 1 9 6 6 ) , C y l i n droteuthis tltemis Crickmay. Huxe Mbt npl{BoAl{ryronficaHue HoBoro BnAa CylindroC.(C.) sp. teuthis, xoropsrft yx<e ynoMuHancc a pa6ore 1966 n. noA Ha3BaHHer',r s p i c z I a r i s . g r . nov. ex
STOLLEY,1919 CYLINDROTEUTHINAE NOACE/VIEfiCTBO P o q C y l i n d r o t e u t h i sB a y l e e t Z e i l l e r , l B T B I-loaPoa .
C Y l i n d r o t e u t h i ss . s t r .
Cylindroteuthis (Cylindtoteuthis) conf essaL Nalnjaeva, sp. nov.
' fa6n.
X r 0r.
1-3,
puc. 31
Ne A7-L57. Myeefi Hfnl CO AH CCCP, Hoaocx6trPcKr'Pera 3exur lonorrn (6acceftH p.Ilensl), nepxutlft 6ar. Pocrp cpeAnnft, crrrrbHoeslraHyrrrfi (na 11OO-12OO), cy6uunrnAnarHos. OceAprrrecrufi. flonepequoe ceqeuue oBanbnoe, cxaroe c 6oroe (EB 77-86). BAn nmttlg noqTH IIeHTpaJIbHas. BHeurHr{e npBsHaKtr. Pocrp oqeHb nocr€neHno cy)t$Baercs K 3a.qHeMy KoHuy. flpnaepuuxnas qacrb cuJrbHo€brrgnyrag, cocraBnser oKorlo Ll4 unrxbt. Beprunna uenTparrbnas, crrlrrruo aaocrpennac, Bepul{HHblfi yron paBen 17-190 B nolepequoM (ra6n, 29r. V aepurnuu Mopusnrn r 6opo3Abr He xa6noAaprcs. ceqeg11u cnHHHag ]r 6pprxuas cropollbr Bbln)ncJlbte, 6oroabte cna6o yn[ouelrbl. [Io' repeqHoe c-eseHl|e y BeputfHbt alrbBeorrbr oBarbHo€, cxaToe c 6oroe (sn z5-83), B flpr{Bepurruroft qacrtr cranoBl{Tcs 6olee oKp}'TJIbIM. npHsHaKH n oHToreHes. Amgeona npsMasrHerny6oxaa, BnyrpeHHr{e cocraBJlser oKorro 1rl5 Aflf,Hbt pocrpa. Bepurnna aJIbBeoIIbt pacnonoxeEa nslTr 35% CE. AnauegTparuruo. Eprorunofi paqnyc y Bepul{Hbr anbBeonbl cocraBflser Beonspgbrfi lron paBen 2Lo. Ha naqanbrrbrx craAEtD( pocrpbl csunnunaprrecrofi 13OO, T.e. flpn nnar'rerpe 4rS r"t"r IIa qocrrraer Qopr"rsr,Bbrrrruyrbre (puc.g1). H@KoJIbKo 6oruue qeM y repocrlbtx oco6efi. e Bq 14Boc r b. HeAocraToqgocrb Marepnana He rlogBonFer noBopl{rs o6 14s 19r noMenqltBocru rrpr{suaroB y ontrcarrnoro BI{Aa. I4r.reroutrecc B Korutexurru pocrpbl AnI{HbI }I creneHbKt oAHorrmHbr u orlrr{qatorcs nrub rone6anneM orltocurengnof, 29). Pocrpu, co6paunue a iuxnerennopefi6oroaoro cxarus pocrpoB (cu.ra6n
*1
Confessa (nar.) - 6eccnopHas.
78
Ta6nnua 29 Her'repenna pocrpoB sp. nov
cy lindroteuthis (cy rindroteuthis confe ssaNarnjaeva, )
lr 87-157, p .3 e ru r
flapaverpu [nrHa flnpeanonaraeMas o6rqag lycraxoarrexHas [nrna nodnearraaeonapHoft
1 s 7 , O( 1 3 7 3 ) 148,5(1291) 126,5(11OO)
TIACTE
flnar'aerp cnrnHo-6prcunofi y Bepu[Hbr grrbBeonbr ,6orosofi y Bepuu_ {uaverp
Orou.rau[e
1 1 7 , O( L 2 5 7| 1 1 2 , O( L 2 O 4 ) 9 2 , s (9 9 4 )
11,5(lOO)
9,3( 1oo)
9,5(ag)
8,o(86)
Hb.I AJIbBEOIIbI
Paauyc 6prorunoft flnnua npaBepuJannoft .racru ,[uaverp cnrsno-6prorunoft a npr{Bepurr{Hxoft qacra Anar,rerp 6oroaofi B rpr{Bep_ urnurof, qaclr{ Anseeonflpxufi yndfl B cnna1o_ 6poruiofi flnocKocrn, fpan. .Bepurruruft lnon a 6orosofi Ilnr)gKc)gaE, rpnrt.
Ne87-158, p. Sexrr
4,o(gs) 4 3 , 5( 3 7 8 ) 8 , 8( 7 6 ) ( L o o)
3,5(37) 3 5 , O( 3 7 6 ) 7, 7 ( 8 2 )( t o o )
8,2(71)(b3)
7 , O ( 7 5) ( 9 1 )
L7
.ra6mruu
flapaverpu
]$ g 7 -1 5 9 , p .3 e rrr
Ite 87-112, YplornrTyrayc
,tlnrsa fnpeanonanaeMas o6uag IVcranoaneHnas l.lrrl{Ha noclearrbaeonrpxofi qaqrr llnavecp cnunno-6prcurnofi y Bepul{lrht albBeorrbr Analrerp 6oroeofi f B€ptrrf,_
163,0( L2O7) 151,O(1123) 130,o(963 )
1s6,ot iz gsI
1 2 O , O( L t 4 2 l
166, (1098) 154, (962 ) L27,5(796)
13,5(lOO)
1 0 , 5 ( l O O)
1 6 , 0 ( 1 O O)
1 O , 5( 7 7 |
9,o(85) 12,O(75) Paanyc 6prcrunofi 4,5(38 ) 7,O(431 Amura npf,Bepurnunofi,racrrr 4 s , o ( 3 3 3 ) 4 0 , O( 3 8 5 ) 48,O (3 OO) .Ilnavetp cnrruor-6prcrunofi a 1 0 , 5( 7 8) ( r o o ) 9 , 9 ( 9 4 ) ( 1 o o ) 1 3 , O ( 8 1 ) ( 1 O O ) nprBeprumroft qacrr .drrarr.rerp6oxoaofi B np[Bep_ 1 O , O( 7 4 ) ( 9 5 ) .9,2(87)(99) 11,O(69)(e+) unnxof, tracr[ Amseongpnufi yrorr B crrrru_ 2t no-6pnurofi nnocxoctrr, rpaA. Bepurnxslf, yron e 6oxoBofi 19 rl,IocKocTl|, rpaA. EbI
AJIbB@'IbI
cKru( orrroxeHrfsx Ha [-oB€ vponrTwyc orn}Iqarorcg 60rgurrr,r 60roauv cr(arrF eM (6875) n 60nee n opo*.oofi nocrearrbBeorspnol vacreo (na=7g6). cpaBHeHre. Har6oJree6nrsox no oHaqe'nrofla u BB c.(c.) spicuraris,spi_ cularis (Phillips) rs KertloB€a- Hrr]rclanorc'Mep,,Axa 3anaq'of, Eaponrr n fuccroft paB'r'br. B ornrtrre or c.(c.) ,orf""iiii-o, BrrAr{Meer pocrp co 3*aer_ 79
Pnc. 3 1. IlponoJrbHoe ceqenue pocrpa Cylindroteuthis -IbUr BeP confessa Nalnjaeva, sp. nov.
(Cylindroteuthis)' ', p. crelo{r ( x
TeJrbHo 6onee KoporKofi npl{Bepurnnofi qacrbro rr scno Bblpaxennofi 6onee M8- . 6prcruxofi 6oposaoft y BepurusI. Pocrpbl , C. (C.) spicularis (Phillifis) rs Kpyrrlrbre. oKc$oPAaPosrpbl C.(C.) obelisca cr{Bnbre I{ Kr{MepuAxa 3anagnofi Eaponu, Fyccxofi pa.Bnf,Hbl u Ceaepo-Bocroxa CCCP u C.(C.) Ienaensis Sachs et Nalnjaeva nelorrxcrono sp)rca Ceaepuofi Cu6npu, cxoAnble no o6uefi 0opr"re I{ ors}l"rcTBrp 6prorurofi 60posAbr, onrocurenbno 6onee yArrrnenbr (IIa 1600-20OO u 14OOa cy6rourvecKylo Sopl,ry. 1600), uMeror ne cy6uunl{HApnecKyp, QauHanbni s npuy poq eHHocru. Ocnoanblecfupbl pocrpoB C.(C.) conf essb. nprrypoqeHbl K pnunlrcro-aneBplrroarru @rurr.,r orKpbh Toro Mops a HugosLax p.Ilexst. Pexe rr ToJrbKo B IruxHeM KerlnoBee Bcrpeqa gr€ rr otrr{c brEaevuft BlrA B 6onee MerrKoboru bt:K [ectraEo-nJrlfHlfctITo xrrBornoe Moxno lyMarb, rrx $auusx Ha n-oBe YpronrsTyMyc. Berro cao6oqHonnaBarouuft o6paa xtlsHu 6uno, cy[a rro yAru{Hernogrs H crpeJroBHAHocru pocrpa, ,@pour[M nnoBIIoM u npeAllottnraflo o6craI HoBKr{ orKpbrrora Mops. Boopacr I{ neoFpa0uqecKoe pacnpocrpaEeHne. BepxCeaepnofi nuft 6ar - Hr{)r(HHfirennogefi (goHa Arcticoceras kochi) Cu6upn. - cSopu Marepran. 8 p o c r p o n u S p a r M e H T b Ir{3 Bepxneno 6ara p.3erur T.kl, KwpuHoft I{ C.B. Meneqrnoft, 3 pocrpa r{s Hr{soB KennoBes n-oBa -Vprcnr-TyMyc - c6oprr T.H. Harunsesoft.
I
(
l
I
I
(
I
H A l I C E ME i i C T B O D U V A L I A C E ASEA C H S E T N A L N J A E V A1, 9 7 0 190?, 1842,STEINMANN, GASTROCOELI ET NOTHOCOELIN'ORNICNY, 1 ,9 4 9B , ELEMNOPSIN AL E T Z K Y ,1 9 6 5 , W E R N E R1,9 1 2 ,S C H W E G L E R JE B E L E M N O P S E I JNEAL E T Z K Y ,1 9 6 6
I
( I I
Pocrpu BepereHoBr'rArrbre,cy6unnnnAplrqecKtre, pexe cy6ronr.recOnucaHEe. Kue c 6opoenar'rr.r B nepeAHeft qacru pocrpa - 6porunofi, y HeKoropslx cer'reftcrs cnunuofi, pexe 6pror:rHofi r{ cnr{HHof f-1tg AByx nap 6oroBux. B arlbBeoJlspnoft .raeTH pocrpa sa6ruoqaercs cnafira, HHotAa nepexoAsr{al B uetlb. Eopoeau B rIpEBepuJItHHoft'racru pocrpa y noAaBlurrouero 6omruuncrBa celteficra orcyrcrB)tlor. 'Opanrvroronbr tlMeror KoHuqecK]'ro 0opuy H no AaHHbIM Eneuxoro (Jeletzky, 1966) xaparTepna)'Iorcs orcyrcrBt{gM KaMepanbHbtx orroxegrfi . BelemnopsidaerNaef, t922i Cerueficrea Duvaliidae Pavlow, 1914; Cocrag. 1967; Dimitobelidae Whitehouse, 1924; BeDicoelitidae Sachs et Nalnjaeva, lemnitellidae Pavlow, LgL4. CpaBHeHne. Pocrpur Duvaliaceae ortrrrarorcs or pocrpoB Cylindroteuthaceae pa3Br{THer.a6opoea u cnafiru B nepeAHeft .{acrH pocIpa. tlpeo6naaaer y Duvaliaceae BepereuoBuAHaq Sopva pocrpa, peAKas y npeAcraeur€nefi Cylindroteuthaceae. @panuoxoHu Duvaliaceae nuruenbl KaMepaJrbubrx orrloxenrfi, ToI'Aa Kar y 1966) Haqanssble KaMepbI Cylindroteuthaceae no AaHHbnuEneqxono (Jeletzky, (xors ecerAa). u He 3anongeHbt ornoxeHugMl{ Sparvoxoga B K f l r o q a e rB c o c r a B o A H o r o [ I 3 c F 3apreqaHtrs. B.A.lycroruecoa (1973) Nreftcre Duvaliaceae, a r{MeHHoBelemnopsidae, noAcer"reficrBoHastitinae c poAaMH Hastites w Rhabdobelus, OcxosaxueM K 3ToMy crryxuT crpoeuue 6orosslx 6oposA, ToqHee nonoc, Ha pocrpax, oKa3brBaroueecg no MHeHuIo B.A. flcror"recoBa cxoAHbrM y co6craeHHo Belemnopsidae n Hastitinae. Oanaro, pocrpbl Hastites u Rhabdobelus nutueHbr 6opoea B nepeAxefi .racrx n noaToMy AotDxnu, Kar( 3To t{ 80
!
t ( I
1
T
T n
( r (
c c u
:::ff:
x ceuer*avHastitidae B %t,,1' ],? "J::#ff"l "::"'".o"o'orbc'I n r e o r p a 0 n q"1.1,: ecKot p"",,pocTpaEeFEe.
BoepacT
$::'--Jnfi
T#T*"in"*
anb'oM rosce (Aa*pan;;'f,ff::f
e p x u H r { x H e Rr o p b r ("u"lp*'i ' nJp.;;;;, Bro,rce, aaaen? -
J"??Ji:*
Bepx'IrftMerrB i",roop.-
CEMEIACTBO DUVALIIDAE P A V L O W1, 9 1 4 on ncaHa e' Pocrpsl Bepere'oBl{A'brer pexe c}dunnrxgp*rec*ne rrec*rre' c nn160roft a cl6xonacnannofi 6_oposaofi, ,o*lrr",o*eftcs or nepeA'ero KOHUa E corrpoBor(aapurefics cnaftK'fi' v naaOo*.ro"."* rpeAcraeureaef, HMeercf, 6pori*rae 6opoaqa' nu6o Haq,,Ha,ouascs r or qacr' pocrpar nr6o rorrh*o a nepeA'efi Ha 60roeulx ciopor* ""prursl, rro*arorcs nap'bre [orrocbr. ocesas Htrs cMeuleHa K 6prouuoft xn_ cropoHe. opa*raoron npar"roft c Kpyrr'brM npoTo'otucoM. flonoxenne 6pnuHon a cnrnnoa cropor poarpa ycra*aBrrnBaeTcs unro. cn0oga c 6prorunofi no pacnorroxe_ cropoxbr Sparuorona. '^ -----Y"v'rq' rr(a Ha u.l(raJlbHblx Ha.{anbubrx CTAAllgx cpaBHIfTerIbHO pocrpbr KOporKI{e.
3 a v eqaHr { q. B c e r,re ftc rs o D u v a l i i d a eB.H .g ry vr T.H .H ans' geaa B*rroq'nr AB8 rsqser.,reftcrsa (1g67aro) co6creemo il;;;"; p"uio*, 1g14 pseudo_ dicoelitinae n u, rtr"in;*u", 196T. Duvariinae ;;;'orro."*rrbrrbre flchs porrre Bepere'oBl{Attblet ro_ iext c}6urnrnqplrqecK[e nnn cl6ro*,*rec*,,e co cnl{HHofi6opoeaoft a nepeq'en pocrpbr, qacr' o o6nt"n' [cKnrotrr{TeJrb'oa Terrqecxo'r rrosce, B ocHoBHoMB CpeqreeMnoMopcxoft o6nacrr, c nogAneft oprr, 6ara, r{ Ao a*a BKrroqErerrb'o.p"uudJi"ouiirin"" BosMoxgo c ;*;;;.1"*ro yArrHeEHbrMr 60agrueftgacrb*, BepereHOBr{AnbrMB pocrpaM[. y 60na*o"ar",
;:ffi::,ff:ff-":ffiffi",'
6prourH'" oqp na;1.'#:'"JTT":J ".oa.oxr 6ar
fi HHffi"H ff:lilJ##xffit#L fl?:Hiliffi;d::l"T;:ff:'
B'A' flcroMecoB orp'uaer cBssb p"urJoai"o.ritln"u-" B*rrorrr{rbrep?brx c'aqara Duvaliinae H nbrrarcs a ceveficrso Hastitia"" e ceMeftcraoDicoelitidae(rvttot".o;;"i;;). i'ny;;.::r, 1966), nosxe oor"*o pseudodicoelitinae valiinae o6naqaror pr{Aov rr Duo6ruux rpr{s'aKoB, orrrr*rapuru( 6enev'uroB. Ha pocrpax ro( * or Bcex Apyn'x npyrrn r.oil""lr""'l"l'^ill-:t** "l
6oposAH. a. qpy,"x6eneun"r::':ffi; ;.J,"#:i,::""::T :f*:l*jJf::, poc'TpoBr Ho BblpaxeH'bre
nepeaHefi qacrl{
3'aqnTenbno
Jlll;"tl'"lt#JrT'*of.""1jtl"e
cna6ee
6pro'*rux. Kporrre nDuvaliinul-B orn,*rreor BcexAplr'x 6s,n' se-
::n":en"#;#'ffi ffrffinn:i:"::rffi:Jil::rr"Iffi fr::,"'ffi ,*ooo'"
ffi :;:f,-,:ffiStrl":#;ffi:T';*: o"'"'"*uo:;:"'ffi:?":' ocHoBaHH.*""11il,:*,:::ffi:f 1'ffi:,#;.T:fiiffi*#ilh;,ffi? cK
oe p a cnp oc rpa HeHHe. Bepxn (r"loj'jiil;r"i,;;Hj:: unx'en rcp' cpeA'efiop", io"rl -""-"#,io Men(ao anra'"***,orerrbHo) #?X"::::? ""rT#. noACEMEHCTBo PSEUDODICOELITINAE SACHS ET NALNJAEVA , 1967
On r c a H n e. por;rpu
npe[MyuecrBeHuo
c nnv6oxoficnnr*rofir6opoeoor r nepeq'eft y
cnaftrofi.
BepereHoBnAHbre, c[rrbHo yArrEBeHHbre vo""le'Hble'
6onuuruHcrBa
"r.J:::",ff"1":-llll:-o_ BaAoe uascq or Bepur{Hbr,, tu6ocaa6o HlffHf:T:ffiif *:*"#:;iXtn 649
Bl
I
rr
B npuBepBbrx cropogar( pocrpa Ea6noqaprcs ABe 6orossle noflocbtr l{cqe3apul{e crBnaqanbrrblx Ha cropoue. .laCir{ nrxus Ocesas ,rrti'on "r"r"t*i:gt]ro"unror c$ronrrecKlfe. cna6o ropoTue, pocl?br oTtrocuTerlbHo aEf,,x PoAtt LenobelusGustomesov, 1966r u Pseudodicoelites cocTaa. Poqoaofi Sachs, 1967. Pocr?rr Pseudodicoelitinae PesKo ortltrratoTcs oT pocrpon DuCpaBHeEue. BI{AoB pasBf,TtreM y 6oruurncTBa valiinae caoefi sgaqn.net6go4 yAmtgeggocltlot npuBepuJl{I{B no,Ioc 6pnqnrol 6OpOOnur EapsAy co cntruHofi, aarylcallueM ooKoBblx qactn. Bepxu xnxnefinprt pacnpocTpaneHue' lr.reorpa$urecroe BospacT ApKrnsecroft' cccPt ceaepo-Bocrora cn6rpn, top. b.t"pnofi (roap) sau6onee Pseudodicoelitinae "p** 3eumr Oparura-.HT"0"' KanaAu, npeAltotloxrrelbgo oAHaI(O cu6nput ceaepnofi HIol(HeM 6afioce B AAneHe lut|potco pafipocryalteHbl npeaeJlSMt{ 3a 6AT€' BepXHeM B n xaliAenu T.H. Harsnceaofi eArH|IrrHue pErpu ne o5napyxegbl' xo$I ne rcK;1K'qegot sTo Eopeanuno"o not"" Pseudodicoelitinae e r Hibolites' no'roxeBHe cuSona so uexoTopbte cpeAHeKrpcKne Qopr.ru, oTttocttMbl HMeTb Ha pocTa geficrs[TenbHocTu MonyT KoTopHx He ycTaEoBneuor sparuoroue Pseudodicoelitinae' par pasBar),r, cnl|Hnylor a He 6porunyrc 6opo3Ay tr rpuHaArlexarb nd
z
t
afl Ta6fi qa BuAoBnoAceMeficTBa P seudodicoeI it i nae OnpegenrTeJtbg 1.
yAnHHeHHbIgr Ba Haqanb'POCrprr npe[MyuIeCTBeEHO BepeTegoB[t4gblet CH'tbHo co cnurnrofi cy6ronrrecKfiet KoporKrret cnerxa Hbrx craaltcr( ornocttrelrbno cnafil| conpoBo)KAanouefics KOltrIa 6opoeaofi, Haqt|BaKrUefics oT nepeAHeFO unu na6rnoaaeTcs pocTpa Bepullfltbt oT Haq[BaeTCs rofi. EprouHas 6oposaa BtrAoB orcyrcrByer' ronbKo B ero nepennefi qacrn, y He*oropbtx
H
c n
LITINAE NOACEMEfiCTBOPSEUDODICOE 2 (13)
3( e ) 4 (8) 5(6) 6(7) ?(8)
8(e)
fl
or Bepull{Hbl' Pocrpul c 6pourxofi 6opoeaofi, na.rrnapuelcg ' ' ' ' ' ' ' ' ' ' '3 aaneH Pog Lcnobelus Gustomesov, 1966t roap BCK' nocneanbBeonosITH unu Bc|o Ha Bpprunag 6opogqa'npoTsruBaeTcs ' ' ' '4 ' ' ' ' ' ' ' ' ' ' ' ' ' qaCTb ' ' pOCfpa ' ' ' t ttgpHylO " ' o ' 5 ' ' ' ' ' ' ' ' PocrpuBgpereHoB!{AEofiSopr"rur ' ' ' ' ' ' ' ' ' c 6oKoe' cxar Pocrp cnerra BepelenosnAHbtfir 66t Ta6r' 6' Oln' 2'31 L.lcnensis Gustomesov ($cror'aecoo, 1966t crp' TOSp - gnxntd €ratreB HanpaBneH'l{' Cneff,a BepeTegga11A'btfir Cxar e cn[HHo-6ppuIHoM POCtp -L.ailigaensis - roap-aaneH (Sachs) (Caxc, 19612r crp'77) no[epeqHblM POCrp oTqeTn[BO BepeTeXOsfqfbrfir-C 6rUrarUM K ogp]tnlroMy ceqeHBeM. Our'9-14) L.sibiricus (Sachs) (Carc, 19612, crp'85, aP-aa'!eH Pocrp cY6ulnrxaPrrecxnfi . crp'68' L.recond.itzs Gustomesov (fycmr"recoa, 1966'
- eepxnnfi ro-
82
o'
a
a tl
u
(
H
E
C
F
z
1
ra6n'6'
0ur' 1l -
nurnrtil roaP (?) npnBeputlnnofi qacTu pocTpa' ( I 13 [ Bproupraa6opoena He BbtxoABT3a npeAelbl nonepeqHbrMceseHueM' oKp)'rnoil{y K fbif f i Pocrp 6ynaaoauanulfic 6rurgxrM - aepxxufi roap - suxnnfi aaneH crp.86) 19612, L.vogt(SachsXCarc, Pocrp BeperenoenAnuft,cxarufi c 6oKoa' 11(f2) - sepx'rfi roap-aaneH L.minoevae(sachsxcarc, 19612, crp. 86) qacrt{ np[Bepruunxoft B Pocrprr ttuueunbte 6opoea 13 Poa PseudodicoelitesSachs, 1967 qacrn pocrpa npncyrcrByer nerny6o14( 17 ) Ha 6prorunoficTopo'e B nepe^Eefi K A g 6 o p q 3 A A t | n u y I I | l O u l g H U €. . . o ' ' '
n
U
T
!(
l
E
u I
) I
15( 16)
Pocrp orqernaBo BepereHoanqHrlft. Ps. bi'dgie'i(Sachs) (Caxc, 1g616, crp.g4, S r n . 1 _ g ) _ aepxnrfi TOap _ xuxsurt aaJreH tG ( fZ ) Pocrp cnerKa BepereroBraHbrfi. P s. platyventriosus sachs (caxc u Harn xseaa, Lg67 2, crp. 2 l ) _ aepxunfi Toap _ snxHuft 6aftoc L7 Ha 6proruHori cropoHe pocrpa 6opoaasl orcyrcTByror. 18 ( 19 ) Pocrp 6ynasoeuAHurft. Ps. clauatoides Saclrssp. nov. _ suxHnfi aanen 19(20) Pocrp cnenra BepereHoBtrAHrrft. ps. hibolitoides sachs (caxc u Hansxeena, Lg672, crp.21, ra6n.2, Sun. 46 ) - aaneH - Hr{xHul-r6aftoc 20 P o c r p u c y 6 u r n H H A p l r r l e c K .u. e . . . . . .. o..,. 2! L2(ZZ) Pocrp cpeqHnrl, cxar c 6oroe grrslonesovi Ps. Sachs sp. nov. - BepxHuft roap _ a6rr€H 22 Pocrp uenruft, cxar s gn[HBo,-6oxoBoM HanpaBrreur{r{. Ps.primoris Sachs sp. nov. - Toap - suxnnfi aarrex P o q L c n o b e l u . sG u s t o m e s o \ . ,1 9 6 6 Dicoelites (pars):Carc, 1g6 L, crp.77, Lertobclus:lycrouecoB, 1966, crp. g4. Tnn poAa. Lenoberus renensis flcrorrrecoB, 1966. Toap p.fleHbr. on ucaH ne. Pocrptl cHiIbHo yAnI{HeHHbre,BepereHoBrrAxbre EnE 6ynaaoalqHble' cxarble c 6oxoa, pexe s cnfilrxo-6ppuHoM HanpaBneHl{r{, uMenr nny6oxpo cnnHHylo 6opo3Ay B nepeAHeft qacru pocrpa n 6pnunryro 6oposqy, Haq11gaKrulyrccs or Bepulrxul' 3ra 6oposaa Moxer nporsnnBarbcs no BceMy pocrpyr Moxer r'pr flBrlsrbcs TonbKo B ero llpuBepunxEofi qacrn. Ha 6oxosbrx cropoxax napgbre nq'nocblt cJIerKa pacxoAfluHecs K 3aAHefi qacru pocrpa E saryxaputte B eno- IrptfBeprunHiioft qacrr{. opanruroxox nparrofi, c cnsonoM c dp,o*ron Beprunna anbBeoJrbl I'I oceBas' nuKus' npn6nnxeHbl K 6prourHoft cropoxe "-poor. pocrpa. V aepruunur aJlbBeoJrbrHa cntrHHoft cropone Ha6ruoAaercc cnafira (ne y Bcex srAoa). Ha Ha_ qaflbHblx crpnrsx pocrp cpaBHI{TerIbHo Koporrrft cJIerKa cl6ronnrecrofi rae c16UnntlHqplrqecxofi Qopuu. Bnaoeoft cocraa. H3 CeaepHofi Cn6npn onlrcaxbr G s[Aos, rro Koropbrx s (L'lenensis, L. ailigaensis, L. sibiricus, L. uist u L.minaevaa) paccuarplrBaprc, Hr{xe. Bun L.reconditus, on[canHblfi B.A.fycror,aecoBbrM (1966) ;; ;;;;;;;r' B Hauux KonneKuntrx orcyrcrByer. B.A. flcroMecoB yKagbrBaer raKxe Ha ro, gro oHToneHe3 Y L'reconditus urler B o6parHov HanpaBnennu (or BeperegoBrutgono pocrpa K cY6uunuHAPFtecxovy). B Hauev Marepr{ale, HactrurbrBapueM oKono 4OO pocrpoa Lenobelus, Hrr{eno noAo6nono He Ha6noAaercg. pocrprl .L.gravis, TaKxe onucaHHble B.A. lycrouecoBblM (1966), nuueubr ocnoBgono np113gaxaBcex Duvaliaceae - 6opoea B flepeAHefi qacru pocrpa r{ noroMy HaME Bbrue or'ece'br K poAy Holcobelus. Hs Apxrrrecrofi KaxaAsr )rrasbrBaercs Eneuxrnr (Jeie;;i;r* 1 9 6 6 1 I . e n o b e l u ss p . ' cpaeHeHHe. or Pseudodicoelites pocrpbr Lenobelus pesKo ornl4ra1grcs paoBI{THeM 6ptouuoft 6oposAsl B rlpl{Bepuruxnoft qacrtl pocrpa. V pseudodicoelites HMelorcs 6opoequ cnHHHas n se y Bcex BuAoa 6prcurxaa ToIrbKo B nepeAgefi qao._ rH pocrpa' Cnaftxa y Bepul{Hbl anbBeoJlbr Bbrpaxena y pocrpo s Lenobelus cna6o, y pocrpoa Pseudodicoelites 3HaqHTeJIbHo n)rlue, B qacrnocrr{ u a 6poruuofl qae TH pocTpa. Bospacr H neorpa0rrqecKoe pacnpocrpaHeHue. B e p x ' s s q a c T bH [ r x HeFo roapa (soHa Dactylioceras communeH Bbrue) aepxH"n .*"" CeaepHoft Cn6wpw, Ceeepo-BocroKa CCCP u Apxrrnrecrofi l(ssaflbro
83
(
L e n r t b c l u s I e r t c n . s i . s( i u s t o m e s o v
c
H
Ta6n. XIl, tfur.5-8 L e n o b e l u s l e n e n . s i s f: y c r o u e c o e , 1 9 6 6 , $ur. 1-2.
crp. 66,
r a 6 n ' 6 , S v r n 2' r 3 ,
ra6n'7 ,
c c
v
Mysefi M|PH, MocKea. Pexa IeHa, noc.KtIrr{aH, f onornn Ns \l-14L/2. H3 KepHa cKBaxHI{bl, roap. cxacnenKa BepereHoBlIAHL'll"Ir Ar,ranHo3. PoCrp or{eub cHnbIIo yAJl}IHeI{Hblt"{, or Bepuny3Kofi HaqHnarcueitcg 6opoagoir, r{ 6prourHofi Tbtrr c 6oroa, c rny{oKoii Hbt H nporrrrgBaourelicq Ao cepeAaubr pocrpa. CnuHHan 6opos.la nny6oKafl B aflb'{acru pocrpa. BeorHpHoii vacTu, 3axoAr.rr B nepeArrrororperb nocneaJlbBeorgpHoli q a c r b p o c r p a 3 a o c r p e H a ' c H n b H oB b l BHeLuuue npl{3HaKH. flpr-loepu-ruHHal T r r H y r a , c o c r a B r u r e r o r 1 / 3 . q o L / 2 r a ( r a 6 a . 3 O ) . B e p u r w H a3 a H l l M a e r u e H T p a n b Hoe nono)Ker{He,BepuuHHblirI'nor rlaxoAurcl B npeAenax 2O-28o. Y aepuunul HHonAa Ha6ruoAaercs rurpr.rxoBqrocrb. Cna6aq creneHb BepereHoBHAHocrHpocrpa ycraHaBn?IBaercrr no 3HarleHnrrMc6, cocraBnapurl{M y nor{TL Bcex 9K3eMnnslpoB gOO-12OO. Eoposaa Ha cnuHHol"I 160-L849o CE npr.r noKa3areflqx na, paBHbrx qacrr pocrpa HMeer nporaxeHHocrr, or 1/3 eo L/2 gnunar cropoHe B nepeAHeft pocrpa. Eopoeaa ua 6prourHortcropoHe rny6oKas v y3Kafl, y HeKoropbtx pocrpoB TsHeTCrror BepuuHbr Ao cepeAHHbr pocTpa, y HeKoTopblx npocnexl'IBaeTcq Lo aJIbBeonrrpHoft .{acru pocrpa BKnpr{u.TenbHo.EoKossle cropoHbl ynnotueHbl, Ha HHx MO)KHOBHAeTbABe napbr napannenbHbrx6oxosrtx noJIOc, 3aTyxapuux B'npHBepllHHHofi sacru pocrpa. flonepevuoe ceqeHlle oKpyl'neHHo-TpaneuueAarlbHoercxaToe c 6oxos. Creneus cxaius y 6onsuuncrBa pocrpoB Bo3pacraer B 3aAHeit n> B rex cnfraqx' Korqa EE 6onuue 71-99). noBr.rHp eocrpa (SS Z8-12O,66 pocrpa B ero anuaeonapxofi o6tgcueHo aeSopvauunMll Mox{er 6slrs 1OO, sTo
A B
E N
E 6 B
,r c c a
B
L
F A
o
qacT'l{.
rl
Anugeona npnMaar 3aHHMaer or npr,t3lraKH n oHToreHe3. BnyrpeHHHe q o c u n b H o c M eueHa x 6ptoulHoft cropoue p o c r p a , B e p u H H a 1/1O Anr.rHbr 1/5
3
Ta6nnua 30 Hs v r epeunsp o c rp o B I.e n o b e l u sl e n e n s i sGustomesov
N e8 7 - 1 6 1 , P. Kenur.anP Anraua fnpeanonanaeMatl o6uras [ycranoaneHHas Anuna nocneanbaeonnpHofi qACTI,I
fiuar.aerp cnnnnoSPtoutHoft Bepul{Hbl
anbBeoJlbl
fl.uauerp
6oroeofi
y
Y
BepIUHHbI
N e 87- 162, p. Kenrunp
9 3 , O( 2 3 2 5 ) 121,0(2200) 90,O (225f]^) 118,O(2145) 1 O 5 , O (1 9 O 9 ) ( r O + O) 8 2 , 9 ( 2 0 7 3 ) ( 1 1 8 4 ) (8e) (87) 4 , O ( ( 1 O O ) 1 0 0 I 5,5 5 , 7 ( 1 O 4)
4,8(12O)
AIIbBEONbI Paanyc
6pr,ourHofi
y
BePuHHbl
ANbBEOIIbI
Anuua npl{BepunuHoft qacrn flnauerp cnuuuo-6pouruoft a Mecre Han6onuuteno yrorlueulln finaverp 6orosoft B Mecre Har6onuuero yrofitueHnn Yron Bepul{HHblft B 6oroeoft
34,O(618)(32) 10,1(184)(rOO) 1 O , O (1 8 2 ) ( 9 9 ) 22
3 1 , O( 7 7 5 ) ( 3 7 ) 7,0(175)(1OO) 7 , 1 ( r 7 8 ) ( 1 O 1) 22
u
(6prorunofi paAayc cocraBrlfier oKoJro r/4 cB). AauaeonapHu',r yfon cg' cnafiKa y BepultHbr anbBeoJrbr He 3aMeprrn* Bbrpaxeua cna60, 3axoAr'r B n H)4or{acrb np'MepHoHa BenrqnHy cB. ocee an- nw*ws.. npra6nnx"r:Tfi"ffffi* cropoHe' nor{Tll np'Maq' Ha rtaqanbHbtx craA'rlx pocTpbr cpaBHkTenbHo KoporKr{e crrerKa cy6rosrqecKr.re (npr.r CB 4,S uv na oxoro 44O). IIpa CE 6 MM pocrp yAnnHgercfl (lla oxono 7Bo) n npr'ro6peraer cy6rrununqpnrecKyp Aanb*eftueM pocre-xr{BOTHono pocrp sopvy. frpu craHoB'rcq- i + zcna60 o, BepeTeHoBrAHbrM,rla Bospacraer nplr c6 612 MM H c6 grg MM Ao fla qo iooo (pnc. s2).
L.tenensi" ; ManonsMeHquBbr. EonruruHcrBo *ff;:#,:::3-. pocrpoBB ";"M "":,.:.1: ;;;;;;; ;r:":::[J:i" J;Hfi:'"' veH'Urefi creneHbro BepereHOBHAHOerr{ "u'.n (po..p
xn g7-762 u Bcrpevarcrcs TaKxe pocrpbr, y Koropblx BepeTeHoBHAHoerb Ap.). BbrpaxeHa 60nee cl{nb'o r{eM y ronorrna, Bnnorb Ao anbBeoaapuofi qacrn FppruHaa 60pocaa nporrrr}rBaercrr
HeHr-rxHera roapasaMerHo or,*"To"; ?T:1"$":rt; fff"ffiT::;"lo"""euuocrblo, cx(aTI{eM'
r^?^Xil;l"
Manofi creneHbro BepereHoB HaHocrH,
pag'nl{ev 6prcurrrori 6oposAbl TonbKo B rorrororr.tM'";ff;:t"bBeo,I'pHoft qacril' a cnlrHHoii 6opos^br B anbBeonapHofi ronbKo gaqHeft
C p aa H eH r{ e. pocrpsr L. lenensis ornrqarorcsr or Apynnx Br{AoB L.enobelusceoefi raub cna6o BblpaxeHHofi aepereHoBHAHorl {opvori, 32. IlpoAorrbHoe cerreHr.re pocrpa L e n o b e l u s l c t t e n s i s i":: Ne 87-163, Gu.stomesov. BepxHnfi roap, p. KefluMqp( x O17)
oqe'b cnnaHoft yAnI{HeHHocrbIo tt cDInbHbrMc}Karl,reM c 6oxos r{aqtu' cneayer oco6exxo B saA'e}-r orM€Tnrr' qro Ha rcHbLK cra'r{sx pocrpbr L. sibirictts HMeIor raKyro xe tt L.uagt fuplry, KaK u pocTpbr om{cbrBaeMonc BHAa" pasnuql.rg Me)Kry 3,TI,IMI,I BIAAMI{ BbIgBJIqIO*r NHUJb HA
BSPOCNbIX
g7- 164, l-,Jb p. Bunroft
6 7, o ( 8 7 O) 8 8 , 3( 7 4 2 4 ) 6 4 , O( 8 3 1 ) 8 8 , 3( t + 2 4 ) ( 8 9 2 ) s 2 , s ( o e e ) (6 4 0 ) ( 7 8 ) 6 , 2 (1 0 0)
7 , 7( 1 o o )
6,o(96)
6,o(78)
L , 7( 2 7 ) 3s,2(Soe)(40) 9 , 9 (1 6 0 ) ( 1 0 0)
25,2(szt)(48) 8 , 2 ( 1 0 6 ) (1 0 0 )
7 , o ( 1 1 3) ( 7 1 )
6,7(87)(82)
20
2a
CTAIII{,D(.
flpnrr.reqa Htle. B ra6aruax. xeuepennft pocrpoB pseudodicoe_ litinae nocne yrasaxus pasMepoB B MM B nepBbr.x cxo6xax, KaK r pa[ee, AaHbr npoqeHTbr rro o(nHoruerHlrro K cnrnno6ppunolvry AnaMerpy y Bepun{Hbr arrbBeoJrbr. B npa$e 'Amrua nocnearrbBeonnpnoft ,racrll, Bo BTopbD( cro6rax AaH npouenr r cnnnno-6pnuHor\dy lttaMsrpy y Mecra nan6onsuero pacumpenrrs poc-Tpa, B Tperbrx cxo6rax _ npc. ueHT K o6ueft Annxe pocrpa. B rpa0e 'Aanna npuaepulrHsoft sactu, KOrOpafl IrOHnMaeTCg, KaK paCCms_ Hne oT Bepurt{Hbr tro M€CT6 rr6n_ 6onsuero yroJuueur, poerpa, Bo BTopbD( cxo6rax AaH npoqenT x AnrHe flocneanbBeonnpnofi qagrrr. ',[uar*erpei B rpa0ax B M€ca€ Haf,r -6onsuerp yTomqeHns, sop"x "o cxo6xax yrasau npouerm K crrtxxo6pnruuoMJr A[aMerpy B Mecre nan6onsuera yrtrrrIrjtenuc.
85
B o c H o B H o Mp o c r p b l c o 6 p a x u B r n u H l { c npr{ypoqeHHocru. OauuanbHas rrrx $aUuflx oTKpbtforo Mops e IIeHo-OneHeKcKov paftone, 1 pocrp AocTaBneH c ;1peAropuft BepxonncKono xpe6ra, Toxe Ha o6nacrn orKpbrroro Mopcr eAltHtfrHble paggeroapcKge pocrpbr AocraBnenbr Hs MenKoBoAHulx {aunfi s Blrnoftcropt s Ana6apcxov pafioHax, 1 paHHeaaneHcxufi pocrp ilponcxoAuT H3 MerIKoBoAHulx $augfi ycnoBHsx orKpbtTono Mopsr Bocro.rHoro Tafiruslpa. Orenr{AHo xnBort{bte xnnH B pocrpoBr xopolxHMl'I nnoBuaMH crpenosEAHofi yAJrHHeHHoft {opue 6srnn, cyAq no n noqrg He 3anrrbrBanu e npu6pexnbre fracrKn Mops. Oco6o cneAyer orMertlTb oTcyTcTBHe HaxoAoK L.lenensis a Oxorcxo-OtuonoHcKoM pafione. p a c n p o c r p a H e H u e . H u x H n f ir o a p ( s o H reorpasuvecKoe Bospacr na Dactylioceras commune H Bbrule) - HuxHnft aaneH CeaepHofi Cra6upn. - HHxEe5.oaaJregape6 Kenuvnp Marepua L 25 pocTpoB u3 BepxHero Toapa C.B. MeneAruofi n lO.l4. MuKnpuHofi, T.H. Hanunreaofi, c6oprr T.H. u Eyop-3eKur peK u Eerngxanr t pocrp ue Jlenbl aaneua Toaps-HnxHero pocrpa r{3 2 Haeaofi, pocrp t|3 Hl{XHetO TOApa Anac6opur T.H. KupuHofi, I HHxHero Toapa p. Bunofi aaneHa p. t{eppocrp H3 Hnxnero 1 6apcxoro 3aJrr{Ba c6opul T.H. HansHReaofi, noxpe6erHoft (Bocroqnuft Taftvulp) - c6opsl C.B.MeneAnHofi. I I-etroltelus vili gaertsi.s(Sachs) Ta6n
Xlll,
Srn,
t-4
Ilibolites .sp.: TYuxoe, 1954' I l o l c o h e l t t . st t i l i g a r n . s i s : c a x c , 1 9 6 l a ,
crp. 77;
L962q,
cTp. 433.
Hoaocn6npcr. Pexa KeMysefi Hff co AH cccP, Nq a7-L66. fonorun (Hnsonbs OneHexa). nuMrrp, Bepxul{ft Toap Pocrp cunbHo ygnuHeuuurfi(IIa oKono 8OO-13OO), cnerKa BepeAuarHo3. (BE orono 115-135), TeHoBr{AHbrii,cxarurft n cnsHno-6prcuHoM HanpaBneHrl{ or BepuuHbl I{ BblnoHaquHaloutefrcg c lunpoKoft n nny6oxoft 6prcurnoft 6opocaoft, .ractu. nrry6oxan B anEF 6oposaa Cnuxnas anbneoncpHoft s6nusu JralKt{Barcrueficc qacTb nocneanbBeongpHyto B r{acTH, HeHaMHOTO nl{lub SAXOAHT aeonepuoft pocrpa. n p r { 3 H a K u . B e p u n H a 3 a o c r p e x a , 3 a H u M a e r u e H T p a n b l r o en o n o BxeuHHe (ra6rt.31). V eepuunut ynon HaxoAurcs B npeaenax 25-32" BepugHHbrfi xeHue, pgcHa6.rnoqarcTcs roHKHe urpllxt{. ilpusepruuHgaa qacrb Ao Mecra Ha[6onuu€Fo BepereHG JrHllIb cJIeFKa Pocrp lla. 3-I/ 2 ugpegurr pocrpa cocraBnqer oKono L/ euAHr,rii,3Haqegnfl c6 cocraBnslor 1o5-135o/o CB lrpu noKa3arensx na 75O-13OO. Boposaa Ha cnuHHol"rcropoHe B nepeAHeii qacrll pocrpa nporqruBaeTcfl npuMepHo Ha O,4 o6ureii ero aru{Hbt, nocreneHHo Bbrnonaxl{Baqcb. EprcruHan 6oposna Ha{IlHaercn y Def)utlrHl,rpocTpa, HHorAa Ha paccTonHuu 3-5 vv or Hee' no Mepe npuqacru 6maxexng K BeplutrHe an6aeoJlt:t BbtlloJlalKunaercn, ItHoI'aa B anbBeongpHoft pocrpa cropoHbl cHoBa ycynr{Baercg. Enaronaprr 6oposnav ctrt{Hnas ll 6ptounal jlJrorrr€Ht,r, 6oxoRr,re BbrnyKnbre. Ha Hux Ha6.ruoAaprcn napannenbHble Aplr apyny napHbre nonocbr, c nororofr nox6llnxoft vexay HltMH, satyxarcurue B cepeAl{He npr{BeputuHnoliqacrn pocrpa. fionepe.ruoe ceqeHlte pocrpa l{Meer xapaKTep oBana, cxaroro B cnuuHo-6pronrHoM HanpaBneHut{, npuqeM cre[eHIJ c)Karuc MeHgers cfl B :]aBucyMocrlr or creneHH pa3BHTI{rrcnusnoft u 6proulroft 6oposa no paSHoMy: B y oaHnx gK3eMnnsFx>B oHa 6onsure y BepuuHbl aflbBeonbl' y apyrux ( nS oT 1 1 3 Ao 1 35' 66 ot MecTe Hau6o.luruero pacuHpeHnc pocrpa LO4 ao 135). BHyrpeHHIre n p l r 3 H a K r . r u o H T o r e H e 3 . A n b a e o n an p l M a l , 3 a n l t M a e r NteHee 1/5 annnbr pocrpa, BepuI{Ha ee 3aMerHo cMeueHa x 6prcruHoft ctopone.
lB"n
86
Ha3IlaH tro p. Bunure,
rae
oH BII€pBtrI€ 6t,ln Haiiaen.
Ta6nuua S1 I,fsvepeuus pocrpoB L e n o b e l u s t,iligaensis (Sachs) (c p. Kenunap) Ilapavetpr,l
N" 87-166
,[l.nuna fnpeanonanaeMarr o6uraq lycru"orfleHHas Anuua nocneaJrbBeonsp_ soft qacrn
]t/,?87-167
L L 7 , o( r o + e ) 1 O 8 , O( r s a g ) (r s s o ) 11O,o 1 0 1 , s( L 4 4 3 1 9 6 , 0 ( r g s o()r r 4 3 ) ( 8 2 ) e o , o ( 1 2 8 6 )( reoo)(es)
Ar.raverp cnrHHo-6ptour_ 7,7(1OO) uofi y BepulrHbr anbBeoJrbr llHaverp 6oxoeofi y Bep_ 8'e(:ril urI{HbranbBeonbr Faauyc 6pou.ruori y Bep_
7,O(100) 8 , 8 (L 2 6 )
IUHHbI AnbBeOJILI
AnHHa npunepu_rHnnoli qacTl{
Auaverp cnnHse6prcuHofi s MecT€ nau6ontueno yromueHue
Auaverp 6oxoaofts-y..-
te Hau6onbueno yronueHujr Yron anbBeonspHrfi a cnrxno6pnurHoft nnoc_ KOCTI{,npaA. Yroa BeputlnHbrfi s 60_ xonofi nnocxocrl{, npaA,
4 6 , 8( C 59 ) ( 4 s l 8 , 4 ( 1 1 8 )( r o o ) 1 1,o ( r s< ) ( 1s 1)
2G
ss,sti, nte7) 7 , s ( 7 C 7 )( r o o ) 9,5(1go)(tzt)'
2A
OroH.{aHue ra6ltrqur rlapaverpr,r
A,.u*ra o6uas
f npeanorraraeMarr
ly"ru"ooreHHarr A.rnna noi.'rearuBeo.rerp_ uoii qactlr Alravetp cntrHHo-6pul_ nofi y BepuuHbl aibBeo/rbr ri[ualrerp IIJIIHbI
fioxosoji
y Bep_
N,g7-169 60,0 (1579) 54,5(L4341
4 e , 2 ( 1 2 e 5 ) ( g e(sa)z 1
8,2(100)
3,8(1OO)
11,O(tlrl1
4,3(113)
2,7(33)
O.'r bBerO.'l br
A.rHua npuBepurrHHo, qacTu Ararlerp cfiuHrro-6p]oru_ xoil g Nrecre uatt6orlr,_ rilcro
8 3 , 5 ( l _ o1 8 ) 64,s (787) (c;os)
A.:IbBEOI'IbI
Paqlryc 6prcuruoii y Bep_ IXIIH bl
N" g7-168
3 6 , 4 ( / l + q) ( 5 6 ) 9,7(118) ( 100)
1 G , 2( 4 2 G ) ( . : . : 1 5 , O (1 3 2 ) ( r o o )
1 1 , 8( t , 1 4 )( L 2 2 )
5 , 2 ( 1 3 6) ( r o < I
yTo.llueHlts
fluarrerp 6oxoeoii B \rec_ re ualt6olt ueFo yro.-tuleHlIn Ynolr a..rL,Beo..tqpnsrii n cnuHuo-6prourHoii
25
r..loc_
KocTr4, I.Pal. Yrofl B(\pllrrHHr.,rii n S9tronoli nflOCKocTn, rpa.g.
32
25
87
yron oKoEprcugroft paAryc y Bepur{Hbr anbBeoJrbl oKono 1/3 CE. AnreeonspHutft MeHee no 2So. Co cnn111oft c,roponbr y Bepulrubr anbBeorlbt ua6ruonaercn cfiafiKa, qacrb Ocesas pocrpa. qeM Ha Benuqr.rHy CB erlcrynarorrvag B nocneanbBeofispx]tlo JrlrHlts ct{JlbHo upu6nuxega K 6prcunrOfi cTopoHer noqTil ltpflMan. Ha naqanbnblx x cy6UuruauAcTa4uirx poqTpbt I{MeIoT cna6o cy6xoullvecKylor npn6ruxapuryrccs puvecxofi {opvy. I-lpu exaveHl{sx CE oxono 3 vv fla paaHo 5OO, npu CE 5r5 r'av oKono 7OO, Sopr"ra cTaHoBHTca cy6Uung11Apu'Iecxoft. Kar n y B3pocJrbt;1 h" pocrp cxar B crrl{HHo-6prcrunou HanpaBneHl{l{, I{Mee'r nny6oKl{e cnI{HII}rn ocofufi B nanbHeftuev pOCTp npuo6peraer cna6o BepeTeHoBI4AHyIO 6opoeaul. 6prourHyro u ao 7 87 npu CE 8r2 vv ila - y pocrpa Ne 87-168 Bo3pacTaeT HecKoIIbI(o $opr"ry, 1OOO-135O' pocrpoB (prc. 33), y Ap)n'I{x ao P o c r p r r L . u i l i g a e n s r s B u e n o M A o B o n b H oo A H o r m x r t . H a Hsrr{eHqrBocru. (e npeaenax psAy c ne6onrrxuMl{ oTKnoHeHEnMI{B 3naqenilqx fla, II8, EE u 66 na6ruoAap'rcs 104-135) 1 13-135, Gso-l2oo, coorBercrBeEHo zso-l4oo, '
eepere1rp6t1eM TonI'Ko B aanege, SOpUu c 6Onee peSKO Bblpax(eHHof,t pocTpbl' npu6H Hao6opoT B Toape BcTpeqarcTcf; HoBI{AHocTblo pocTpa. B Toape oTMeqaloTcfl x cy6unnuHApuqecxuv. no fupve nuxalouHectr B cpeAHeI'"I qacTg 6opoSnst 6pnuruofi cnna)KeHHocTbto co TaKx(e SOpVsl Har{HHaeTcq He pocTpbr, y KoTopbtx 6prcuHan 6oposaa pocTpa. Ectr MM or Hee. BepuII{HbI' a B 3-5 O n I{CbIBaeMoro BHAa pe3Ko oTnl{qaloTcq PoCrpbl CpaBHeHue. no Hanrr{ilrc c}IJIbHono cnHHLertobelus poaa B[raoB Bcex or [pyr.fix 6prorunoft vt Anlruuoft I{ paSBHTHIo tultpoxofi c)KaTHs uO-6prouuOnO
y caMofi
6opoeasl. 3aveqaHHs. par L.uiligaensis rrMerr pocrpoB
olnec pocrflepBouaqanbHo v-v- Ty'rxoe (t'gsa) ue a B.H. Caxc (f gO1a,6), x poAY Ilibolites, c coxpaHueuJeftcg anbBeonnpHofi .racrblo I{ cnltuuoit 6o-
P n c . 3 3 . f l p o a o J r b H o ec e q e H u e p o c r p a L c n < t b e l u su t l i g a e n ' s i . s( S a c l r ' . ) . oepxuttft Toap' p. Kenurtsp( x Or75) Ne 87-168,
c6opoe 6onee npeAcraBl{po3Aoft, BKntoqun qanusrft Buq B Wu Holcobelus.l-locne oTHOCHTCg K pOry TeIIITHOT.OMATepHana cTano oqeBl{aHo, t{To BI{a t,iligaell.sr.s Lenobc lus. Han6onee tlacro pocrpbt L.uiligaensi.s rb. rr nprypoqeHHoc OauuanbHa Mopn e JleHo-OneHeKcKoM pat"IoHe' B nnrHucrr,rx @uuax orKpbrroro HepeaKo ouu o6sapy)KuBarorcrr I{ B npel{N{yuecrBeHHo I'IIuHHCTLTx {nuunx OxoTpaftoua Bganu or ropcKono MarepI,IKa. EAnultqHrt HaxoaKu L.uicxo-OvonoHcKono paiioHa, rpprrupoligaensis e 6onee MeJrKoBoaHbrx $aunnx Ana6apo-Xaraurcxoro ppcKorro ct{HCxaeaHHoe no3Bonqsr MaTeplrKa. no6epexsn e6nugld BaBuruxcrr r{To L. ailigaensis x(It3Ht{ rt npeano.i o6pas Ben cBofuaHonnaealouuft Tarb, quran 3annbman B nplt6pexrurrJb II3peaKa Moprr, orr(pbrroro o6ctaHoax[ BcrpeqarcTcrr
urre
pafiowu.
pacnpocrpaHeHtte. H t r x u u i ir o a p ( s e r . rn e o n p a $ u v e c K o e C u 6 u ptl. B e p x H t n ' r a a n e x C e e e p H o f i Ha Zugodactylite.s braunianus) 43 pocrpa r.r3 BepxHeno roapa-BepxHero aaneHa pp. Kerulrnp Marepuan. Bospacr
- c6opur T.14. HanuHqeeoii, T.[{. Kupnuofi, P.A. Elrax(IleBa H pocrpa u3 Bepxner-o roapa - aoJIBHa p,Ilenltcxe u 2 pocrpa /vtenear{Hoii,3 9 pocrr{3 Toapa Ha pa3BeAoqHbrx nnouaarrx Ha p.Ilene - c6opr,l T.1,1. ltrpuuoir, Ana6apcxoii ry6l,t lt p o B u 3 B e p x o B H r . t ) K H e n or o a p a - B e p x H e r o r o a p a n o 6 e p e x s n pocrpoB It \rHol'o o6nortxon rr-oBd Yprour--TyMyc - c6opr,r T.Il. HanuuRenoii,5 - t'6ono6eppxlfl H3 BepxHeno roapc - oaJr€ga 6acceiiHa Konl,nrrr,r lr Oxorcxoro
u Eyop-3eKHT C.B.
pur A.C.
88
Aarrrca.
L e n ob e l u s . s i b i r i c r r s( S a c h s) I ) Ta6n. XlV,4rnr. I-4 D i < - o cl i t e . s s j | - l i r i c r r s: C a K c , 1 9 6 1 a , c r . p . g S , $rn. 9-14; crp. 433. I-tnobclrrs si&iric'rrs: fycrouecoa, 1g66, crp. 65.
Carc, 19616,
Ne 82-6. Myseft Hrf CO AH CCCP, Honocu6rpcn pera Eyop_ ,.^.,1: i"'':1l^i (Husonsqp.JIeHu), uuxuufi aaneu. 9eKuT A u a n H o 3. Pocrp cl{nbHo yanl{HeHHrrft, eepereHoBuAurrft c nonepeqHbrM ceqeHI'IeM, npr6nuxarcuHMcg K oKpyrnoMy, c 6prouurofi 6opoeaofi, Haqr{Haroueficrr oT Bepul'rHbr I{ crnaxl4Baloueficn npu npu6nrxenl{r.r K aJrbBeofispgoft vacrrr. CruHHas 60posAa npocne)Kl.{Baercs B nepequefi ilonoBr{He pocrpa. BHeuHI{e npI{3HaKa. llpuoepuuHHarr qacrb KoporKarr, cocraBnser oKoJro I/4-V3 l-la, Bepur{Ha 3aocrpena, 3aHr{Mae,r 6nusKoe K ueHTpanbHoMy [ono]Ke_ HI{Io' I'IHoI'aa cnerKa cMeueHa K cnuHHoft cropore. BepurunHurft -ynon Haxoalrrcrl B npe^enax (ta6n. 26-290 82). BepereuoBuaHarr pocrpa noATBep)KAa_ sopva ercg 3HarleHltsvr c6, cocraBnslotul{Ml{ 160-2207o CE npn floxa3areJrsx na oKoJro 75O-85O. Cnrssan 6oposaa nny6oxas u y3Karr, Haq[Haercrr or repeAgeno Kpas I'l l'IAer noqrl'I no cepeAl'rHbl pocrpa. Bprcruuan 6oposaa HaqrHaercsr or Bepurgbr, BHaqane raKx(e rny6oxan u ysKac' pacuupqercs H Bbmona)Kl{Baercrr nplrMepgo B cepeal{He nocJleanbBeongpHoft 'racru pocrpa I{ fipocnexl{Baercrr ao aJrrrBeonnpnoft qacrl.t eno' cnr'uag cropoHa pocrpa e .saAHefi eno ,,onoBEHe craHoBr{Tcrr cnenKa esmyxnoft' Eoxosrle cropoHbr ynnoueHbr, Ha HI{x Moxgo Br{Aerb napaJurenbHbre napHble noJlocbl' cnel'Ka pacul{pglouuecsl B fipl{Bepurrnoft qacrt,r pocrpa u garyxabLut{e npn upu6ru{)KeHl4l{ K BepuI{He. llonepevuoe cerreHr{e pocrpa y BepuuHbr aJrt, BeoJIbr 6nrsxoe K oKpyrneuxo-cy6KBaapaTnoMy (y vnorux pocrpoB ono cxaTo B cnrHno-6prcuHoM HanpaBneulru. BcneAcrBr{e nosAHeftruux Ae$opvau}rfi). B gaagefi qacrl'l pocrpa nonepequoto $opva ceqeul{e raK)Ke oKpynneutrecy6xaaaparHas, pery' HarIpI{Mep, y roJroTErra, pocTp B 3aA}reft qacrn c)KaT c 6oxoe.
Bnyr peHHlre npHsH aKu t4oHTo,- eHee. Ansseor,, ,rp"rlr, saHrlMaer oKono L/ 10 anuHbr pocrpa, BepuHHa cMerueHa r 6prcurHofi (6prou*rofi panrrvc oKono o,ss cE). AnbseonspHsrfi yron pa:::P:l% BeH z 5 . cnairxa rra6nroaaercrr B O6nacru Bepur{Hbr anbBeonbr co c,,nnH_ uofi cropoHbl' saxoAr{T B nocneanbBeonepHylo qac?b npr{MepHo Ha qnB_ Hy arrbBeon'. ocesas nr{Hlrs nor{Tt nprrMarr, npu6nuxeHa K 6prorunofi cropo'e' Ha Ha'{anbHbrx cra'Hrrx pocrpbr HMercT Cy6unnuuApr{trecKyro 0oprury, npa CE 3,8 vv fla 860, upr CE S MM pocrp craHoBr{Tcs cna6o BepereHoBlrAHbrMc IIa oxono l goo , ,"ornrquM or MoJroAbrx pocrpoB L.lenensis (pnc. g4). I,lev e Hr{ r.rBo crb. pocrpsl L.sibiricus rloBonbno oAHorr{nHbrno ceoefi 0opr're. Hapnry c npeo6naAarcur{Mu B KonJreKur{H BepereHoBua_ HbrMr{ pocrpaMr{ (pocrp NfegZ_176 u np.) Bcrpeqaprcs pocrp', 6onee Puc. 34. N,r 87-184,
flpogoJrbHoe ceqeHr{e pocrpa Lettobeltts sibiricus (Sachs) aepxuufi aaJren, p. Keanvrp ( x
O,ZS)
3aocTpeHHble'
c MeHblullM BepuI{HHbIM yl'noMr c MeHee scHo BblpaxegHofi eepereuoK raKI{M pocrpaM orHocrlrcsr ronorr{n, onucaHnufi g 1g61 B.H. Caxcov HuqneHrryanluoft oco6ennocrblo nonornna rrBnrrsrcq Hanr{que 6prcruno-6oxoaofi [puBepullHHofi 6opoeqbr c neBofi ctoponbr y qplryr{x pocrpoB pocTpa. B nsyqennoft Kon_ neKIII{u TaKas 6oposqa se ua6nrcaanacb. HsverqHebr crerreHb yAnr{HeHHocrH npl{BepBI'IAHocrbto'
-
Sibiricus
cu6rpcxufi.
89
r.
Ta6nrrua 32 I lcrlepeuurr pocrpoB
L c n o h c l u s s i l t i r ic u s( S a < ; h s )
N'82-6, p. Byop-3exur
Ilapalterpt,t
,[,nlr tta (I n pennonoFae I\raq o6urag IycrauonneHHan Anlrua nocJleailuneonnpuoii qacrrl cnuuuo-6ppurIltralrerp uoir y BepuI,IHr,r anbBeo-
N.87-176, p.Ke,nur.rnp
65,O(95G)
82,8tlo esI
65,0(956)(err)
78,3( rSO6)(8OO)
6 , 8( 1 0 0 )
5,2 ( 100)
6 , 5 (9 6 )
6,O(11-5)
nbl Auarnrerp 6oxosoii lXl{HIrl anbBeOJIbI Paauyc
y Bep-
6prorunoii y Bep-
rullHbr onbBeonbl
2 r , 7 ( 3 1 9 )( z s 1
AnHHa npueepulrHuoii qacrll cnunue6pnu,[.uaverp Hofi e Mecre uatt6onlueno ytofiueHurr
1O,81 ( 5 9 )( 1 0 0 )
A l r a v e Tp 6 o x o e o f i B N r € c - 8 , 8 ( 1 2 9 ) re uau6onbrxeno yroaue-
(81)
2 2 , 7( 4 3 7) ( 2 9 ) 9,8(188)( 1oo) 9,6(18s)(98)
HVlfl,
Vnon
anbBeoJlrrpur,lfi
e
cnlIHHo- 6prourHoft nJlocKOCTH, rpaA. Ynon xoeofi
BeplxllHHbrft
e 60-
nJrocKocru,
rpaA.
26
38
urunuofi r{aeru pocrpa (ot 33 ao 39o/o f]a), nporsxeHHocrbcnuHuofi n 6prorunofi 6oposq. C p a r H e H Ide. Poctpur L.sibiricus no caoeft BeperenosuAHoft Sopve 3aMerHo orJrr{qarorce or Apynilx BI{AoB Lenobelus. Enuxe Bcero oHIr crorrr K pocrpaM L.ninacuae, oaHaKo nocneaHl{e pegKo Bbraenrltorcrr rro HaJII.tr{uIo nulub xopoTxoft 6ptotuHoft 6opoeasl B npnBepruunnofi qacrn pocrpa. O cxoqcrBe pocrpon [-" sihiricus, L"lenensis u L.uagt Ha ronbrx craaurrx fooopltJrocb Bbtute. P o c r p t r L . s i b i r i c u s o q e H b r r a c r o a e $ o p v n p o B a H b r .B u e y u e H 3ave.raHLIfl. HOfi XOnneXUUU7olo pOCTpOBIr3OnHyTO r{nu CnaBneHO, Y Oyeub MHon[x sxgevnnspoB B cBssu c o6utuv yronueHueM pocrpa B nepeaneft eno qacrll, ora qacrb o6nouaua, qro HHonIIa clKara, npHBoEHT K I{cKax(eHl{gtvr B 3Haqenusx CE. Q a u n a n b H a q n p H y p o q e H H o c r r r . I l o q r [ B c e p o c r p b r L . s i b i r i c t L sc o 6 p a x r r B rnl{Hl{ctsx @uunJ( orKpblTono Mope e IIeHo-OneHeKcKov pafioHe, sgecb gror BI'IA Bcrpeqaercn qaue Bcex apynl{x Br{aos Pseudodicoelitinae. Oqnr eKseMnnsp L.sibiricus Haftaen a Bumoftcxov paftoHe B MenKoBo[nulx npu6pexnbrx @unnx. HeussectHbl Haxo[xn L"si&iricrrs B MelrKoBoaHbrx@uuex AHa6apo-Xararu.cxono pafioHa H e Oxo'rcxo-KonrtMcKoM paftoue. Bce cxasaHnoe noaBoJurer cql{Tarb, 'r'ro L,sibrricrrs aen cso6oAHonnaBaroutuiio6pas x
N 87-177, p.Kenuvnp
N e8 7 - 1 7 8 , p.Kenurtlnp
1 1 2 , O( 1 5 5 6 ) LO7,3(1490)
80,5(rzeg)
1 O O , O ( 1 3 9 0 () e o o 1 ( B e ) 7 s , 4 ( 1 6 7 6 ) ( 7 5 4 )
N" g7-194, p.Kenrvnp
IOO,O(1493) 93,4 ( 13 94) 87,6(rgo7l(796)(88)
7,2(100)
4,5 ( 1oo )
6,7( 1oo)
7,3(101)
5 , 7 ( 1 2 7)
7,O(1O4) 2,4(36)
3 8 , 5 ( S S S )( S g )
2s,5(s67)(S+1
24,2(3Or)(za1
1 2 , s( L 7 4 )( r o o l
1 o , o( z Z z )( r O o y
1 1 , O( t a q ) ( r o o )
L 2 , 4 ( L 72 ) ( g g )
9 , s( 2 1 1 )( 9 5 )
1 0 , 6 (1 5 8 ) ( g o )
25 39
39
45
Marepuan. O x o n o 1 S O pocrpoB t{3 Bepxxeno Toapa_aalreua peK. Keaurraap u Eyop-3eKHT - c6oprr T.H. HansHRenofi, T.H. KnprHofi, p.A. E'Axuesa, C.B. Me_ aeatnofi tI lO.H. MrHaeaofi, 1 pocrp rf3 Ht{)KHenoaarreHa p.TroHn (6acceftx Bu_ mos) - c6opur T.H. KrpnHofi.
Lenob e l u su c g t (Sa c [s )I Ta6n. XV, $rn. 4-7 Dic oelit es uagt : C a x c , 1 9 6 1 L ,^ c l p . g 6 , C arc, 19612, crp. 4gg. L enobelusv agt : l y c ro v e c o u ,.1 9 6 6 , .c rp . 6 ; . lonornn Ns 87-L71. Myeefi l4rf co AH cccp, Honocu6rpcn pexa Ke_ Jrr{Msp(Hnsosr,s Onerexa), sepxrrn Toap. .Euannos. Poctp yAJrnneuxrrft,6ynasosuAHuft,c 6nnexr{MK oxp}rr.rroMyrrG nepeqHblMceqeHlreM,c uupoxofi u uerny6oxofi 6prcurHofi5opoe[ofi, uavusarouefi-
cg oT BepuIIrHbI l{ ltcqe3arcureft B cepeaune nocJlearbBeoJurpHofi qacT}t. CuuHHas 6oposaa pasBI{Ta B anbBeonapnoft H nepeaHeft nonoetlHe nocneanbBeofispHoft vacr'. 'v'agt - BAlr - BcecorosHrtrt aeporeononnqecK[ft rpect, neonoru KoTopono BilepBbre aocTaBunn pocTpbr aaHHOno BnAa.
9l
Ta6nr.rua 33 HevepeuHrr pocrpoB
Lenobelus uagt (Sachs) (c
N J b8 7 -l _ 7 1
flapave'rprr
Kenuvnp)
87-17 2
80,o (1481) InpennonaraeMas lvcraHosJreHHarr 7 4 , 6 ( 1 3 8 1 ) 7 \,O(L775\ t" ( S e + 1 ( a r 1 6 4 , 8 ( 1ZOO) 64,r ( 1603 ) (7 L2) Anuua nocneanbeeonspHofi
finuna o6uras
TIACTI4
AHave'rp cnHHHo.6prouruoft y Bepul{Hbr anbBeonbr
5,4(1OO)
4 , O (1 O O )
,[luaverp 6oxosoii llJI'IHbI anbBeonbl
7,5(139)
4 , 5 ( 1 1 3)
Paauyc
6prouuofi
y BepBep;
!
IIIHHITIanbBeonbl npHBepuuHuoft
Anrua
2 r , 2 ( S g S )( a S 1
1 7 , 5( 4 3 8 )( Z t 7
TIACTII
Auaverp cnuuHo-6prcruuoii 1 1 ,1 ( Z O O () r O O l B Mecre Halr6onsrueno yTonrueHI,rsr Auartre'rp 6oxoeofi B N{€c- L L , 2 2 O 7 ) ( 1 0 1 ) re ualr6onbruero yron-
9 , O ( 2 2 s )( r O O ) 9 , O ( 2 2 s )( 1 O O )
tueHllg
Yron BepuunHbrft e 6oxosoil nnocKocrlr, rpaa.
42
39
OxoH.{aHHe ra6nuuu
N" 87-173
nperronaracruarr f I ycraHoeneHHafl
finnua o6uraq
L
A.nuHa nocne anbBeonlpHblr"I r{acTI{ fiuatrretp cnusuo-6puuuoti y BeprullHbl arurBeoJlbl 6oxoeoli
fiuarrerp
y Bep-
8 7- L 7 4
s 8 , 2f t i z t l
2 8 , 5t r i o e l
53,8(rZS6) (579)
73,3 ( r rZe) (637)
3,1(1OO)
6,5( 1OO)
3 , 5 (1 1 3 )
6,1(94)
aJILBeOJIbI
Ull{Hl,I
Paauyc 6prouuoti y Bep-
1 , 8( 2 8)
anbBeonbl
txlIHbI
A.rurra npuBepulrsrroii qacTu firrarterp B Nrecre
cnuurro-6prcurroii
L 9 , 2 ( O r g )( g O )
2 O , s( 3 1 6 ) ( 2 8 )
9 , 3 ( 3 O O )( r O O l
11,s(ttz)(roo)
8,G(277)(92)
LI,4(rZS)(99)
Hatt6o.Burero
yTo.;IueHuq 6oxosoii B \lccfiuarrtcrp te gatt6o,'rbrrieno yroJrLufflilt
tl
Yro.lt r3epultlrrr,tii u 6oxonoii rlnocKocrrr, npirn.
92
44
BueruHue
E p I ' I g H a K l I . l l p r , r a e p u l r H H a r rq a c r b K o p o r K a s , c o c r a B r r g e r oKono BepuI{Ha 3aHl,rMaer ueHTpanb'oe nono)Ke'I,re, aaocrpeHa. Bepruuugrrft ynon paBen 37-440 (ra6n. 33). o 6ynaaoeuaHori Qopve pocrpoB cBgAer€nbcr,B)ruor 3Har{eHI'l'I c6, xone6ruouueca B npeAenax 175-3OO"/o CB npg noKa3aTensx na oKono 580-720. CnuuHas 6opoeaa, rny6oxar u y3Kas B anbBeonspHoft qacTr{ r B Har{ane nocJreaJlbBeonspHofi vacr}I pocrpa, 6ucrpo BbrnoJra)KuBaercrl u ucqe3aer K cepenrHe nocneaJllBeoJlfipHoii rracrl,I. EpnuHan 6opoena HAer or BepuI,IHbr H llcr{e3aer npI'IMepHo raN{-)Ke, I'Ae HarruHaercrr cnI.rHHarI 6oposEa. EprcurHax crG poHa B nepeaHeft nonoBHHe nocneanbBeorurpHofi qacru pocrpa I{ B eno anbBeonfipuofi qacru ynnoueHa I cnI{HHa9 cropoHa s eaAHefi noJroBHHe nocJreaJrbBeoJrspHofi qacru pocrpa BblrryKnafl. Eoxoeole cropoubr cJIenKa ynnouleHbr, Ha Hux orle'b HerrerKo npoqBntnorcg [apannenbHble napHble lroJlocbt, I,Icr{e3arouae B npl{Bepul{HHoft .{acrr pocrpa' flonepeuuoe ceqenl'Ie oKpynneHuo rrpflMoyronr,Hoe 6es caverHoro c)Karrrr B cnnHHo--6prcruuov u 6oxosoM HanpaBreHlrrrx. 3Haqenuq 66 saxoarrrce B 11peA&. JIax 92-1O1, snaveHns BE qacro ItcKaxeHIrI BcJreAcraue ae$opMauuli anbBeonqpHoft qacrt{ pocrpa, BtyrpeHHI'Ie ''pu3HaKH u oHTo.eHe3. A n s g e o n a n p r r M a f r ,G a H H M a e r L/5 anuubl pocTpa' BepuI'IHa cMerueHa x 6prouuofi cropoHe (6prougror paqlryc nopnaxa Or3 CE), anbBeonspur,lr? ynon 3aMepuTb He yAanocb BcneqcrBr.re Ae6opvauuft s o6nacrll arbBeontrt. Cuaftxa y Bepul{Hbr oJlbB€onbr co cnnuuofi cropoHbr BblAaercrr B [ocneaJrbBeoJrrrpHyro r{acrb pocrpa npnMepHo ua AnI{Hy afibBeoJrrrq Oce_ Bag JII'IHI'Iqnor{Tl'l npgMafl' npu6nrxena K 6proruuorl cropoHe, nocreneHHo oTxonI,IT or Hee e canuel:r qacrl'I pocrpa. Ha HaqanbHr,rx craalrrrx pocrpr,r nprr ALIaMerpe oKg. no 2 MM I'IMeIor cna6o Bb[pax(eHHyro cy6xoHHqecKyro fla oxono 4OO-4SO +opvy, B JanE'Hel:truev pocrp craHoBl{Tca cy6uunnHApiltrecKnN,r, [pI,I ar{aN{erpe 3 N{M lla Bo3pacraer Ao 12OO. flpu C6 4vv l-la coctaBn rer 15OO, pocrp craHoBr.rrcrr cnefKa BepereHoBlAHbllt'I (c6 6 vrvl, na IOOO) a no Soprue He oTJrr4r{r.rMoT MonorllrLx pocrpoB rlpyflrx BI'rtroB I.errobelu.s (a nepByr.o oqepeab L.nttrtrtet,at, To)Ke c KopoTKoii 6prctuHoir 6opocaoir). flocne ororo npolrcxoauT 3HaquTenL,Hoe B3tryrr.re pocrpa B el-o saaseir noJroBr{He, xapaKTepHoe AJrs AaHHoro Br{aa (puc. 35). l'lcveHquBocrFr. Hau6onsrueii ncveuql{Bocr}r noABep)KeHa creneHb B3ayrlrrr pocrpoB B nocJlearbBeonqpxoii qacru. t{apany c pocrpaMr.r yMepeHHo B3Ayrr:rMH r/ 3
fla,
t,* oo.) r*ercrcrr oK3eN{ilrrepbr 6onee B.qyrbre (pocrp
lnlo)_*-t:tl \,r 87-173 v Ap.), a TaKxe N{eHeeo"ryroro, ^;;;"-;;;;; #;:_ napuoii qacrblo (pocrp N 87-1 74 u ap.). I4sveHqusa " raKxe crene'b pa3BrITlIn 6proruuoii 6oposAr,l, ecrb pocrpbr, y Koropbrx 6prcuruaR 6opos_ 'qa 3axonltr B 3a,a'HrcIono.rIoBI4Hynoc.teaJrL,Beo.lqplrol'rqacrtt pocrpa. c p a o H e rr lr e. Poc'p st [ ..7'n91 pe3Ko or.T'rqarcTcs or !p]T!rx BI'aoB pola /.r',obelus no cnoer-r 6y.'ranonuanoii Soplre. Kponro ,o"o, 6pnuuaa 6opoaaa y HIrx He 3axorl{T Aanee cepenuHbr nocJrea.lL,ReonirpHor'r qacrll. oauua'rrbHarr n puypoqeHHocrl. pocrpi,l L.uagr o6Hapyx<errul TOJIbKOB npeAe.qax ,rleuo4neHeKcKono paiiona B n.rruHlrcrbrxoauusx Plrc. iJ5. Ilpoao.lruxoe ceqeHHe pocrpa eopxrlrii Toap, p. Kealrwrrp( x O,ZS)'
L e n o b e L u su a g t I S a c h s ) .N q8 7 - L 7 4 ,
orKpl,trono
Nlopq.. Hu oanrono pocrpa u 6o.ree \teJrKoRoAHbrx tr nprr6pexrrr,lx 0auusx o'{eell.0.Ilo pacc\ttlrpltBaelu,rfi RIi[! Ber ceo6oAHonlaaanurttir o6pa:l X\U3HII B yC.-IOBUqx OTKpr,rTofo \rops. Haltrlt
IIe yA''l"Iocl'
Bospitcr If reoI- patfrrrqecKoe pacnpocrli..tHcHlte. Bepxuuii TOilJllrtrxHui.i aaJrcrr C-eoeptror.i Crr6rrprr. A 1a r e p I I a n ' 4o pocrpoB II3 BepxHer-o Toapa-HlrxHefo o . 1 l . n e t . rp p . K e n t r r r r i r y rl r - c6opr,t T-u- Ilrtrtr,rrrenori, T.ll. [iyoP-3exllr KrrllrHoii, t).A. Elraxlrena lr c.u,.,\te_ Jrc.fI,fHoH.
93
L e n o b e l u s n t i t t ae u a e ( S a c h s ) I
Ta6n. XV, $nn. 1-3 D i c o e l i t e s m i n a e l , a e: C a K c , 1 9 6 1 1 , c r p . 86; C axc, 1961_2,crp. 433. -L966, crp. 65, ra6n. 6, $un. 6, ta6n 7, Lenobelus ntinaevae .' I'ycror'aecoB, 1 3 . $ur. I'ono'run Ne 87-L79. Myeeft Hff CO AH CCCP, Hoaoctr6upcx. Pexa KeJrr{Msp (Hnsoerg Oaeuexa), aepxuuft Toap. Poctp cHJrbHoyanuHeHHr,rft,aepereuonuAxurft, cxaT r,tir c 6oxon, Ar,rarHog. c npuBepuunnoft 6prourHofi 6oposaofi. Cmassan 6oposaa HaqlaHaercn or nepeAHeno KOHUa pOCTpa I{ 3aXOAttT nr{tub HeHaMHOnOB nocneanbBeoJlslpHylo r{acTb pocTpa. n p u 3 H a K u . f l p u n e p u t { H n a r rq a c r b K o p o T K a r I ,c o c r a B J t n e r o K o J r o BHeusue fla; aepuruna sanuMaer ueHTpanbHoe noJloxeHlle, 3aocrpeHa, Beput'IHL/4-L/3 (ra6n. 34). O neperexoeunHofi Sopr"re uuft yron HaxoAuTcrr B rpeaeJrax 34-470 pocrpa noBopsr suaqennrr c6, cocTaBneroruue135-L75"/" CE npu noKa3arennx na or 600 go 95O. CnuuHas 6opocaa rny6oxan u ysKas B anbBeonnpuofi r{acr[, sareM 6rrcrpo Bbr[onaxunaercrr Ir saxoAI{T B [ocneanbBeoncpuyro qacrb y 6onree AJrtrHbr.Eprcuruan 6oposaa HaqnHaercn ruuHcrBa pocrpoB Ju{rxb Ha L/4-1l5 y BepuuHur, 6urcrpo BbmoflalK[Baercrr u He 3axoALIT aanee L/ 5-I/ 3 nocJleanbBeonspHoft qacru pocrpa. 3a npeaenaMl{ pacnpocrpaHeHun sr}tx 6opoca 6prcuHan I{ cnI{HHaa croponbr BbrnyKnure. EoxoBr,re cropoHbr cnenKa ynnoueHlrl, Ha Hltx npo-
Ta6nuua 34 H s v e p e x N f l p o c r p o B I - n r c b e l u s n r i r t a ( t ' t t cS a c h s
N'87-179, p.KenuttnP
\l 87-180, p.Kerurvnp
1 1 7 , O( 1 8 8 7 ) 111,5(180O)
Annsa f np.ono*raeMarr o6uag 1 u"ru"ouneHHarr
106,0 ( rooo) 1 0 G , O( L O 6 O)
ArurHa nocneanbBeonqpHoft qacru
8 2 , e ( 8 2 e ) ( 6 1 4 ) ( 7 8 ) 9 3 , 3 ( r S O S )( A S 6 )( 8 0 )
Anar.aerp cnuxxo-6prcuHoft y 'BepuHHbI anr'Beonbr Auaverp 6oxonoii y BepuuHbr anbBeonbt Paauyc 6prcruuofi y Bepul{Hbr anbBeonbl nnuua npl{BepuunHol"r
1 O , O (1 O O )
6,2(100)
8,8(88)
5,3(85)
4,8(49)
2,4(39)
(-
29,9(299)(3C)
2 8 , L ( + eO )( 3 0 )
13,s(rss)(roo)
1 O , 9 (r Z O ) ( r O C l
qacTLl
,[,uar'rerp c nu HHo-6pburxof,r s Mecre ualt6onuueFO yTOnrueHHrI Arar'rerp 6oxoaoir e N{ecre ualt6onuruero yronueHurr Ynon BepruuHHblfre 60xoeoft nnocKocru, npaA. lBuo
94
Ha3BaH B qecTb
1 1 , 8 (1 1 8 ) ( 8 7 )
I'eonoFa
47
9 , 7 (1 5 6 ) ( 8 6 )
34
K J . l l . A \ r t H a e n o i i , n e p e o t ' l AoCTaBIIBIUOI-!
eI.O pOCTJ)I'I.
Puc. 36. flpoaoauuoe ceqeHxe p o c r p a L e n o b e l u s m i n a e a a e( S a c h s ) Nfe87-180, Hu)KHHftaaJreB p. Keauvap (x Or7)
CfleX{I{BAIOIC5I [apannenlrHbre napHbre nOnOCbr, paCxOAgtItUeCsB npr{Bep_ ruHHHOftqAC?r pocTpa u EcqeSarcrrlHe npn np[6nr{}KeHr{r{ K BepumHe. B nonepeqHoM ceqeu[n pocrp no Bcefi Anr{He cnerKa c)Kar c fuKoB (EE 85-97, 66 86-94). oopua norepeqHono ceqe'r{rr oKpyrrre'HonpnMoyreJrbHas. BnyrpeHHIfe nplrgHaKI{ I{ oHToneHeg. Arrb'eonaflpgMas, 3aHltMaer' oKono 1/5 lnrnbl pocrpa, BepuI{Ha cnenKa cMerrleHa x 6prcuxofi cropoue (6pnruHoft paa'yc gg-4g% c6). Ansaeongpn'fi yren He yAanocb ttgMeptlTb BcneAcrBue aefupMaur.rfi g o6nacru aJrbB@rrbr. cnafixa. sa6nrcnaercs co cnusHofi qropoHbr poopa, nepercoAur B nocneanr'B@Jlrrpryp eno qaerb npHMepHO Ha Arrmry anbBeonbr, OceaaR nvilurl noqr[ npsMas, npn6nnxeHa r 6prougofi cropoHe. Ha Ha.fanbnbrx craqngx pocrpbr r{Menr{ cy6unnrnAprqecKyb fup}w (npr cE s,z r,.nu IIa coc_ TaBnsna 113o)r B AanbHefiruev pocrpbl npro6peralrrr cnaqara BepecnegoBs4gylo 0opvy (nps CE 5r2 vv IIa - lgSS; c6 - 1g1, rd - lOSg), a sar€M n sc'o BbrpaxeHHyro BepereHoBnllH)'ro o0pr"ry, csoficrseHrr)4o B3pocnbrM oco6su (p"".- io), Hsr'aeH qaBoc rs. Han6o[ee ,,sMe'quBbr y paccMarp[BaeMoro BuAa creuelrb B3ayrl{s nocneaJlbBeoJlspHoft qacrll H cr€rreub c)rcrrrrs aJrbBeoJrspxoft qacr1, pocr! . pa. B ueyvexEofi-KoJrJreKul{u r{MetoTcg SKSeMnngpbI, Jlulllb cna6o BeperenoBtrtrHbte, nprr.6nnx
95
P o c r p u L . m i n a e u a e o 6 H a p y x e H bBl o c npuypoqeHHocrr,. OauuarbHarr HoBHoM B r.nr{Hr{crsx @uurrx orKpbrrolo Moprr s IeHo-OneHeKcKov paftone u pe)Ke B npeAropr'nx BepxorrHcKono xpe6ra. EauHH.{Hbrepocrprt Hafinegbl B 6onee MeJrKoBo,q,Hbrx$auusx na p.Moroptryne s6flfisll 6epera ppcKoro Marepl{Ka. Cneaoeao6pas )KI{3HI,IH npeatro.Iltram{ o6TeJrbHo, )KHBorHbre BenH ceo6onHonnaearcultfi pafioMops. B NaenxoeoaHbrx@urnx Aua6apo-XaraHrcxono craHoBKr.r orxprrrbro paftone oror BIl[ He HafiAeH. Ha, a raK)Ke s Oxorcxo-OvonoscKoM p a c n p o c r p a H e H t { e . B e p x n u f iT q a p Boepacr u reot.pa0ntlecKoe nnxuuft aareH Ceaepuoft Cu6xpn. M a r. e p u a n, 47 pocrpoB I{s BepxHero roapo-HnxH€I-o aaJIeHa pp. Kenuvep n Eyop-SeKldr - c6opnr T.H. HannuReaofi, T.H. Kupruofi, C.B. Menearnofi, P.A. EunxleBa u lO.V. Mnnaeaof,t, 2 pocrpa I{3 Bepxnel'o roapa p. Jlenucxe, 2 pocrpa us HI{)KHenoaaneua p. MoroprryHbt - c6opul T.H. Kupuuofi.
CTI
0u
3e co qel
P o a P s e u d o d i c o e l i t e sS a c h s , 1 9 6 7 Dicoelites (pars): Caxc, 1961a, crp. 7 7 . I l y c r o M e c o B , 1 9 6 6 c r p. e4. Lenobelus (pars) : Pseudodicoelites; C.axc, 1967 1, crp. 44O, 19672, crp. 20.
nq
BE tul
pe lx?
poaa. Dicoelites bidgieui Canc, 1961a. Hnxnuft aanes neBono rtpuTun toxa lleubr - p.3exnT. Onuc aHI{e. Pocrpu cunbHo yaJruHeHnbre,BepereHoBl{Aunte,6ynaBoBl{aHble unu cy6uunruApuqecKue, cxarbre c 6oxos urru c oKp)rnnblM nonepeqHbtM c€H€HIreM, c rny6oxofi cnuHuofi 6oposaofi u cnafixoft e nepeauefi tracTu pocrpa. B saAHefi qacru pocrpa 6oposAr,r uet. BprcruHan 6opoeaa lrJrl{ ynnoueHue Ha 6prcurnofi cropoHe B nepeaH€fi qacru pocrpa sa6ruoaarcTcrr He y Bcex BHAoB. Ha 6oroerrx cTopoHax pa3BuTbr napannenbHbre napHbre floJrocbr, pacxoastunecs u 3aTyxapuHe B npnBepu:unnofi rracrrr pocrpa.
CI
Ps. platyuentriosus, Ps. hibolitoides, Ps. clauatoides, Ps. gustomesovi, Ps"primoris). Beponruo Haryque ceAbMoro BI{aa, ofincbrBaeMoro Hllxe KaK P'seudod,icoelites sp. 'Hibolithes" He Apxruqecxofi KanaAsr Eneuxuu (Jeletzky, 1966) yxa3bmaercs sp. u3 Toapa-aarreHa, Hatro nonaFarr,, orHocsru[fics x poay P'seudodicoelites" CpaeHeHue. Cpaaxenus c poAoM Lenobelus Aaxbr nplr onucanl{u rrocneArlero. flpr orcyrcrtsuu AaHHbrx o [o[o)Keuur{ cu@Ha ao $paruoKone tr ocesoft nrflocnee Brru pocrpa Pseudndicoelites nerKo Moryr 6urs crryraxbl c Hibolites. BHelUHeTOBr{AA, HO OTJIl{qaIouIIecs paSBur[eM B HUe I{MeIOT pOCTpbr raKonrxe nepeAueft qacru pocrpa He crrsurroft, a 6prcrurofi 6oposlu H coorBercrBeuuo cMerueurreM ocesofi $trnuu .n cropony erofi 6oposAbr, a He uao6opot Kax y Pseudodicoelites. pactrpocrpaHeHrre.Toap - nnxrrft Bospacr u neorpaSsqecKoe CCCP u npetrrlonoxurertbrlo AIrKm6aftoc Cesepnofi Cn6npu, CeaepeBocrora .recxofi KaHaAu. Ha cenepe Cu6rpu T.H. Hanungesofi c[erlaxbl elu{Hnqrlbte HaxoIF MeKt{ pocrpos Pseudodicoelites sp. B Bepxxeur 6are. K Pseudodicoelites xer orHocrlTbcs pocrp t{s Bepxueno aanena 3er'rnx Qpaxua Hocn@, oupeAenennrtfi (lln6nep u lllyntr'rxa, 1960). xax Hibolites (?) cf.. beyrichi Oppel
B€
KA
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HO KA AE
po ro II(
HC
K€
e€
c1
M
AJ MI
ul Qt
6( p( 6e
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1 c' II(
1 96
P s e u d o d i co e l i t es b i d g i e u i ( S a c h s )l ) Ta6n XVI, $r.m. I-4. Dicoelites bidgieui-: caxc, 196la, crp. g4, sun. 1_g; caxc, 19616, crp. 433. Lenobelus bidgieui.. lycrorraecoB, 1g66, crp. 65, ra6n 7, $ur. L4. Pseudodicoelites bidgievi.' caxc r HanraRena, 1g676, crp. 2r, Ta6r. 2, Srr. 1-3. lonorrn Nq B2-L. Mysefi Hfr co AH cccp, Hoaocu6Npcr. pexa Eyop_ Sexzr (Hueoarg p.Ileusl), HrxRf,ft aaneH. au ar H O 3. Poctp Bepe?e'oauluurft, cuJrbHo yIrnl{He'Hsrft, cxa*,rri c 6oxon, co clll{Hlroft 6oposaofi, rAyurefi ot nepeAHeno KoHua u fipoTrrrusarcuefics 6onee qeM Ha [onoBlrHy nocneanrrBeonspHoft qacru pocrpa. EprcruHaa 6opoeaa oqerrb He._ nny6oxas, na6rnoaaercrr B rrepeanefi rperu pocrpa. BseruHue ''pr{3HaKr.r. [puaepu,"r".r.""t, oa'uMaer oT L/4 Ao L/g [a, BepuI{Ha pacrono)KeHa ueHTpaJIbHo l{rut cnenKa cMeuteHa K cnuguofi cropoge, BepurrH'rrfi yron paBeH 31-380 (ra6n.35). Beprunra pocrpa ryrIb cJrenKa 38ocrpeHa y ocrpl{s' na6ruonarcTcn upoaonbHble MopurHKu. Crenesr Har6onbueno pactxlrpeHl{s pocrpa oflpeAeJuercrr TeM, qro c6 cocTaBneeT 160_2007o, peAKo iso.t" cE npu sHaqeH[sx na 760-920. cmluHas 6oposaa Ha BceM nporsxeguu nny6oKas !! ysKas, pacu[psercs JII{lrb npu npr6JruxeHl,r[ K nepeaxeMy KoHrIy pocrpa. Boposaa gTa flpocfle)Kl{Baerc s Ha 5/ I anaw'r lla. Epror,uxaa 6opoeaa B aJrbBeonspxoft qact,n pocrpa uMeer Butr tuupo*oro y,,norue',.s, onpa'r4rre'Horo aByMs B.trrlrKaM[ r saxoat{T B nocneaJlbBeoJurpHyro qacrb Jrr{rub Ha L/ Lo ee anrrr{bL 3a npeaenaM[ pagBllrllg 6oposa clI{HHas u 6prcruuafl cropoHbr Bbrrly6nbre. Eoxosure crr po'bl ynnolue'bl' Ha HI{x BI{IIH'' napaJrnelbHbre naplrbre [onocbr, rrAyr![e BAonb Bcsno pocrpa' cJIenKa pacxoAgul[ecg I{ Tepffolrluecs B nplBepulungoft eno gacTl{. rloneperxoe cetreHl{e oKpyrne'Ho-Tpaneur{eAanb'oe, cxaToe c 6oxoa, c pacuupe'xofi 6poruroft cropoxofi. BnyrpeHH,ne n p I { g H a K r r u o H T o n e H e s . A n s B e o J r an o q r [ [ p r r M a s ,c n e n Ka I{3oI'Hyra K 6prcrunoft cTopoHe, saxl{Maer oKono 7/ G annft,r pocrpa, BepuI{Ha ee cnenKa cMeueHa x 6prcuJlroii. cropoHe (6prournofi paArryc gg..47.i. CE). AarBeongpHbfi yron paBelr 2o-24o. cnarlxa Ha6rnoaaercs Ha cmansoft cropoge, Bbrcryllag uottrl't BABoe 6oagrue trJrI{IfbI anbBeoJlbr B nocJreanbBeonsp}rylo qacrb pocrpa, Megee qerKo Bbpa)Ke'Has cnafixa, Bbraarouagcs B nocJreanbBeoJrsplrl4o qacrb Ha lm{lry aJrbBeoJrbr,t{Meercn Ha 6prcuuroft cropoHe pocrpa. Ocesaq ilfiHus noqrr rrpsMas' cnenra npn6Jru)l(eHa r 6prcuaofi ctopoue. Ha na.ranbHbuc craAr,lcx pocTp cy6ultrlrHApuvecxufi, cpaBHnT.,enbrroroporxnfi (npr CE 2r7 uu sHalleHlre IIa oxoao 460), aarlee crnbrlo ylrrrrrser", irp, CS 5,S vr.a rla oKono 12OO) rl rpr CE 6onee 5 r.au upro6peraer caa6o BeperegoBnArr)rrc, Oopvy. pesxo Bbrpaxelrnas Be.per€HoBI{IlHocTb rlosBJlsercs Jll{lllb npu npz6Jtrtx(enl{I{ K pasMepaM Bspocnbu( oco6efi (prc. g7), H g ve Hqf, Bo c rs. Ha6rnoaalorcc pasflrqus B cr€[exu Bbrpa]Keuxocrt{ 6prou'rofi 6opoelrt, f,Iroraa HaMeqarcueftcg B Blrae ynnouenrrs pocrpa B ero nepeagefi tracrr{, l|Honlla xe Bblpa)Kerurofi locTarotlrto oTtleTrrt{Bo, Ho BcenAa rurpoxofi n uerny6oxofi. CmrHHas 6opoe[a, Kar( [paBnno, nrqr6oKas r{ ysxas, y HexoTopbr.x pocrpoB (wp 87-186 u ap.) Toxe 6rrsaer urpoxofi n nenny6orofi rr saxaglrrrBaeres, He aoxotg Ao cepe[[nbr rr@flearrbBeoJrspnofi qacrr. Baprrpyer r crerrerrb BsAyr\rs pocTpa B eno [ocJleanbBeoJrspnofi sacrlr, llpeo6naaaror.snaqegtrs c6 uopaaxa 175-180' Ho ecrb pocrpbt (m az-rgi),' y *otopor* c6 6onee 25o u coorBerL crBeHHo 0opva pocrpa craxoBrrrcs 6ynaaos[Anofi. b"r", xao6opor, pocrpbr c c6 [opsAKa 160 (Ne 87-188), BepereHoBt{axocrb Koropbrx BHpaxega cna6o. Ke
-
1 tBna
7 4 9
HagBaH B qecrb reonora
P.A.
Enax,.eBa,
BnepBbre AocraB]rBrr,er-o eno pocrpbr.
97
Ta6nuua 35 oelites bidsieai Sachs I4svepenns pocTpos P seudod.ic
ls g7-195, P.EYoP-$eKrr
f upeanonaFaeMas A'MHA ,| o6uas'l IYCrauoaneHHas Annna [ocneanbBeounPnofi qacT'l
Y Auaverp cnnuro-6Prcunofi BepulrHbl albBeotrbl Auaverp 6oroeofi Y BePut{ItbI
1 5 5 , O( 2 2 L 4 1
1 1 4 , Or(A r o )
1 5 1 , O( 2 L 5 7 ' )
1 1 4 , O( 1 8 1 0 )
1 3 2 , O ( r a e o 1 ( 9 1 9 )s L , 7( r + s o )( e r e )( 8 o ) (as1 7 , O ( 1 O O)
6,3 ( 1OO)
6,5(93)
6,3( 1OO)
ANbBEOJIbI
Paaryc 6prcunofi Y BepunHbr anbBeoJbt Arnura nplrBepumnoft qacfl{ cnnnno-5Prcurnofi Ataverp B Mecre uan6onguero YToJtF ueEgg. .E,uarr,retp 6oroaofi B Mecre xau6onsurero yroJltuentrfl 'B cIrI{HYron aabBeoJlgpnrtft uo-6pnunoft nnocxocrl{r rPaA' Vron Bepuulrnbd s 6oxoaofi nJI@KOCTil, rpaa.
35,9(513) (27) 1 2 , 3 . ( 1 7 6 )( r O O l
Lt,2(too)(91)
2,9(461 '28,1 GqA)(26)
L \ 2 ( r z e )( r o o l 10,8( 1? 1) (96) 20
32
32
nnrsnaubr Beposrlro ae$opt.tau[sMu pocrpa B ne6aHus B snatrensu EE (gS-fOO) rracTll. ero aJll'Beonspnofi Pocrpsr Ps.bid.gieui auaersrorcc cpealr apynruc BIraoB Pseud,oCpaaHeHue. Bblpaxensoft BepeTexoauAxofi $opve u [plcyTcTBl{lo B oTqeTrrl{Bo no dicoeldtes nepeanefi qacru pocrpa 6prcurxoft 6oposArt. O aura nbHa s npuypoq eHHocru. Pocrpu Ps,bidgieui s octtoBuoM co6pauu B rnuHuctorx $aUunx oTKpbr1org Mopc s JIeEO-OneHeKCKoM pafione. Equnu.rHrre pocrpbr o6napyxexbl B Bepxonnre u Ea uo6epexre Oxoccxoro MOps n SaUunx ClTKpbrToro MOps. OrAensxue EaxOAKtI neb Becrgbr e pafione Ana6pcxofi ry6rr u Ha neBbtx np[ToKax llenst s 6onee MenKoBo[Hux Sauuex. OqesuaHo xuBcrrHble Benu cso6oaHonnasapulufi o6pae )Kr{gtru B ycnoBucx qrKpblToro Mops. BoepacT H reorpaQu.recKoe pacnpocrpaHeHue. Bepxxrfi Toap-Hr{xunfi aanen ceaepxofi cu6npu. n. '62 pocTpa us Bepxgera Toaps-HI{XH€r! aanega peK Matepua n Eyop-3eKur - c6opsr T.V1. HanbEsesofi, T.H. Kupuuofi, Kenm,rnp P.A. Ert[xueBa u c.B. MeneAnxofi, 1 pocTp Il3 BepXHerO roapS - llHX((npaa51fi npI{ToK J]ensr) - c6opst T.H. KupuHere aanega p.$eruAxan aanena Ana6apcxofi ry6u - c6opbt T.LI. HarnpocTp HuxHeFo lts Hofi, 1 HReaofi, 8 pocrpoB lrs Hr{xxer.o aarrena p.Bnnrnu (no6epexse O:torcrorp Mops) - c6opu A.C. Aalnca.
Puc. 32. I-lpoaorrbHoeceqeHr{e pocrpa P'seudodicoelites bidsieui (Sachs). Huxuuft aaJIeHr p. Eyop-3ers{r ( x O,8) lS 87-185,
98
II e 87- 186, p.Kennvrp
N e 8 7 -1 8 7 , p.Kenuvnp
N e 87-188, p.Kernrvnp
(rzoe) 123,O 123,O(17O8) 1 O 2 , O( t + t 7 ) ( 8 1 6 ) (83) 7,2(1OO)
ro6,4 t inruI
1 0 6 , 4( 1 9 3 5) ( z a s I
9 O , O( 1 5 O O) 88,O(1467)(eoz)
5,5( loo )
6 , 0 ( 1 o o)
7,8(1O8)
6 , 0 ( 1 1 0)
5 , 3( 8 8 )
3,4(47)
2 , 3( 4 2 )
2,9(48)
3 4 , O ( + Z Z )( S e 1
1 2 , 5r(z a )( r o o )
3 2 , 2 ( S e s )( 3 O ) 1 3 , 9 ( Z S g () r o o )
3 2 , 6 ( S q g ) ( s z )\ 9,7 ( L62l ( roo )
LL,1(163)(g+)
r L , 7( e r s ) ( 8 4 )
8,3(138)(ao1
31
31
3B
P s e u d o d i c o el i t e s p l a t y v e n t z i o s u , sS a c h s l Ta6n. XVII,
0rlr,
L-4.
Pseudodicoelites platyuentriosus..
Carc u HansneeBa, Lg67
2,
crp.
2L.
fonorrn Ns 87-189. Mysefi l4ff co AH CCCP, Hoaocu6rpcn pera KenuMsp (Husonss p.OneHex ),-Huxnrfi aareH. AuanHos' Pocrp cJIenKa BepereHoaraurrfi, noqrr{ cy6unnnrApuqecKnfi, oqeHb cuJrbHo yAm{HeHHUft, cXaGlft c 60xon, co cm{Hxoft 60posaofi, uayurefi oT nepea_ HeI-o KoHua llo cepeal{Hbl nocneaJrbBeorupHofi vacru. Ha 6pnruxoft cropoge B rrg.peanefi qacra poc?'pa y[nouearre unv Herny60xaR 60posAa. Bseutnue npI{3HaKI{. rlpunepurHHas qacrb pocrpa saHuMaer oKoJroL/4L/ 3' I-la' BeputtrHa pacflonoxe'a ue'Tparb'o, cner*a oKpJrrrreHa, Bepufin'brft ]rnon (ra6n' 1&230 sd). Husxas creneHb Bepere'oBua'ocrr{ pocrpoB o[peaensercs sHaqe'Hsvu c6 14s-166% cB np' oqeHb BbrcoKr{xnoKasaTensx na (11oo_ 1350)' cnurnraa 6oposaa Ha BceN{ npoTrrxeHur rrry6oKas r{ ysKas, B nocneanEF BeoJltlpHoft qactu pocrpa HecKonbKo pacu,Hpsercs r{ aanee BbrfloJlaxl{Baercg. 3a npeaenaNrn ee pasBlrrl{s cnHHHas cropoHa BbnryKJr€rs.Epuuuae 6oposaa oqeub MeJrKas I{ lul{poKas B anbBeonepHofi qacTu pocrpa, saxoar{T Ha L/ IO-L/S e nocneanbBeonspHylc' rracrb' y MHorHx sK3eMnnspoB npeAcraBJrser xulxb ynnoruegue ua 6proruHor?cropoHe pocrpa, Koropoe 6urcrpo r{crresaer. Erutxe K sa'Heg. nono_ I
Plorrurntriosus
(nar.)
- nnocxo-rry3aruft.
99
Ta6nnua 36 Hsuepexus pocrpoB Pseudodicoe lite s platyv entriosus Sachs I.Ie87- 190, p.Kerur'lnp
),le87-189, p.Kernrunp'
flaparuetpu (tp"atto*naeMas ANI|HA
'
o6ruag 1 LycraHoBneHHas Anuxa nocneanbBeoflspllof, .ractn ,[nar.,retp cmrgno-6ptoulHofi y BePuuHbl a'IbBeo-
L22,L(24421 1 1 O , 3( Z Z O A )( 1 3 2 9 )
95,O (2O2Ll-
92,O(rgsz)(ress1
5 , O ( 1 O O)
4 , 7 ( 1 O O)
4,7 (94)
4,8 ( 102 )
JIbI
.[uaverp IUltHbI
Y BeP-
6oroaofi
atlbBeolbl
2,O(431
Paaryc 6prcrunofi y BePlIIllHbI anbBeonbl
Ilnrxa upnnepurnxofi
2 5 , 1 ( S O e) ( 2 3 )
32,4(689) (3s)
qacTlf
Ilnar'rerp cmmno-6Prcuuofi B Mecre gau6onuruero yTorlrueHrls .[rar.,rerp 6oxoeoft B Mecre gan6oasuero yroJIIIreHxs Vnon anLaeongpurrft a nnoccnnrgo.6prcuuofi xocru, rpaA. Vron BepuuuHufi a 6oxoBofi nnocKocrl{, rpa&
8,3 ( 166) ( 1OO)
6,8( 14s) ( 100)
7 , O( 1 4 0 ) ( e + )
6,0(149)(88)
19
23
B11Hepocrpa 6prcunras cropoua BbrnyKnag. EoroBble croponbl HecxorlbKo MeHee Bbrrryxtrbte, B [epeAxefi 'ractn pocrpa ynnouleHbl, napanJtenbHble.napHbte nonocbt tlacrlr pocrpa HeHa rrur( Bbrpa)Kegbr cna6o, upn npu6mtxenrl{ K npuBepurururoft cKolrbKo
pacrcoA*Tcs
It, He Aoxogs
Ao BepIul{HbI'
SaTyxaKtT.
floneper{Hoe
ceqeul{e pocTpa a6nuen BepuJllHbl anbBeonbl oKpyrnenuo-npqMoyronbnoe c rny6oroft sget{xofi co cnmruofi croponst. B6nn3rr Mecra uas6onrurero pagugpeHus pgcTpa erto no[epeqgoe ceqexue I{MeeT SopW cxaTono c 6oroa oBana, pexe oKpyrnoe. B X y T p e H H U e n p n g H a K U H O H T O r e H e S . A 6 s e 6 n a f l p s r v r a s3r 8 HuMaer oKono Or 1 anunbr pocrpaf BepuMHa ee He3HaqurenlrHo cMeuleHa. x 6prcuruofi cropoxe (6prcuruofi paqryc 6onee O,4 CE). AnbBeonnpustft yron paBeH 18o. Cnafixa xa6npAaercq TOnbKO Ha cnxnnOfi sropone pocTpa, BbteTynas b ctopOny nocneaJllBeonspHofi qacln npuMepno B 2 L / 2 paga AaJrbue [nuHbl aJrl'Beonrt. Ocesas nuggg floqTu npgMas' npn6nnxena K 6prcruxofi qropoHe. Ha naqaruublx cTagusx pocTp cy6UunuHApnqecxnfi yAnIIHeHHrtfi, yxe npu CE 2r2 uu fla orono 2OOO. Cna6o esrpaxeEEyro BepeTeHoBualryp oopr"ry pocTp npno6peraeT, nuub npn6nrxaccl' K pasMepaM aepocnoft oco6u (puc. 38).
pnc. Sachs.
r00
3 8 . I l p o n o J r b H o ec e q e n u e p o c r p a P s e u d o d i c o e l i t e s P l a t y u e n t r i o s t r s ( x O,8) nepxunfi Toap-Hlrxsnfi aaaeH, p' Eyop-3ers{t lS 87-192,
I J b87- 191, p.Moropqysa
N e 8 7 -1 92, P.Eyop-$eru"
6 4 , O( 1 8 8 2 )
L O 7, O( 2 O1 9 )
60,9(1791)
5 6 , 9 ( r o z s( )r r s a )( e g ) 3,4(1.oo)
tot,7(1938) e6,o(rer1)(1116)(90) 5,3 (1oo)
3 , 7( 1 o g )
5 , 7 ( 1 o 8) 2,2(421-
1 3 , 3( g e r ) ( z g )
2 9 , 4 ( S S S )( s r )
s , o ( 1 4 7 )( 1 O O )
8,6 ( L62) ( roo )
4 , s ( L 4 4 )( g e )
7,8(L47)(91) 18
22
18
I'|sw"t**ocrb' HsMeFg[BocrIf no'Bepxena o6uag Qopr.ra pocrpoB oT [oEru cy6unnmlp[qecxoft Ao oTqerJrltBo BepereuogtrAxoft, CoorgercrBexgo sHaqexf,s c6 vesspTcs oT 145 ao L70. g"pr"py* y'n,,He','ocrr pocTpoB _ I-Ia -oo uMeer sHase'r{s or 1620 "r"n""" ao 22oo, ;; - or-uoo lgso: Mergercg ?arxe cTeneHb 60rosono cxaT,ns pocrpoB - EE g4-10g, g4_gg. g pasrunrrof, 66 neHI{ Bblpaxe'a 6prcruxan 6opoeaa cTs - oT y[noule'I{s B aJrbBeoaapnoft qacrf, pocrr. Pa ao uuporofi u uearofi 6opogAst, [pocnexlBarouefics '-vqvr'v4 p r no"t""rrbBeoJrgp]rFo *\rurre qacl.b pocTpa aa
or2
ee
AJrrHbI.
cpa eHeH Ire' Pocrprt Ps. platyventrl'oszs oTtrernrBo oTflItqarorcc or BltAoB po'a Pseudodicdelites Apynlx ceoefi Jltrtrrr' cnenxa Bepere'onnruofr, ''orrrr cy6uuflrHlpuqecxofi fuprrrofi rrpa oqenb cuJrbtrofi ynmrnenrtocTt{, Har6onee 6rugor no o6uteft fuprvre pocrpa Ps'hibolitoides, y .*"pono o[Haxo Her Ha pocrpe 6pp'rHofi 6oposrlbl' caMuft pocrp MeHee yAJrrmelrrtrrft u 6onee orqerJrrrBo Bepere[oBn'F
t0l
Eyop3O pocrpoB I{3 BepxHeno Toapa-aaneHa peK Kenuvnp u Marepr{aJr. pocrp us l Meleaunoft, c.B. U KUpUnOft T.H, HantHqenofi, 3exnr - C6Opu T.14. 6al"toca p'MotoptlyHbl Bepxger.o Toapa-aaneHa p.IIewIcKe H 1 pocrp I{3 Hu)KHero Toapa-HI{)l(F BepxHero I{3 pocrpa (npuroxll p.Ileurr) - c6oprr T.H. KrpuHofi, 2 A'C' Aanuca' c6opul Hero aaneHa 6accefina Ovonoua s SachsI P s eudodic oe li t es hibo Ii toi d'e Ta6n. XlX,
Sur.
1-3
3' Hastites claaata.' BopoHeu, L962,Ta6n' 6O, $ur' 1 9 6 7 6, crp. 27', ta6n' 2' P s e u d o d , i c o e l i t e sh i b o l i t o i d e s . ' c a x c , H a n t H R e e a , 3' fiut" 1-3' ra6n' 56, crp' LgTg' HanunleBa, Srr. /*;6; Menenrsa u '
Ioxnuft 6epen N.: 85-6. Mysefi Hlf co AH CCCP,Hoeocu6Hpcr. fonorun Ana6apcrono 3anrrBa' aa.neu. c POCrp CJterKa BepeTeHonlaAnurft,CUnbHo yAnI{HeHHufi' Cxalrtft AnarnOS. eacpOCTpa' B npueepu:uHuoft UOnOAEHe 6oxoa, co cnunuoft 6oposnofi B nepeAuefi neT. ynnoueHufi HmI Tr{ pocTpa H Ha ero 6prcuLnofl cropoHe 6opo3A o K o J I oO ' 3 n p I { 3 H a K I { . f l p u a e p u l { H H a gq a c T b p o c T p a 3 a H I { M a e T Buerusue K cnugsofi CTocMerUena cnerKa I{JII{ UeHTpanbHo pagngfloxeHa Or4 IIa, BepuII{Ha g2-4Lo (ra6n' 37)' B6mlsu poge, cnefKa oKpynneHa, Bepullrsssrft ypon oKoJIo B3AyTnq rtclCteneHs MoplUI{HKI{' Beput4Hbr na6moqarctcs cna6ue npUBepuI{HH51g C6 115- A1Y" CE Uptl 3HarIeHI{eIUU qacTr{ pocl'pa onpeaeJuleTce cneanr,Beolspuofi Ta6nu:ua 37 Sachs I4euepeuurr pocrpo a P seudod.icoelite s hibolitoides Ne 85-6' Ana6apcxufi garn{n
Ilapavetput
Ne 85-7' Ana6apcxuft 3arMB
1 1 O , O( 1 5 2 8 ) (-tp"anottaraeMa' 1 2 9 , O( r o z s ) Anr{Hal o6tuanl 105,O(1458) 1 2 1 , 8( 1 s 8 2 ) ycra'oBne'Ha' ( 8 5 ) 87,s (.L225) ( rozg) (80) ( ( t + 2 1 ) 1 0 6 2 ) 1 0 9 , 4 ,[.nrua nocneanbBeonsPHofi .racrt{ '7 ( 7,2(1OO) ,7 1OO ) Anar.aetp cmasso-6puuxoft y BepuI{HbI aJrbBeoJrbl 6,2(86) 616(86) h,"ur"tp 6oxosofi Y BepurHbI
anIrBeOJIbI
Paauyc 6ptoutHoft Y BepuuHbI anbBeonbl [nuHa npl{Bepurxuoft qacrl{ Auavetp cmauso-6prcurnofi B Mecre uau6ontureno yToJlrueHU'I Auaverp 6orosofi B Mecr€ Hal{6onbueno YToJIIueHua Vron arlbBeonsPurfi n cnuHno-6prcuHofi finocKocr]I' rpaA, Vron BepIuHHHblfis 6oroeoft IrrrocKocrl{, rpaA. L
Hiboliroides (nat).
102
2 , 6( 3 6 ) 3 1 , 8 ( 4 1 3 )( 2 9 ) 1O,3(rga)(1oo) 9,4(t22) (91)
32,7 (454) (37) 8,5(118)(1OO) 7 , 8 ( 1 0 8 ) ( O Z1
22
35
- noao6Hufi Hibolites
32
noKasaTerlsx na B rlpeAenax 1060-1270. cmrnHas 6opo3Aa ys*a e u rrrykoKas B nepeA'eft tracr' pocrpa, AaJIee pacu,I{psercs, BbruoJra)KrBaeTcs gesaer, He loxoAs u 6rrcrpo rcu Ao cepeArxrr lla. nonoxenre 6oposabr Ha cnunuofi cropDlre ycra'aBnr{BAercs ''o Haxo)KAeHr{ro Ha npoTDrBOnoJro)KHOii cropoue cr@Ha ua Moxo'e pocrpa )e g7-195 $pan_ (ra6n. XI*,orr.-s). g noio"*" pocTpa cffiH_ Has cropona BbrrrJrxnas. Eprcruuan cropoga BbrrryKnag""or"ft no ecefi AJrl.rHepocTpa, BKJTIG. qas eno anbBeonsp')'o sacTb' Eoxoeuie cna6o y[noue'br, no Hr{M npoxc lgr napanneJlbHble nap'ble nonocbt' "toffi, cnenKa pacul{pff,ou,necs B rar6onee urpennoft qacrl{ pocrpa pacv I'cqesapque ,,pr K Bepur{He. rlonepeqHoe Cer{eHUe OKp)frrreHHO-IIpgMOyfOnbHOenpu6nuxerru B nepeAnefi rraCTI{ pOcTpA, Toe c 6oroe unu pe)Ke CX&* B cnunno-6ppur'oM HanpaBne'r{I{, cTaHoBrITcs oKpyfiJrbrM rlnu cna60 cxa?brM C 60xoe or"nr*"B 3AAHefi qacT|t pocrpa. orBercTBeHHo sHaqeHas co_ EE Bapbl{pyror B npeAenax g 6-1 10, sHatrexfis 66 _ 91-96. BnyTpeHHue npI{sHaKn n oHToneHeg. Anageona npsMas,garlltMaer oKono 15-20% aru{Hbr pocTpa, Bepunra ee Hq3xaqllTenbro cMerrleHa x 6prournofi cropoHe (6prcrunofi pa[xyc sg-46yo cE). air..orrrpHsrfi ynon Haxo'r.rrcs B nps leJlax 2o-22o' cnafixa xa6rnoaaercs ron.xo Ha curHrofi cropo'e e o6aacrz anbBeoJlbr.t{ IlpocnexuBaeTcs B I
il:":'#J#*r":ffi:;:,.":o:mo :ffi ::x';:",*nff J*;"'"ff nprcE4 rurv naaoctunaer ;;.'i"""'?'i: ?HH#"t."ffi:fffi":::#lt?;
6n,nsrzx x 4 vv Al{aMerpax Haqrr'aer -gg):---"' yxe [posBJrsracc perenoBuAnocTb a fiopr"re pocrpa (pac.
Ne 87-1 94, P.Eyop-3e1rT
cna6o Bbrpa)KesHas Bs,
]t 8 7 _ 1 9 5, p.Kenuvep
1 O 5 , O( 1 9 0 9 )
1 O 2 , o( 1 8 5 4) 9 2 , o ( r o z s )( 1 1 7 9 ) (88) s , s (1 o o ) 5,8 ( 105 )
LO7,9(1798) (tz tz) (L272) 1O3,O 6 , o( 1 o o ) 6 , 6( 1 1 0 ) 2,5(42)
2s,2rr]:. ) (s2)
7 , 8 ( L 4 2 () r o o l 7,3 (133) (sq)
38
3 I , o ( o s o () s e ) 8 , 1 ( 1 3 5) ( . r o o ) 7,8(130)(ea)
34
r03
ugMeHtrnBocrl{ crenenb 6oxonoro I{Jrr{crIuHHo-HsveHqHBocrr.Iloaaep)Kelra qacrlr. Pocrpu us Ana6apcKoro pafiapocrpoB B ux nepeatrefi cxarr{q 6prcuHoro 6oxes-"€e+P5t ra g 6omurncrBe HanpaBneurrfi, uMelor a 6onee c)Karbr B cnrnuo-6prounov An}Iua cnnnnofi 6opoensr, X Hexourr{poKyro cn}rnrryro 6opoeny. Baprnpyet TopbD( pocTpoB OHa Aoxo1ttT [o CepeAuIIbI pocTpa, y EeKoTOpsx 6onee KOpcrrKas. Mensercs u o6qac Qopva pocrpoB c/T orqernrrBo BeperenognAnoft ao noqru cy6uunuuApu.recxofi. flc $opve pocrpa Ps.hibolitoides 6nuxe Bcero K Ps. CpaBHeHue. Orrnrqus yKasanbr npu onrrcaHuu nocneAHero. Pocrprt platyaentriosus. poaa HiboliP s.hi bolitoides oqeub HarroMlrHaroTpocrpbr npe[craerrenefi qacTu a pocTpa He CrrIIHEaa y 6prcunas, nepeAueft nOCneAHI{x B Eo .tes, cnfun Spanvoxona Haxonrrcc Y Hibolites Ha 6opoela. CooteercrnenHo tOfi xe CTOpOHe,qTO u 6opOega, B sTy x'e crOpogy CMeIqeHA OCeBAs truHHg.
O a u u a n b H a c r p E y p o r r e H H o c r b . E o n b u J l n r c r B op o c r p o B P s . h i b o l i t o i fles co6panbr B nnr{Hncrbrx $auugx orKpbrroro Mopc a fleno-OneueKcKoM pafioxe. Pexe sror BuA Bcrpeqaercs B npu6pexnux MerIKoBoaI{bx $aulUlx pafioxa r{ neao6epexsg llenrt. EAuHlrrHbIe pocrpbl Ana6apo-Xarairrcxoro HafiAenbr a @Unnx oTKpblTorp Mopc a Bepxornbe E a Oxorcxov pafione.
P u c. 3 9. flpoaoJrbuoe ceqcHue pocrpa P s e u d . o d i c o e l i t e sh i b o l i t o i d e s S a c h s Ne 85-7, aaneH, AHa6apcuft eaaua ( x o , 9)
cneayer cqr{Tarb, .rio Ps.h ibolitoides Ben ceo6oqHo[naBaroluufi o6pas xI{sHIr' npeanoql{Tan o6ctanoeKu orKpbrrorp Mopg, Ho galrnbrBarl r{ B npn6pexubre yqacrKl{ Mopg' H u x x u f ia a n e H pacrlpocrpaHeHue. u reonpasuqecKoe Bospacr Cu6upu. Huxnuft 6aftoc Cenepnoi.i 62 poclpa Hs oalr€Ha-Hl{xuero 6afioca per KemlMtrp u EyopMarepr{aJr. Oerur - c6opur T.H. HanunRenofi, T.V., KupuHofi, C.B. MenearHoft u P.A. Eun)Kr{eBa, 10 pocrpoB r{s a€Jr€Ha-Hlrxnero 6afioca neeo6epexss llesrr (pexu Me ToprryHa, C16nnrcae) n 8 pocrpoB I{s aaneHa npaao6epexbtt llesrt (pexu Eenua)KaH H llenucxe) - c6opr,r T.H. KuprHoiir 8 pocrpoB ns aaJIeHa-HHx(Hero6afioca Ana6apcxoft ry6ur - c6opsr T.H. HanrnReaoft, 1 pocrp us ltlrxneno aaneua p.Bnnunn (Oxorcxoe no6epexse) - c6opu A.C. ,0.anuca. P s e u d o d . i c o e l i t e sc l a a a t o i d e sS a c h s ,
Ta6n. Xvlll,
"p.
nou.l
Sun,"1-4
Mysefi Hlf CO AH CCCP, Honocu6upcx. Pexa EyopNr 87-197. fonornn 3exur (Hueoebs p.JIeHr,r), Hllxuufi aaJIeH. A u a r H o s. Poc.rp 6ynanoeuauurft, cl{nbHo yAnI{HeHHttfi, cxarufi c 6oxon, pe3B nepeaHeft qacrn, co cnI,IHHoft 6oposaoft, uayueft or nepeAF yroHbqarcuruftcg Ko 'Hero KoHua 6onee qeM Ha nonoBrlHy anrnbr pocrpa. B npuaepuurnofi qacru poctpa lr Ha ero 6prcurHoii cropoHe 6opocn nnu ynnoureuufi uet. n p u 3 H a K H . f l p u a e p u r l { H H aqr ra c r b p o c r p a o a H l { M a e r o K o J I oO r 2 Bueu;Hue Or3 l-la, BepuuHa pacnono)xeHa ueHTpanbHo ltntl cnerKa cMeurera K cnungoft croyron HaxoauTcg B npeaenax 35-510(ta6n.38 poHe, cJIenKa oKpyrneua, Bepu:ltHHbrft
L
clorntoide's (nat.)
104
- noao6Hrrft npynne t-lattati
(6ynanoeuanurfi)'
T a 6 n a u a 38 Hsvepeuua pocrpoB Pseudodicoelites clauatoides Sachsrsp.
nov.
-,T.
r
F->
rDF--E___
JlJ-b 87- Lg7, P.Eyop-3eK1aT
llapaverpu
Anuna ycTauoBneHras A,runa nocJreaJrbaeonnpuofi
1 1 8 , O( 3 2 7 8 ) 1OO,O(3333) 1 1 3 , O ( S r s 9 )( . 1 0 4 6 ) 9 6 , 0 ( g z o o()r o s s )
qacTI{
,fluar,rer.pcnnrso-6prorunofi y BepUII{HbI
AJrrrBeOflbt
Auarurer.p 6oxoaofi y Bepur{_ Hbr anbBeonbr Paaryc 6proruHofi y Bepur_ HbI
lr 87_198, p.Eyop-3eKr{T
3 , 6 ( 1 o o)
3 , o( 1 0 0 )
3,s(97)
2,8(93)
7,6(44)
AJIbBEOJIbI
.0nyHa npr{Bepuasnori qacrn Auaverp sm{Hno-Sprcuuoft e MecTe uar6onsuero yroJl. ueHI{'I AHaverp 6oxoeoft B Mecre trau6onuuero yTonueHnrr Vnon Bepur.rHubrfis 6oxoeofi nnocKOCTH,rpan.
2 4 , 8 ( 6 8 9 )( z z l 1 0 , 8( 3 O O () r o o )
18,3(oro)(rg)
e , 7( 2 7 0 ) ( 9 0 )
8 , 7( 2 9 0 )( e o )
9,1(3O3)(1OO)
51
Oronqaune ta6nuurr
Ntu87_200, p.Eyop-OeKHT
AnuHa ycraHoBnenHas Anusa nocneaJrbeeonapuoli qacTu Auave:.p cnuHHo-6proruHoli y B,epuLIHbraJlbBeonbl luaverp 6oroaofi y Bepuu_ HbI
AJIbBEOJIbI
Paquyc HbI
6proruHofi y Bepu?t_
AJIbBEOJIbI
lnlura npr.rBepuusuoli qacrlr fluarr.rerp cnuHuo-6prcuHoli n Mecre Hau6oasrueno yron_
73,7( 1992) -73,7 (1992) (egA)
N, 87-1 99, p.Kennvnp
L O 4 , 6 ( 3 4 8 7)
9 9 , 6 (e s e o )( r o a s 1
3 , 7( 1 0 0 ) 1
3 , o( 1 o o )
4,O(1Oe,;1
3,1(103)
1 , 3( g s ) 1 4{
2 2 , O( s e s ) ( s o 1 8 , 8 ( ' 2 3 e )( 1 O o )
2 3 , 2 ( tt s ) ( z s ) 9 , 3( 3 1 0 ) ( 1 0 0 )
TUEHI,Ig
luarnrerp 6oxoeoli B Nrecre Hau6oru,rueno yronrueHurr Ynon BepuJrlHHbrria 6oxosoti nnocKOCTH,PpaA.
7,6('295)(aoy
8 , 3 (2 7 7 ) ( 8 9 )
35
1y "o*pnHr.rBule.ocrr
nepe^Her-o xoHua pocTpa
r05
CE e 3 u 6onsrue pasa npl{ onaqec6 npeaocxoasr suaqexus cnnbuo Bsayr, I{Aer or lrep€fiCrnurHas 6oposaa ysKas u rny6oxanr na oKono 1OOO-11OO. aJlbBeonbl u y gaAHeno oKoHqaHlrn' 3aHlrMaer B o6nacru Heno KoHIllBr pacu,upqercg EprorucuuHHag cropoHa BblnyKnag. oKorlo nonoBr{Hbr f]a. B saaHefi rlacrl{ pocrpa Bo3F BblnyKnocru crelleHb pocrpa nputleM BbrnyKJras, rro aceft AJrIrHe Harr cropoHa cropoubt ynnoueHbL napaJlJlenbuble [apHble pacraer aJTbBeonbr. Eoroerre e o6flacru Pocrp Hltsx
B npI{BepuI{Hnonocbr 3aMeTHbr Ju{rxb Ha etrHHl{rrHbrx. pOCTpaX, oHLI paculupgloTcs ceqeHlle oKp}rneHno-npnMoflonepevuoe r{ sareM Hofi qacTg pocrpa ""r."uo{. qac'rtl pocrpa, n o6nacrl{ anEF B rrocneanbBeongpHofi c 6oxoa yr-onbHoe, cxaroe BblfiyKnocrl{ Ir Bospacraxl{Io 6opoeau c Bepul{Hoit na 6prcuuot @pr"ry rpeyroJlbnuKa 3uarleut{g EE 93-108 B [peaenax 86-96' 3uaqeHns 66 xone6nrorcrr cropoge. anseeoiulpHofi cyxeHnofi (neo6xoauMo ae$opvau[fi c BeposrHocrbro crrr{Tarbcq pacujl{peHr.lrc
BeoJrbr 6naroaaps 6prcrunofi
qacrl{
cnl{Hnofi
npuo6peraer
cropoHbr
poctpa).
Anteeona He6onbuas, 3oHt{u oHTor'eHes. pocrpa, BepHI{Ha cMeluena x 6ptotu1 Or veHee aJuIHbI Maer npearronoxnrenbHo ynon 3aMepI{Tb (6proulrofi Anrseongpusrft CE). paArryc Or45 oKono cropoue noft B croBblcrynaer Cnafixa Ha6rnoAaercrr Ha cnl{HHofi cropoHe pocrpa' He yganocs. npI{3HaKI{
BuyrpeHHne
poHy nocneanbBeoJulpHoft eno qacrld. Oceeaq nI{HlIn noqrl{ npnMas, He3Haql4Tenbpocrpbl cy6uurunrApl{I-{a Ha.ranbHblx craannx K 6prcuruofi cropoHe. Ho cMeueHa oKono 2OOO' HaqaHaer fla cocraBJlaer qecKge, npu 3garrenl,rex CE 215-3 vv pocrpbl pocrpa. Ha e'rux craal{sx n {opve yxe flposBJurrbcrr BepereHoB]rllHocl'b E y n a e o n u a H y l o p o c r p o B o r P s . h i b o l i t o i d e s . Qopnty Heorrrr{qHMbr Ps,claaatoides 6nilsxux K pa3Mepalvl BspocJlblx ocopocrp npuo6pe'raeT TonbKo npu pa3Mepax, 6eft (puc.
40), Bce pocrpbr B u3yqenHofi xonneKul{n B3nrbl B oauo}vl HsveHqlrBocrr. ypoBHe I{ B lI€ge6oJtburorvr paiioHe, Ha oaHotvt crparunpa$r'recxoM cpaBHHTengHo noM aoBonbHo oAHorlrnHbr. I4sN{eHeHus B sHaqeHuRx rla, [o, EF, 66 u c6 e o6HeBeruKll.
ueM
M lit fit Irl
onuCpegu lpy-nux BllAoB poAa Pseudodicoelitcs CpaBHeHr.le. CblBaeMbrfi eUA BbIAenneTCs no 6ynaeoeu$oft QopMe pocTpa' cI'IJIbHoN,fy yToHbueHHIo pgcTpa B eno nepeauefi rIaCTu u oTCyTcTBuro 6prcurP's.clauatoides c P.s.lrrnpI{sHaK c6nuxaet Hofi 6opoeAbl. flocnennuft Ho y nocneAHeno bolitoides, peTeHoBI{AHaq.
o6utag
Qopva
pocTpa
JTHtub cJIenKa Be-
Ps.clauatoides PoCrpu Oa Ur{aJrbHarr npr4ypor{eHHocTr,, Mops e neoTKpblToro B InI{HucTrx I{cKJIrcq?ITeJIbHo Hafi[esu SaUusx oTCyTCTByIoT oHI{ B MenKoBof paftoHe. CoeepueHHg no{neueKgKoM a Cr.r6upcroft nnar$poMbl' Hbrx r{ npu6pexHsrx Sauunx Ha repufupura MaccI,IBa. OqeeuAHo )KI{BorHoe BeJIo OMonoHcKono Mopg I{ He orKpblTono o6pas xlr3Hlr B ycnoBl{nx ceo6c4luonnasarcuuft e npu6pe)KrryIo 3orry. saxoruno
TaKx(e e o6nacru
40. flpoaoJrbHoe ceqeHue pocrpa P.seudctdicocIites
Boapacr Ceeepuofi
u
neol. pa+uqecKoe
pacnpocrpaHeHtte.
21 pocrp lr3 Hnlruet.o aaJIeHa peK Eyop-3exr4T MarepHan. c6opsl T.14. HanruReeofi, T.L4. Knpunoii u C.B. MenenuHoir.
106
HuxHuiiaaJleH
Cu6upu. vt Kernrvnp -
(
I
\
P s e u d o d i c o e l i t e s g u s t o m e s o u iS a c h s ,
"p.
nou.l
Ta6n. XIX, $ur. 4-6 Lenobelus sp.: Srn. 16, L7.
fonotun
lycrovecoa,
N' 87-202,
196G, crp. 68, ra6n. 6, Sur-. grg, ra6n. 7,
Myseft Hll CO AH CCCP, Hoeocr6upcx. IOxnufi 6e-
pen AHa6apcKono oaJrllBa, nepxrnft aaJreH. Anar Ho3. Poctp cltnbHo yAnHneHuuft (IIa 7OO-1170), cy6uunnnapn.recxrfi, cnerKa cxaruft c 6oxoa, co crrLIHHoft6opoeaofi, nporcn[Balouefics or uepeAgeno, KoHua Ao nonoBHHbI AJtrtHbIpocrpa. EprcurHafr cropona Bbrrryrnas 6es 6oposa rru ynnouleHutr. BHeluHue n p I { 3 H a K L I . B e p u r t a n H a rqra c r b s a u u M a e r g 6 - 4 1 7 o I I a , B e p u 1 a . . Ha pacnonoxeHa ueHTpanbHo, saocrpeHa, BepuuHgbd yron HaxoAI{Tcg B rrpeAeJrax (ra6n. 39). oxono ocrpl{a ua6moqarorcs npoaonbHbre Nroprur{H1rr.Cnux20-48" Han 60po3Aa y3Kan u nny60Kas, fiporrrnnBaercs Ao cepearrHbr pocrpa, }IIronAa Konqaercs, He AoxoArr Ao Hee. Cnrguag cropoHa e aaaHefi nonoBuHe pocrpa BbF. nyKJras, 6prcurHan cropoHa BbrnyKnas no scef, lJrI{He pocrpa. EorosHe cTopoubr Tox€ BbrrryKnbte, Ho B ueHuruefi cTenenr.r. Ha gnx cna6o BbrlleJurroTcs napanJlenbHbre napHble nonocbl' B npI'IBepuHHHofi qacttl paculllpsrcquecc I{ garyxalou[e. flgF. neperrHoe ceqeHl'Ie HN{eer Ha BceM [porrrxeHr.rn pocrpa $opury oBana, cnenKa cxaF ToFo c 6oxos. 3Ha.reHug EE u 66 no cyuecrBy oarnaxoBbr (OO 91-98, 66 9297 ), BnyrpeHrlue rrpI{gHaKu v oHToreHes. A n s B e o n as a H u M a e r o K o r r o Or3 anrubl pocTpa, BepuuHa ee cnefKa cMeueHa x 6prounofi cropoHe (6prcumofi paAl{yc cocraBnset 42-437. CE). Anuaeonnpnrrft ynon He saMepsJrcs. Cnafira pasBLITa Ha cntlHHofi cropoHe pocrpa, Bbrgaercs B [ocfieaJlbBeoJrflpl{yrc erc tracTl 6onee qeM Ha anl{Hy anbBeonbr (pnc. 4Ll. Ocesag nuHus notrr14 rrpsMas, H€orrarruTeJlbHo clvterlleHa x 6prcuuoft cropoue. Ha Haqanbnbrx craAlrsx pocTpbr cy6UunI{HApI{qecKI{e' oTHoeuT€nbHo 6onee yAnI{HeHHbreqeM y Bgpocnbx }KItBorHbIx (V pocrpa Ns 87-205 npn cE oxono 4,5 vu lla oxono 14oo, upu cE 1219 r.4.a oKono 775). IOsue pocrpbl Ps.gustomesovi Heorrril{Mbr or roHbrJK pocrpog ps.[ibolitoides u Ps.claaatoides, To)Ke rtrruleggbrx 6prourHofi 6opoeau. HaveHquBocrb. I4prepurecs B nauefi KonneKurrug pocrpoa ps. gustomesovi u 2 pocrpa, ueo6pa)KeuHbteB.A. flcror"recobslr"r (1966), o6uapyxtlBaloT 6onruryro ugMenquBocrb B orxoueHr{r{ crelregr{ yAnuHegns (eHaqeHuc [Ia ot 5oo y pocrpa, ueo6paxennoro B.A, I]crovecoBbrM na $un. 8, Ao 1170) u o6ueft fupr,,rsr (or cna6o cy6xonuqecroft IIo cnenKa BepeTeHOeH[HOftC HeKOTOpbTM cyxeHueM B anbBeonspnoft qacru). Mensercg o.';:rocurenbHarr Annga cnunnofi 6opoegsr y HeKoTopbH eKSeMnnspoB oHa KoporKaq u nl,Irub HeSHaquT€JrbHoBbrxoAI{T 3a npeAenbr aJlbBeonapnoft qacrr{, y Apyrux 6onee anHHBac, aocrr{papuas cepe[r{Hbr pocrpa. Bapsup}4or TaKxe e 6oru,u}Ix npeAenax suaqeuus (or 20 Ao 48o). Bepumuuoro yrna ,[onycrnrb, rrTo olrucbrBaeMbte pocTpbr ItpIrHaArre)KaT paSHbrM BItAaM npa orpaHr{qeHxocrtr HMerouefocc MaTepuana Aocraroqubrx ocEosaEf,fi (f gOe ) neo6paxeHxbre uM ABa pocrpa Her. B.A. IlctoMecoB
Puc. 4I. IlpoAoasrroe ceqenue pocrpa ,Pseudodicoelites gustomesovi Sachs, sp. nov. lle 87-205, aepxmrfi aanetr, Ana6apcruft sarus( x or75.l -1Bua
HasBaH B qecrb
BIIepBbre onncaBrxero
eno pocrpbr
B.A.
IlcroMecoBa;
107
Ta6nnua 39 l4svepeuufi pocrpoB P s e t t d o t l ic o e l i t e s g t t . s l o n t e s o i ' l S a c l t . s r s p . n o r ' .
N 87-202,
ilapaverprr
sarmg
N 87-203, p.Eyop-3eKuT
100,0 ( 1000 )
npeanonar€leNfaq
Anuna o6uraq Anuua
AHa6apcxt{fi
ycTaHoBneHHatt nocneanbeeonspuoii
96,7(gOZ) 7 0 , 5 ( 7 O 5 )( t Z ) ( Z r )
77,5(775) 7 7 , 3 ( 7 73 )
10,o ( 100)
1 O , O (1 O O )
qacTll
,[.uaverp y
BepuLlHbI
c nrnno-6pnuruoii anbBeoJIIJl
tuItHbI
9,1(91)
9,3(93)
Auarrerp 6oxoeofi y BepanbBeoJlbl
Paanyc 6prcruuof, Anuua rrpllBeputlnHoft qac'ril fluauerp cnHHno-6prcumofi B npr.rBepuuuuofr qacrr Auarurerp 6oroeofi B npuBep[r]tHHofi .{acru Vron BepulrHubtft e 6oxGeoft nnocKocrl,r, rpaa.
28,6 (286 ) (40 ) 9,7(97)(rOOl
4,3(43) 28,6(286)(3O1 10,c(roo)(100)
9,1(91)(94)
s,2(92)(92) 21
48
Oxonqanue ta6nuuu
Ni"87-204' p.Kenuvnp
llapauerper Anuna o6ruac
[-nPennottaraeMas ) \ lycraHoBneHHarr .il.nuua ttocheanoeeonnpuofi qacTu
fiuaverp cnsuno-6pnruuofi y BepuuHbr anbBeoJrbr fiuaverp 6oxoeofi y Beplul{Hbl anbBeonbr Paqryc 6pgruuofi nnuua npuBepufinnofi qacrl{ fiuaverp cnunno-6prcuruofi B flpl{Bepuunnoft qacr}t fiuaverp 6oxosofi B npr{Bepurl{HHofi .racru
68,9( 1170) 68,9(117O)
craBngercrr,
rrro
CIII{TaII Arg
eanlre
95,8 (7 43) >95,8 (743)
5,9(1OO)
12,9(100)
5,8(96)
L 2 , O( g S )
2,5(42)
5,5(43 ) 34,OQA+\ (<SS1 L2,8(gg)(1OO)
2 8 , s ( + e s( < ) r) 5,9(1OO)(1OO) 5,7(97)
12,O(SS1(ea1
20
Vnon BepulluHblfi e 6o10sofi nnocKocrl{, fpaA.
npeAnoJIoxI{TeJlbHO
N r 87-205, AHa6apcxufi
oTHOCtTUIEMUC g
o6ocHoeaHlrg
raKol-o
[PeAHau K AByM pasHbIM BuaM. MHeHI{g Hy)KHO I{N{eTb 3HaqI{TenbHO
6onsruufi MarepuarL Pocrprt Ps.gustomesouipesKo orJll{qarorcrr or Bcex paHee oilllCpaeHeHue. caussx BlrAoB poaa Pseudodicoelites ceoeft cy6unnuHapur{ecKoit unu 6nusxofi x cy6uunrnApuqecKoft $opr'roft. Pocrpu onl{cbrBaervroroHI{xe nuta Ps.primoris sp. nov. TaKXe cy6uunrnapilqecKoft @pvu, ornl{qarorcrr or pocrpoB Ps.gustomesoai HaJu{rrneNt3HaqllTenbHoro cuuuno-6oKoBoro cx(aTl{s.
r08
-:
QaullaJILHas
npI4ypoqeHHocrb. H s u e e e c r H F , r x1 0 p o c , r p o a p s . . g 1 1 .sl.trne[oJroBr.rna co6paHa B MeJrKoBoAHbrx npu6pexHbrx necqaHo_r_Jrl{Hr.rcTbrx a AHa6apcKoM paftoHe n 0aurrflx Ha JreBbrx rpl,rroxax ,enrr, noJroBrrHa _
soui
u'sx or*pbrroro Mopqe teHc-one'eKcKon, ouoo"" , u e"p-"n1.:"ii#:T.:T
@uuernrr oTKpbIToI'o,tvlopq cBq3aHbr 6onee NreJrKr,re sKgeN{nJlrpu. MoxHo npeanoJranarb' qro xIlBoTHoe Beno ceo6oa'onnaBa,ou,rr oop"3 x'3'r.r KaK B o6cr.aHoaxax orKpblTol'o Mops (Nronoaue oco6H)' TaK, npnopexHbrx qacrrrx MopcKoro 6acceftxa (6onee Bspocnrrre oco6u). " u neof pasuvecKoe pacnpocrpaHeHHe. AaneucenepHofi a"j;;pacr Marepr{aJr. 2 pocrpa r{3 BepxHeno aaJreua Aua6apc*ofo sarrr{Ba_ c6oprl T.U. Hansrgesoft, 4 pocrpa ao Hr,rxuero aaJrena peK Kenrvep u Eyop_OBXhr _ c6oprr T'H' HansHgeeofi u T.H, KrpuHori, I t;;rp H3 Hrrx'er-o aareHa p.crcH_ rroae (neno6epe.xue ,euu) n l pocrp u3 Hr{)KHefo aane'a p.Jleuucxe (npaao6epexse p..l'lensr) - c6opr,r T.H,'KzpuHofi. pseudodicoelites
printoris Sachs, "p.
nou.l
Ta6n. XVI, 6zn. 4_s foaotan
Nr 87-207. Myeefi Hfr co AH cccp.
Hoeocu6rpcx. llorryocrpoe Yprcrr-Tyvyc, uuxlrufi Toap, soHa Dactylioceras commune. AuanHos' Pocrp ge6onsuofircnnuuo yanr.tHelrHr,irl, cnerKa BepereHoeaaHnrfi, Ilortrrl cy6uunauApuvecrnfi, B nepea'efi qacru cHJrbHo cxar-o'i B cnl{HHo-6prourHov ' HanpaBneHHI{'B .a'Eeft nonoBl{He c 6nusxrM K oKpyrnoMy no[epeqHbrM ceqeHueM. Hveercg c'',nHlras 6opoeaa B nepeaHefi qacro, pocTpa. Eprcu,Haa c?.opo'a Jrr{ue'a 6opoearr. BxeIxHI{e npI{sHa KI't. l-Iprnepul{HHas qacrb yAJrr{HeHHafl, cocraBnser HeMHorlM Me'ee uonoannu fla (+g-qqyi. Beprur.r'a saocrpe'a, 3aH'Nraer panbHoe roJroxeHr{e, Bepur{HHbrfi ueHTr ynon paBeu 2Lg2o (ra6n 40). Cna6o Bepere_. HOBr{AHassoprrla pocrpa o[pereJrsercrr reM, qro c6 cocTaBnser 102_1197. c6 npr na 76+930' cnn'nas 6opooaa ysKafl N rny6oxas B o6nacrn anbBeoJrbrl 3oxoAI{T B TlocJleanbBeonspHylo rracrb pocrpa Ha 7/ I amltsr IIa. Cnugxas cropo'a ga IIpeAenaMI{ ee pasBr{Ttrs B cpeauefi qacrlr pocrpa ynJrorueHa, crauoBl.trcs Bbr_ nyxaofi B [p[Beputrunofi ero qacrl{. Ep'uHaa cropoHa BbrnyKnas, cJrer*a ynnGueua B anbBeoaspHoft qacr,n pocTpa. Eorogrre cTopo'br Bbrrryr(Jrbre, gerrtrfBo ButrIfbI [apHble Ha HHx oT_ noJlocbl' uAyu[e napatnenbHo apyn Apyry Ha npoTrr]KeHr{a 6onsrueft 'racta linrnri po3rpa, cJrenxa pacxoatcb B npr,rBepul4gnori ero qacTr{ ' SaTyxas npr npl6J'II{x(eHI{H K ocrpl{p. rlonepetr'oe cegeHue I{NreeT B saru{eft rracTt pocrpa oKpynJrlm 00pvy, B nepe'Heft nonoarrHe pocTpa npuo6peraeT oBaJIa, c)t(aToro_ a crmlrHo-6pbuJHoM dOpvy HallpaBneultu. coorr"r"r""HHo 3gaqesus EE tL3-L24, 66 g7_1O9. BnyrpeHHEe ,,p,ngHaKIt u oHToreHeg. A n s B e o J I ag a ' I { r \ , r a e ro K o J r o Or2 nnnubl pocTpa, npsMas, aJrbBeoJrspnrrfi w.on DaBeIr 24o. Beprunsa cnenKa cMelueHa x 6prcuruofi cropone (6pourxofi p"ory-" y pocTpa Ne g7-2 Lo 49o6 cE). cnafira Bblpaxe'a cna6o co c,,r'Hofi cropo'ri s6nuer{ Bepur{Hbr aJrbBeonsr. oceBag Jrtflrl{g [otITIt upsMas' cnenxa ll3ol'Hyra H cMeuleua x 6prorrxori cropo'e. Tone'eg o[[cblBaeMoro BI{!a OHIr3-'34 HeAocraToq'ocru MaTepl{aJra He npocnexeH. LlauegtrnBocrb. paHseroapcKae pocTpbr c [_oBa Vpronr_Tyuyc poTl([e' qeM pa[HeaaJle'cxl{fi 6oaee Ko_ poctp c p.Kenrrvgp (nu 7go-gso n 1111), upreprj?p" Orursxpo r cy6uuanrAprrqecKofi"ootr"r"t"""ro (c6 cooteercTBeu_ topr*y Ho 102-116 u 119), 6onee *opor*1* cnr{HgJrro 6oposay-("oo"u.rcraeuuo n Or5 ArIIfIrbI pocrpa), Menee Or3 cxaTbr s cnmro_dprcrJgoM-HarpaBneglrg B npuBep_ L
Pr;^or;,
(aar. ) - neparrfi, nepeaoeofi.
t09
Ta6nuua
40
Hsuepeuurr pocrpoB Pseudodicoelites primoris Sachs, sp. nov.
]s g7-2O7, rr-oB VprcHr'-Tyvyc
( lTrpeanolnaraevac Aru{Ha{ o6uagl lycraHoBlnexnas Anr,rga nocneanbBeonspHoft .rac-
I',{e87-2O8' rr-oB Yprcnr-Tyr'ayc
5 4 , s( 9 9 1 ) 5 4 , s( 9 9 1 )
)s g7-2O9, p.Keruruap 5 1 , O( t + t Z l
50,4 ( 925 )
4 2 , 8 ( 7 2A ) ( ZA + 1 4 6 , 9 ( 8 s 3 ) ( Z S S 1 ( 7e )
48,4(L344)
4O,O(rrr1)(eso)(ze
TIt
Auarurerp cIruHHG6puuxoft y BeptxHHbt aJrbBeoflbr Auar.aerp 6oroaoii y Bepulrubr anbBeoJtbr Anuna npuBepurrsgofi qacrn Anaverp cnunno6pronrHoft B Mecre uau6oJrbr.uero yTomlleHI{tr Auaverp 6oxoeoft a Mecre H6I{6onruero yroth
5 , 5 ( l O O) 6 , 2 ( 1 1 3) 1 8 , 6 ( 3 3 8 )( + S )
6,8( L24)
20,7(376) (441
3 , 6 ( l O O)
4,L(LL4,
L 7, 5( + e O\ G + l
5,6(102)(rOO) 6,4(116)(100)
4,3(119)(1OO)
s , 6 ( 1 O O () O e 1 6 , 2 ( 1 1 3 ) ( O Z 1
4 , 7( 1 3 1 )( r O O1
luEHId'I
Yron aepuruHHrrfi e 6oxoaofi
5,5 ( 1oo )
23
32
2L
IIJIOCKOCTlI, npaA.
uurnroft qacrl{ (66 coorsercrBeHHo 97-98 u 1Og). Iloerovy o6teauuenue Bcex oTI{x pocrpoB B oaI{H BI{a sBnrrercrl npeanonoxl{TenbHblM. He ucxruoqeHo, lITo pocrp Nb 87-209 c p.Kermvnp [peacraBJr.ser rcHrrft eKseNrnJurpap]rnoro BuAa P seudodicoelites. Cpa eHeHH e. OnncuBael,rbre pocrpbr rro Hanl{qurc ctlnbuoro cnuruo-6prcurHono c)Karuq pesKo oTru{qarorcrr or Bcex ap}'nux BI{AoB poaa Pseud,odicoelites I{ B Torvr trI,IcJIeoT aaneHcKoro Ps.gustomesoui, I{Meloueno roxe 6rueKl,rfi r cy6urnuutr puqecKoMy pocrp, Ho cx(arbrft c 6oxoe. Her ocHoeaHnfi paccMarplBarb ux n KaK Monoab KaKoro-nu6o apynoro BIrAa Pseudodicoelites, repr 6onee qro B paHHeM Toape Apyrux BI{AoB Pseudodicoelites Mbr I{ He 3HaeM. Onncr,rBaeNrbre pocrpbr c6nuxarctcn no caoeft @pvre c Lenobelus ailigaensis. Pocrpu nocneaHero raK)Ke cxarbl B cnHHHo-6prcuruoM HarrpaBneHI{I{, Ho, KaK Caoftcrge}rHo BceNry poay Lenobelus, r{Meror Haqr{Harouyrccq or Bepurxsr 6prcuru1xr 6opoeay. OauuaJrbHarr nptlypoqeHHocrb.3 pocrpa P's"primoris saft[esu B MeJrKoBoaHblx npu6pexHbrx necqaHo-I-JrlrHlrcrbrx @unax Ha n-oBe Vpur-TyMyc H rrutxl) 1 pocrp B nnuHltcrux @uunx orKpbrror-o Nrops B HH3oBbgx Oneuexa. OrcrcAa Mo)t{F Ho npeAnonararb, rrro onucbrsaevrlft Bt{a npeanorruTan MenKoBoagrre o6ctaBoBKI{, Hor cyas no yaru{HeHHocrH pocrpoB, aen cao6oAHorrJraBapurfi o6pas }Kr{3HI{tt g€rnJIr,rBaJILr B orKpbrroe Mope. BoevoxHo, rrro B paHHeM Toape Ps.primoris o6nraa
1t0
B npu6pe)KHoft soue Mops rr nuulb lrosxe nepecenuncs B o6nacrb orKpbrTOrA MOpg. Bospacr u reonpa+nqecKoe p a c n p o c r p a H e H H e . H u x u u f ir o a p (soua Dactylioceras commune) - Huxarafi aaneu Cenepuofi Cu6upu, Marepuan. S pocrpa r,rs Hr{x(Hero Toapa tr-oBa yprcur-Tyruryc, l pocr.p ne Huxgero aaneHa p.Kenrvnp - c6opbr T,V. HanuHqesoft.
P s eu d o d i c o el i t e s s p . B ocHnu nopoa Bepxnero 6ara Ha BocrorrHov no6epexse Ana6apcxofi r-y6rr BMecTe c xapaKTepHbIMHAns 6ara pocrpaMl{ Pararnegateuthis T.Z. HansHgeeofi 6unr saftAeHsr 2 Henonublx pocrpa, rplrHaAnexaruue 6eccnopxo Pseudodicoelitinae u cKopee Bcero poAy Pseudodicoelites (poc,rprr NNe B7-211 u 87-212), Hexoropag HeyBepeuHocrb B poAoBofi npuHaanexHocrr{ pocrpoB Bbr3brBaerce reM qro oTcyrcrByeT npl'IBepuuHHarrqacrb t4 coorBercrBeHHo Henb3rr Karenopl4qecKr yrBepxAarb, trro y BepuI{HbI He 6rtno csoftctseHuoft Lenobelus 6puuHoft 6oposasL Y poctpoB He coxpaHllnucb KaI( npr{BepulrHHarr, TaK H anbBeoJurpHafrqacru. @opva pocrpoB cy6urnuHnpuqecKarr c HeKoropbrM cylKeHneNrB nepeAgef, ux qacru. IlonepevHoe cer{eHl{e oBanbHoe. B nepeaHefi qacrr.r pocrpoB Ha6.rnoaaercn rny6oKas cnraHan 6oposAa, [ono)KeHl{e Koropoft yctauaBnr.rBaercs ilo cNrerueg1ro x 6prcr:rxofi ctopoHe oceBoir nunuu H no ga6ruoqaeMoMy B rlpoAonbnolvr cer{eHgr.r nofloxeHuIo BepuI{HbI oaHofi ue HaqaJlbHbtxcraaufi pocrpa. Eproruuan cropona BbrnyKnarr, Ha Heft He BI{aHo HItKaKI{x cneAoB 6opoeasr unu ynnoueHuq. ,0,nuHa coxpaHuBurefterr qacru 6onee Kpym{ono pocTpa N" 87-211-110 vu, CE (a6nusu Bepur{Hbr anbBeo_ n r r ) - 1 2 , 5 M M , E E - 1 1 , 1 r , r v ( 8 g ) , c O ( n v e c f e H a n 6 o n u r u e r op a c u . i u o e u u a ) _ 1 5 , O v v ( 1 2 O ) ( f O O ; , 6 6 - 1 3 , 2 v v ( 1 0 6 ) ( g g ) f l o c n e a l r b B e o n e p H a r rq a c T b pocrpa npearono)Kr.rrenrrHo cocraBnrrna oKoJro 170 vv (rgoo) ( rrss). Eprculrrof, panl{yc a6nueu Bepur{HbranbBeonbl paneu 5,3 MNI (42). OrMctlsaeNlble pocrpbr Henb3tr orHecru Hu K oAHoMy I{o l.rgBecrHbrx BI{aoB pseudodicoelites. Enuxe Bceno no o6ruefi @prvre pocrpa Ps.hibotitoides r{3 a€Jr€HaHI{xHero 6aftoca, xoropttfi oAHaKo uMeer pocrpbr c 6onee ortrernl{Bofi nepereHoBI'IAHocTbIoa MeHee NtaccllBHbte. MoxHo ayMarb, qro pocrpbr npr{HaanexaT HoBolvty eue HeH3BecrHoMyBI{ay Pseudodicoelites, Ao)KLrBueMyao no3AHeno 6ara. llono6Hsre pocrpbl Hl{t.ae 6onee B ornox(eHr.tqx 6ara s CenepHofi Cr,r6rapnHe 6rrnn HaftaeHu. 3ro voxHo npeanonox(uTengHo o6rgcHnTb o6HranueN{ xHBorHbrX B orKpblTOt\l
MOpe.
CIICIEMATI,IKA IOPCK}IX I,I MEJIOBbIX EEJIEMHIITOB
I I
(
r
h
)
I t
t l
Hsy.rerue KoJureKuufi ropcxux rl pargeMenoBblx 6eneuguroB ceBepa Eepaenl{ Mbt aBTopbI Csou Bgrrtsabl no gTolvly Bo[pocy saKaHqnBaeM o6sopolvr I{x cucreMaruKl. nsJraranu pagee (1967a,6, 197O). Hcxons tls npexlrllx npugur{noB B oueHKe npusHaKOB TrpI{ BbTAetteHUUTaKCOHOBpa3JrUqHOrOparra, aBTOpbr npeAJElaror yTOqHeHByro KnaccuQuxaunro, ocxoBanryro Ha vouorpa$uqecxofi o6pa6orxe Bcex rpynn 6eJIeMHIToB, Bcrpeqennbu( Ha ceBepe Cu6upu, a raKlKe Ha norrBuBlill{xcrr ga nocne!HI{e roabr HoBbrx nl{TepaTypHblx aaH}IbDC. flosg5eN.renoBbre 6enevHurbr Ha ceBepe Eopasur BcrpeqarcTcs oqeHb peaKo lr HaMu lte ugyqanucb. Hx cucreMar[Ka [pHBoaI{Tcs Bceueno no truTeparypHblM AaH-
I
( I t
e
HbIM.
MoHorpa0rtrecKoe rrcytreHge 6onuurofi KonneK[I{t{ pocrpon 6opeanbubtx 6enevsuToB npI{BeJIo Hac K BbrBoay, rrro Bce IopcKI{e H Meflogrte 6eneMHI{TbI MoFyr 6uttu o6reauueubr B aBe rpyrmu ceveftcrn (nna naaceMefictaa): uepsas - Passaloteuthidae, Hastitidae, Cylindroteuthidae, Oxyteuthidae; Bropas - Duvaliidae, BelemTaxuv o6paeorr, oduen nopsidae, Dicoelitidae, Dimitobelidae u Belemnitellidae. L842), I'.fl.Kpnrvceveficrea Belemnitidae a roHuMauun a'Op6nruu (dbrbigny, u prrAa lpyrux aBTopoB rfJrr{HaAcerureficrsa Passaloteuthaceae roJrbua (1958) cooteercrByer B.H. Caxca u HansHsesofi (1967a,6) AByl"r xaacelreftcrBaNr:Cylindroteuthaceae n Duvaliaceae (noapo6xee cM. Caxc u HanrnReaa, 197O). Eneqxxft (Jeletzky, 1965, 1966; Westermann, 1973)' HaseaHxbre HaAceureficrBa paccrv,arpuBaer KaK noaorpsabr Belemnitina (= Cylindroteuthaceae) u Belemnopseina (=Duvaliaceae). Ograro raKoe peueuue rpe6yer nepeBo[a o6urenpaHsroro noaorpsaa Belemnoidea (Belemnitina) B pam orpsla u coorBercrBeHHo [epecMorpa scefi cucreMarfiKu KnaccaCephalopoda rrrm no rpafixefi Mepe noAKnacca Dibranchiata ('Coleoidea ). llogrotuty uenecoo6pa3Hee rpyrrrbr cer"reficre 6eneunuroe o6belMxsrb B HaAceMeficrna, BKJrroqaeNoreB olun noAorpsA Belemnoidea. Heo6xoarMo yql{TbrBarb ro, trro rrpe![onaraeMasr xnaccu0rxauus ocnoBaga Ha IIpl|3HaKax pocTpoB, cocTaBnslout[x qacrb BHyTpeHHeI-o cKetreTa xI{BorHoFo, u noroNry lfMeer ugBecrubrfi gneMeHT ycnoBxocrl{. 197O (Belemnitina Haacerueficrso Cylindroteuthaceae Sachs et Nalnjaeva, Pocrpr,r cy6xoHlruecxl{e, cy6uunHHa1965, 1966; !\estermann, 1973). Jeletzky, plrqecxue, pexe BepereHoBl{aHbre c 6oposaaMI{, Harrl{HarourlrMltcrror Bepurl{Hbr, pexe BoBce Jrl{ueHubre 6opoea. QpanvoroHbr l{Meror KoHuqecK}'ro dopvy, saAHue KaMepbr 3anom{egbl (no ne scerga) KaMepHbrMHorJro)Kenl{rrNrl{olopa-anr. C e r n r e f t c r g oP a s s a l o t e u t h i d a e N a e f . 1 9 2 2 ( B e l e m n i t i d a e d ' O r b i g n y , 1 8 4 5 (pars), Jeletzky, 1965, 1966). Pocrpu nnaArre rrru co cna6o BbrpaxexubrMu npl{BepuuHnbrMn 6opoeaarru, c [apHbrNil{ 6oxogrrvlt nonocal_vru,c6rtrlxabul{N4ncn B rlpuBepuruxofi qacru. lertann - HH3br KeJIJroBes. IloaceNreficrBo Coeloteuthinae Naef, 1922. Pocrpu cy6ronnvecxoft @pvrr, npeacraBnglol[rre Jrutub cpaBHr{Tenbno roxrnft noKpoB, o6nerapuuft {parr'aoKoH, KaK y Sopru Ha Haqaff'HbDc craar{gx pagBl{Tr{s, TaK r{ y Bgpocrrbrx xr{Borrrbrx. Ornocurenbnan anuxbl pocrpa. Cuuerupp - nnrHc6ax. anuHa nocneaJrbBeonspHoftqactu 1/1O-1/15
lt2
c c t
,
I ]
l a E l
v
F
r u
T li
l ri tl
c o e l o t e u t h i sL i s s a i o u s , 1 9 1 2 . T r n p o A a - B e r e m n i t e se x c a a a t * sp h i l lips, 186s-1g71, nuxHro .r".rop Axraru. pocrpsrcy6xoHuqecroft @pvrr c clKarblec 6oxog
ffiiffit:tffi:::'#:o'
t{ru{c or(pvrrrbM nonepeqHbrM ceqe'r{eM.
llonceveficrBo Nannobelinae sachs et Nalnjaeva, 1967. pocrprr c naaaxofi noaepxiocrblo' 6es eaver'blx 6oposa, cy6xoHr.recxofi urur cy6uruunrpnrecrofi MbI' Ha HarrarbHblx cTaAu'x cy6xoH'tlec*ue' 6opoT'oct{Teauxo 6onee yaJluHelr'bre, geM y R3pocnbD( )KI{BOTHbrx. N a n n o b e r u sp a v r o w , 1 9 1 4 ( p r o t o t e u t h r s Lemoine, 1g1s). Tun poAa _ Belemnites acutus Miller, 1g23, A'rmru. Koporxue pocrpbr cy6ro'r_ qecxofi $opvu c aJrbBeonofi, "n""r,of earruprarcutefi oxono [oJroBlrgbr trnuHbr pocrpa uJu 6onee. feTrarl - gurcrHfi aaJreH. C l a s t o t e u t l r i sL a n g , 1 9 2 8 . T u n p o a a - C l a s t o t e u t h i sa b r u p t a Lang, 192g, ur.rxHuri nnnuc6ax Aunnu6 GeHu KopoTKue, cy6xoHuqecroft @p'*r pocTpbr c [pr_ TynneHHOfi eepur'Hofi. Anuseona 3a'r{Maer 60nee nonoBlr*br lJnf'br pocrpa. Ilmnre 6ax - ullxsrufi aaJreH. B r a c h y b e l u sN a e f , L g 2 2 ( A n g e l o t e u t h i s Lang, 192g). Trn poAa _ Belem_ nites brevilormis Voltz, 183o, Toap Bocroqsofi opaxunr. Koporxue pocrpbr cy6ur{nr{H[pr{qecxofi $oprurrr. Anuaeona oaHr{Nraer or 1/B ao i i--o*br pocrpa. flnuue 6ax - 6afioc - ? nuxullfi xeanoaeft. floapoa Brachybelus Naef, Lg22. BepuuHa pocrpa c ,6rursxrM K ue'TpanEr HOlvly nOnOXeHI{eNl, noneperrHOe CeqeHHe OKp},TnOe Hm{ Cna6O Cx(aTOe C 6OKOB. Ilnrxc6ax-6afioc - ? uuxHufi xennosefi. I l o n p o a A r c o b _ e l u sS a c h s , 1 9 6 2 . Tun poAa _ Dactyloteuthisdolosa Voronez, . 1962, Toap ceaepHot cr6npu. pocrpul -6oxoa, cxaTbre c co cMe_ trIeHHblM K cnl{Hgofi cropoHe saanrltM KoHuoM. Toap _ truxnrft aaJreg. lloaceveficTBo Passaloteuthinae Naef. pocrpr, t' giz. Bepere'oBrraxofi, cy6uunuuap[q66kofi unu cy6xouuvecxof "*oo cxaTbre c doxoa, @purr, t.nalxtre, co cna60 3aMer'brMg npnBepur{HHbrMr{ 60poeaavrf, Ha Haqanb'brx cra[[sJ( cy6uuarnr^pl{qecKue, oTHoc]rrerbHo 6onee yAJrItHeHHbre, qeNt y Bgpocnbrx ]KttBorHgrx. Fetr.
TaHI'
-
aaJIeH.
Passaroteuthis Lissajous, 1g1s ( H o r c o t e u t h i ss t o l l e y , 191g). Trn po_ lI8 - Belemnites paxillosus Schlotheim, 1819, nanrc6ax roxsofi tracr[ opr. Pocrpu yMepenHo BbITstHyTble, cna6o BepereHoBrranbre c [pnTyrrnesHoft aepunrof,. Anbeeona cocraBJlseT oT 2/ 5 anwtbr pocrpa I{ Me'ee. rerrann - Ht.x'nfi aaJle[. Catateuthis Nalnjaeva, 196Z_. Tun poo" _ Catateuthis atricaNalnjaeva, 1967' roap ceeepuofi cu6rpu. Poctprt Bbrrs'yTbre cy6unrurxaprrqecrue unu cy6xoHt{qecKse c ocrpofi aepurHofi. ""t"ro Aasaeona cocraBJrrrer L/4 uwt Me'ee anr{Hbr pocrpa. flrg{Hc6ax _ Htrxntlfi aaJreuo Orthobelus Nalnjaeva, 1gZO. Trn poAa _ Orthobelus obscurus Nalnjaeva, 197ot roap ceaepHofi cu6upu Poctpu *o"n*."* ofi unu cy6xonr.recrofi yMepeHHo BbITgHyrbIe' 3aocrpeHHble. 4opr,ru, Eorogrle [orlocbr Bbrpaxegbr cra6o. flon* peqHoe ceqeHl{e cxaro c 6oxon I{Jru oxpynJloe. Bepxu nrnnrc6axS a€lroH. Doctvloteuthis Bayle et Zeiller, 1878. Tan p o A a - B e l e m n i t e si r r e g u l a ris schlotheim' 1813, roap pxcuofi sacr' oPr. pocrprr cy6roxr.recxofi rru cyG u,rrrrH^puqecrofi @pruru c rJrnbrM saaHr{M KOHTIOM,cxarbre c 60*os. HHOnAa a lpl{Bepu,ttrHofi sacTl{ pocrpa pagBltBabrcs 6prcun at n B Me'rruefi crenenu cE rb Ho-6oxoaue 6oposAbr. ll4s11c6lx _ TOap. Pleurobelzs Naef, Lg22. Tnn poAA_ Belemnites compressus Stahl, t824t unrsc6ax pxsofi qacrlr oPf. Pocrprr *p"r."oe,,qsofi Qoprr,rul,crnsno c)r*Tbre c 60ro8, co cna60 Bbrpaxerr'brMn c[r'HO-60nossrvu 60pooAaMr. rrrnryc6ax_!0€p. Castrobelus Naef, Lg22. Tun p o t r a _ B e l e m n i t e sa e n t r o p l a n u s Voltz, 1830' nrunrc6ax Bocroq'oft opa'unn. Pocrprr cy6xoxrrqecroft nru cy6urJu'Apliqecrofi Sopvu, cx€Tbre a cmnuro_6ppu'oM HarrpaBne trnrr, c ynnouennoft 6prcur nofi cropoHoft. floacerraeficrBo Megateuthinae sichs et Nalnjaeva, 1 9167. pocrprr c xoporuo pa3B[TbIMu [pIrBepuIt{HHbIMIt 6oposaarr'rr, cy6turruoraprr.recrofi rru cy6rosrqecxof,
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KoporKHe' cy6roHnvecKl'Ier no Nlepe droprrlr, Ha HaqanL,HbrxcraAl,tflx orHocnT€nbHo o6paayercsl orrnpocrp. lerHepearo B atrMHy. I)ocra xgBorHbrx BbrTqrl{Balorcri KenJIOBefI' Hh3bi CUHerulrcp ? TaHF p o A a - B t ' l e n m i t e s o . \ ) ' ( ' o t r l l .lsl e h l i n A c r o c o e l i t e . sL i s s a j o u s , 1 9 1 5 . T u n .{acru lln}I cy6ul'lruHAput{ecxlle Cy6xoHttvecKue OPf. Z.ieten, 183O, Toap rc)KHofi npHBepu[HHbiMI{ 60Ir cnfiHHo-6oKoBblMI'I 6prcu:Hoii Bbrpa)KerrHoii xopouto pocrpr.,t c - nnHHc6ax - aa'neH. po3AaNt[. lettanr? p o A a - B e l e n t n i t e .ssc a b r o s t r s P h i l P s e t t d o l r a s l i l eN s aef, Ig22. Tnn yanl{HeHHbtepocrprt cna6o Beperq Cuasuo AHrnuu. nrMHc6ax lips, 1865-1871, HoBI4AHoftr{nrl cy6[unl4HApuqecKofr @pvt l, co cnI'IHHo-6oKoBbIMlI I{ 6pxluxoft 6e y BePut{HbI. I]nugc6ax. po3AaNrH ' poS a l p i n g o t e u t h i .[s, i s s a j o u s , 1 9 1 5 ( C , u s p i t e u t h i sA b e l , 1 9 1 6 ) . T u n qi a c r u O P f . r c > r i H o r T o a p 1 8 4 9 , u e n t r i c o s t . s a c r r a r i r r . s Belenntite.s Quenstedt, Aa Pocr.prr ctnbHo Bblrqgyrbte c enfipocrpoM I{ KopoTKofi anueeoJlol"{rHa HaqanbHblx c,1aalrrrx KoporKue cy6uu,runaprdrrecKue uflu cy6xoun'recKue' c 6pnuuofi 6oposnoit y BepuI{Hu. IlrnHc6ax - ToaP. p o A a - B e l e m n i t e sr h e n a n r r sO p p e l , p e s o t e t t t h i sL i s s a j o u s , 1 9 1 5 . T H n aaneH rcxsoft qacrr.r OPf. Poctpsr cy6xoxtlvecroft @p*'ru, c)Karble 1858, Hn)KH14ft Cuc 6oKoa, co cnl,rggo-6oxonsrMr.r6oposAaMll. EprcruHan 6oposna orcyrcrByet. 6afioc. HeMroP p o A a - B e l e n t r t i t e ss p i n a t u s Q u e n s t e d t , H o n t a l o t e t t t h iS s tolley, 1g19. Tun qacrr{ g, pxxofi OPl, Pocrpu cy6uununaprlqecKoft @pvu c ocrpofi aaneH 184 co cna6o BbtpaxeHHbIMI{cnlnHHo-6oxosuvu 6o6oxoa, c aepurnHog, cna6o cxarbte pogaaMlt. Aanen. p o a a - P a r a m e g a t e u t h i si s h n e n Tun p a r a m e g a t e u t h i sG u s t o n e s o v , 1 9 5 6 . fle.ropu. He6ontu-rue cy6xo6accefisa sis Gustomesov, 1960, suxnuft xennoaeft Passurnt L/ 3 pocrpa. oKono cocraBnsrcuefi anbeeonoft, pocrpbr c HlrqecKr{e I{ 6prcurnas. Eafioc - xuxsuft Ke[cnruxo-6oKoBble npr{BeprurrHubre 6opoensr noBefi. poAa - Belemnitesgiganteus 1878. Tun h l e g a t e u t h i sB a y l e e t Z e i l l e r , Schlotheim, 182O, Bepxglrfi aaneH OPl. Poctpsl cy6uunrHApuqecKoir utwr cy6xoxu'recKoii $opvur, c oBaJrbHbrMnonepeqHblM ceqeHueMr cI{JrbHo BbITqHyrbIe' Ha HaqanbHbrx craaurrx KoporKne, cy6xoguqecKue. llpreepuruHHbte 6opoeaut xopolxo pa3BUTbt - 6ptougac n O6llqHO IIBe napbl 6oxoetx. AaneH - 6al. poaa - Belemnitesmunieri Deslongchamps, H o l c o b e l u sS t o l l e y , L g 2 7 , T n n L877, aaneg QpgsuHll. Pocrpur cy6uunun[purrecxoit utw cy6Konilqecxofi @pltu, c 6prcruxofi 6oposnoft, nauuuaroruefics y Bepurubt I{ aocrnrarcuefi anbBeonspnofi qaCTU. V naqana anlrBegnbl I{HOrAa xa6rnOqaetCs Cnaftxa. TOap - 6aftOc' Ceueficrao Hastitidae Naef. 1922 (nom. transl. ex Hastitinae Naef, L922l.. Crnaxo BbrrrHyrbre BepereHoBullnL,re, caa6oaepereuoBuAnbte unu cy6uulluHApnqecKl,le pocrpbr. Ha 6oxosbrx cropouax napHble rlonocbl, pacxoAluuecq a gaaxefi gacrl,t pocrpa u garer\r ucqe3arouue. Cbeeas fitrrltrs saHuMaer uenTpaflbHoe [orloxeHue. flnunc6ax - Aanes - HlDKHlrfi 6afioc. floAcer"reftcreo Hastitinae Naef, L922. Pocrpu cnnbHo BbrrgHyrble, BepereHoBtrluroft unn 6ynaeosuaxofi @prru, He llNterouue 6oxosux 6opoeA, llnrxc6ax - HII)Kp
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R h a b d o b e l u sN a e f, I g 2 2 , T n n p o a L _ B e l e n t , , i t e se x i L s d,Orbigny, Ig42, Toap fuauuut't' cuasHo BbtrslHyTble cy6uunl{H.prqecKlre }rnh crm6o BepereHoB,AHbre Xoporuo pa3Br{rbr 6oxoaue 6opocaur, He Aocrarouue Bepur.rHbi. Toap _ :fi:::t. Parahastites Nalnjae'a, 7967. T r E n 'cr6upu. p o q a - p a r a r t a s l i r e . sn t o r c r t a e t t s i s Nalnjaeva ' Lg67 ' cpeauzfi Toap ceneprori pocrpsr yrviepelrHo Br,rrqH).T,L,e Bepere,oauanofr I{nH naHueroB}IJl'oir Sopvu, cx€Tbre c 6oxoe. Ha 6oxoeo,* Hax xopono saMeTHbI€ aBol'rHrte 6oposau, -"ufrn. B anbBeonspHoii vac "r.oo* oTxoaquue apyn orqr;yr;'n rr,rrl pacurup"""ot "6nr*"""Lre _ Hr.rxHr.rri Toap aaneH. 3aueqaHI{s' B'A'' lycrorutecon (-1.g73), o.rro"o,Baflcb Ha pa3nuvunx Ha pocrpax, npeAnoxun B ceMer?crae 6opoea l{astitija" BbrAerrr.rrbeue aBa noqceveft_ crBa sachsibelinae u Parahastitinae. H"r-np.ocraBflqercfr 'Ho"tii"*"",'iu"""r"rBarcrueM Heuenecoo6pasrrrul yBefiutil{Ba?b qlrcno noaceN.,etic's e ceNtefictu" qeTbrpe poAa, xoTrl Bceno Ha[o [pI{3HaTb, qTo Bce qeTbrpe poAA, cocTaBJlgloulue gTo MeficrBo' cyulecrBeH'o pasnxqalorcs ce_ Mexay co6oft no xapaKreily 6opos, Ha pocrpax. Hatlr'a.ueficJr
pocrprr c 6prcuruofr 6oposaofi, oT Bepur't'I ,n Bbtpaxe'Hofi e pa3n'q'ori creneur. Ha 6oxoebrx cron9{99t1' c6nuxarourecs u Ecqesaroure B npr.rBepulr{nxoftqacru.
::H: floAceveficrao cylindroteuthinae stolley, 1g1g. pocrpul Ha Haqarrb'brx cra'All'Ix pagBETltg cI{JtbHo BblTsHyrbte, cy6urmulplqecKr{e r{Jrl.rBeperexoBlrA'bre, no pocra dc{BoTHbrx oTnocrrrenbno Mepe yTom{arorcs. Eaftoc (?) _ 6ar _ t.orept.lg. cvlindroteuthis Bayle et Z e i i l e r , i g t S l '-Fo"rpo, r-rn poaa B e r e n t n i t e sp u z o s i o n's dorbigny, 1842: o*"oopo tDpaxuun. BbrrsHyrble, c orHoc'Terssoft aruruoft nocneaJrbBeonspHofiqacru or 3OO_SOO qo 2OOO. Aaseeona 3aHuMaer or 2/3 anrHrrr pocrpa I{ Me'bure, BepulrHa anbBeoflbr Jvv'tDr n I{ *,Le,,asl oceBag fiIi*r4s cMeue'br x 6prcurxofi cropoue. Eaftoc (?) _ noTeprBo floapoa cylindroteuthis Bayle et Zeil)er, 182g. pocrpu cr(arbre c 6oxoe, l'ol'epeqHoe ceqe'ue oBaJIbHoe I{JII' 6nlsxoe K HeMy. Ear-xl.rxg{Dft Bana,,xr'. Iloapoa CunmunicobelusGustomesov, 1964. Tun rroapoad _ Cylindroteu_ this (communicobelus) subextensoides Gustomesov, 1964, nnxnrft xennoeefi PseaHcrofi o6n' Pocrpsr c oKpynJIbIM nonepeq'brM ceqe'HeM, c aJrbBeonofi uarcuefi He Meuee L(? osHr{pocrpa. KernoBefi _ ? oxc$opa. --i l*y, floapoa ArctoteutAis S achs N " i n ;'igot, """;; ,,64, Trn noopoAa _ cytind_ roteuhis septentrionaris Bodylevsky, "i rlp*rr* KrMepuA,. Tafivsrpa. pocr_ pbt c)K{rTbi B cltrf'te6prcruHoM HA,,paBre'uu, nonepeqHoe cetle'ne oKp]rnJroe I{JII' 6nnexoe K HeNryo Eaftoc (?) _ t.orepuB. 3aveqaHug._ Brraenenrre poaa Rhaphibelas Naef, Lg22, Mbr cturTaeNrnpexa_ AeBpeMeHHbtM' A'Hes ourcan ,,oa gTIiM HagBa'Ireu He6onsuge ErnoBl{arlbre pocrffi::
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Spanioteuthis Gustomesov, Tnu poAa - Spanioteuthis okscheven_ g6O. sls GustonEsov, 1960, Heo*orvrLpyccroli prrrlo,ur. pocrprr, cxarbre B cnr.r'Ho_ 6prcuHornl HalpaBJre'rrt{, KOpOTKU., c arrbBeonofi, cMerue'Hoft 'rv. o x vcnuxgoft cropo'e u noqru npevofi oceaofi nr',nefi. Heorov. floaceveftcreo Pachyteuthinau Stoil"y, 1g1g ( nom. transl. ex pachyteu_ thidae stolley, 1919). F;il;; Ha Hasarrb'brx craAr{sx pasBurr.e or'ocrreJrbrro KoporKrre, cy6uurnrnApnqecxue ufin caa6o BepeTeHoBtrAubre. Eartoc _ slrrr P a c h y t e u t l z a sB a y l e e t Z e i l l e r , lg7g. Tun poIa _ Belemnites excentralis Younget Bird, Lg22, [email protected]. Fo"rp"r Koporrs{e un' yMepe'Ho yArrn_ '"J,r5 HeHHbre c orHocrfren'rofi arursofi qo 400_500, cmruno-6ptorunoft AltaMerp 6oarute .nocrrearrbr.o*pr-on-;;;;; 6oxoaoro ,nn,n paBe' eMy, BeprrIEHa alrr,Beonbr lf oceBas rrunus HegxaqnTenbno cNreueubr r 6prcrunofi cropoge. Lono" _ r.or€pt{B; rloapoa Pachyt.eutftis Bayle et Zeillern i67g. poctpu yMepe'Ho yAnlrre'Hbre,
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noApoAs - Belemnites bteaiaf l o a p o A S i m o b e l u sG u s t o m e s o v , 1 9 5 6 . T n n Pocrpnt KopoTKIre c IIp!paBHIIHbI. Pyccrofi oKcoop[ Bepxllufi 1892, rrs Pavlow, qacTtr 125-250. TynrreHHo* aepuruHoft, oruocuTeJlr'sas IJIIUrfi nocneanbBeonspHga Kenaoaeft - 6eppuac. poaa - Belemnites magnificus Lagonibelu,s Quslemesov, 1956. Tnn paBH11HbI.Pocrpsr arr 1845, HuxHeBon)IcKI{fi nogbspyc Pyccrofi dbrbigny, qacfli or 3OO-5O0 no TsHyTbIe, c oTHocuTeIIbIIofi AJIUHoft nocfleanbBeoflspnofi oTJII{qaeTCs pOCTpa He AJITmA OTIIOC[TenbHas pasBIrTUs 14OO, Ba roHbrx cTaAUsX U OceBag III;HUg 11IrUMegbtue, qeM y BgpOCnOro )KUBOTHOTO.BeputrHa anbpegJrbl for€PlrB' cMeuleHbt x 6prcruxofi cropoue. Kennosefi lrnr c)KaTbtM c Ilonpon Lagonibelus Gustomesov' 1956. Pocrpu c olp]r1.tlbuvt y e3pocnbD( seM Kopoqer cTaAusx Ha llaqallbHblx ceqeur{eM, doxoa nonepeqgbrM xrrBornblx. orcc$opa - for€Pl{B. noApoAa - Belemnites I T o a p o n H o l c g , b e l o i d e sG u s t o m e s o v , 1 9 5 6 . T u n Pocrpbl c cllclrblM B crlltl|Qparurun.' KetlnoBeft L842, beaumontianus dtorbigny , 6proruxofi 6opoeaofit c AJII{HHofi ceqenueM, nonepeqxblM HanpaBJreHI{r{ uo.bprounroNr 6eppUaC' KennOeefi Ha Haqallbgbtx cTaAUsx yMepeggo BbrTtlHyTbIe. poAa - Belemnites subquodratus Roemer' Acroteuthis Stolley, 1919. Tun tlacTu OPf. Poctpu KopoTlqe urut yMepengo ceBepsofi rorepuB 1836, Hu)KIIufi qacTu oT 15O AO 4OOyAJrI{HeHHbterc oTHocHTeJIbHofi AnUHoft uocneanl'Beotupnoft aJIbBeOJtbIt{ oceBas BepulElra 6orosoro' 5OO, cmrnHo-6ploujHofi AI{AMerp MeHr,lxe SIIT' Kennoggfi CtOpOHe. 6ptotuHofi ITItglg CUnbBO CNtetueHblK 1919. Pocrpur KpyIIHbIe' yMepegno ctraTble B Ilonpo1 Acroteuthis Stolley, HanpaBtrenllu, 6ppumas 6Opoeaa pasBUTa cna6Ot Ea HAqAJIbtlbLK CmTHHO-6pptuHor\,t - sllro pocrpbr orHocurenbno cna6o ynrurneubl. Huxgreeon)Kcrrfi rlo[bgpyc cra[usx P a c h y t e u t h is n o A p o A s T u n 1 9 5 6 , l l o a p o a M i c r o b e l u sG u s t o m e s o v , (Microb-elus) hrimholzi Gustomesov, 1960, cpelHlril KennoBefi fuccrofi paBEulrbt' pocrpu He6oflbtxue, cgnbHo cxarbre B cmtuuo-6prcuruov HallpaBneHulr' co cna6o yMepeHHo yAF BbrpaxeHHoft 6prcuruofi 6opoeAoft, Ha HaqarbBbuc c"aAIffIX pasBtlrlrs I'OTepI{B. turHeHHbF. Kennosefi roApoAs - AcroI-loApoa Boreioteutliis Sachs et Nalnjaeva, 1966. Tun roAtHrxHeBorlxcKxfi 1966, teuthis (Boreioteuthis) niiga Sachs et Nalnjaeva, S CnUHHO-6ppu.rHoM c)KaTbIe crJlbgo Kpymrbre, spyc ceBepHoft cm6uptt. Poc,rprl Ha HaqaJIbHbI'XCTauarrpaBneHr{H, c xopoluo pasaurofi annnxofi 6prouxofi 6opoeaofi' 6appev. trrlsx pa3Burus ornocurenbno cunbno yaJrlrlrernbte. Otc0opa 1919. Poctprt c 6prcur1rofi6opoaAofi, Ha-Cerueftcrso Oxyteuthidae Stolley, .IuHaloueficfl or BepuruHbr u Bbrpa)Kengofi B pagtrutlHofi cteueHB. Ha 6orogblx c,G x o.9p9.9p a gaqHefi rracru pocrpa clo4gggg-9fl-:y*re ponax rrapHbre wgt, 6op9say. Ocea6; rrllgus cMeuena x 6prcur dbpurrHgF_o[onyp uue Aoxoaguyrr'ffi nofi' Ciopbire.'EanaHxuu - alrr. p o A a - B e l e m n i t e s b r u n s u i c e n s i sS t r o m Oxyieuthis Stolley, 1911.. Tun beck, 1861, Bepxguft 6apper"l ceBepHoft qacrl{ OPf. Pocrprt AJrllllHbre, cna6o BepeTeHoBIIAHbIeurulcy6uunEHApltqecKuecoKp}rnnblMnonepeqHblMceqeH[eM.Eprcrrr tIaCTU. BanagXfOr - SftT' nar 6OpO3Aa pa3Brra Cna69r TOnbKO B npgBepurrggoft BbtaeJutrb B poae Oxyteuthis IIpe&' Mu ue cotrnu BosNro)KHbIM I-lpnveqaHue. noApoabr Validoteuthis tr Fususiteuthis" no)Ke'Hbre A.E. Inasynoaoft (1969) BKnoqeH B cocTaB Acroteuthis s's" [tts gTI{x 6u1' [oApoAoB Aonxeg flepnurft I{3 ocHOBaHItfi' elge HeAOCTaTOTIHO BUA' OAIIH coAepxaqeno BToponO, BbTAeJIeHI{e Belemnites absolutiformis poAa A u l o c o t e u t f t i sS t o l l e y , 1 9 1 1 . T n n l{ru| Sinzovr., L877, eepxxufi 6appev floeonxbs. Poctput cna6o BepeTeHosltruIofi aflbBeonqpxofi aocrurarcurefi 6oposnofr, 6prcrrrHoii c cy6UurutHgpt{qecxofi $oplrsl, qacrlt. forepun - 6apPeu. noAoTpsA C.Eneuxufi (Jeletzky, 1965, 1966) npeanoxun B BblAenseurtfi HM ceueftcrsa BKllotlttTb Nalnjaeva et ) Belemnitina (=Cylindroteuthaceae Sachs u Bayanoteuthidae. Aaropst, Belemnoteuthidae, Chondroteuthidae, Diplobelidae gTuM I'pynnaM 6enevnutoa, a 1967 t.o BbICI(AHe pacnonaraq MaTepuanoM no n Bayanoteuthidae HapaBHe qro6br Belemnoteuthidae ceNaeficraa 3anr{cr, 3a ro, il6
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c paccl\rarpl'BaeNtbl'rt{ SAecb npynlaNrl{ 6eneungroB nepeBecrr,r B paHr HaaceNlerlcre Belemnoteuthaceae Buyunoteuthaceae. B xacroflruee BpeMs B€cKlrx oc" HosaHtrfi lrJrs nepecMoTpa srono nofloxeglrs aBTopbr He BnAsr. B nepaoe Lg Hag_ BalIlIbIx saAcer"reficrB Moryr 6rtrs BKJrIoqeHbr ceueftcraa Chondroteuthida" Oifiou"lidae. " (Belemnopsina Jeletz_ L,, /aF n K y , rl rv. r o b' B elemnopseina Jeretzky, rgoo, w".t".r-*l-1gzg). pocrprr Beps TeHoBI{aHste' cy6u,nuHnp[rtecKl{e' pexe cy6roHr.recKl{e c 6opoeaaM[ B nepeaneft rracrlr' B ansaeonapHofi qacrln pocrpa na6rnoqaercR cnaftKa, r'on[a [epexo'gqarr B uteJrb. CpanuoxoHbr t{Meror KoxrqecK}iro 0opvy H rro AaHHbrM IO. Eneuxoro (Jeletzky, 1966) xapax'epusylorcrr oTcyrcrBr{eM KaNrepHbrx orJroxe'ufi. Toap -
MEIL
pocrprr co crranHofi 6oposnofi, HagH_ garcuefics or nepeAHeno KoHua Lr colpoBoxAaloueftcc cnafixofi. CbeBafl Jnruug cMF ueHa x 6prcurxoft cropone. Toap _ 8rIT. lloacelaefi.r"o_ 1962. pocrpnr ae_ peteiionuhurte, yAnI{HeHHbte'Ha ", HarraJibHbrxciaqusx Me'ee.yaJII{HeHHbre. Hapgny co cwIHHoft 6opoeloft qacro ua6ruoqaercs 6prcrunan 6opoena rnu y11noure'uer Ha cnnHnoft cropoue cnaftxa. Toap _ sllrcrufi 6afioc ( ? 6ar). L e n o b e l u s G u s t o m e s o v ,1 g G G . T u n p o A a _ L e n o b e l u s l e n e n s i sG u s t o m e _ sov' 1966' roap ceaepHoft cu6Hpu. Poc'pu Bepere'oBua'bre rnu cy6uunrH'pnqecr^I{e' cnuHnas 6opoaaa pasBI{Ta B nepeaHeft qactu pocTpa, 6prcuuan Hay}rHaercs or Bepu[Hbl I{ He Bcel'aa aoxoAltr ao anbBeoJurpHofi .racrl{ pocrpa. Toap aaJIeH. P s e u d o d i c o e l i t e sS a c h s . T u n poaa _ Dicoelitesbidgieui Sachs, 1961, suxHufi aaneH ceeepnoft cr6r'rpr'r. Pocrprr BepereHoBr.u'bre nmr cy6uunu'AprrqecKre' c xopotroi'i ansseonoft. Hapray co clruuuori 6opoeaofi s nepeAgefi qacr[ pocrpa ecrb cna6o Bbrpax(enHan 6prcruHaa 6oposaa, y HeKoropbu( Bt{aoB nepe_ xoasuas B yttnouextle LIJrI{BoBce oTcyrcrBJnouas. I}prrneprunxHarr qacrb rnalF KaR, 6es 6opoea. Toap - HHxHHfi 6arioc. Duvaliinae P+u&o* . 'roAceveficrno 1914. orHocurenb'o KopoTKr{e pocrpb, co cnl{HHofi 6oposnofi,"ffi-nbHbrx craAusx 6onee y'rn{HeH'bre, qeM B3pocnbre. Kenaonefi - €nT. Rhopaloteuthis Lissajous, 1g15 (= protltn,a/ja Schn,egler, l94g) , TunoM poAa npeAnanaercs cqHTaTb Rhopalotetttltis dilucidtts Gu.rtor""oo, 196g, BepxHnfi xennoBeft-Hn)KHrfi oxc{opa Kprrva, llpennoxesHuft M.Jbrccaxy Tr{rroBo' BuA Belenmites saut'|anau.id'orbigny, cyArr no pucyH*aru caMoro a,op6uxsa, r{Nreer pocrp He co cnuuhot?, a c 6prournofi 6opostrofi. orxocHrenbHo KopoTKr{e pocrpbr 6ynaeonuanoii nnll BepereHoBl'raHorl co cnr{HHoft 6opoeaofi B anbBeonapuofi {opvr,r qacrll I{ tuenblo' ua 6prcruuoft cropoHe pacnoJroxeHo cyxarcueecrr B,,epeAn y,,nG rueHr{e, onpaHr.rqeHuo6cnaor,;ffi*paexoagurrv'cs 60po.50"r". E;; _ oxcQopa. rlpnvevaliiite. AnfciB:r-* 1967 r;" RcKlnoqiltt iiho,palobelus r{s rrrrcrrapoAoB Duvaliinae, BxrIIot{I{Bel'o B crrHoHr{Mrrr(y poAa Hibolite.s bo"u*o Jluccaxy (Lissajous, 1 9 1 5 , 1 9 2 S ) r . rn o e x e f i y r . a v e B c K a q( p u g a c z e w s k a , 1gS7, 1961), B.A. lycrorrecoB H E.A. ycneHcxag (tg6s); i{.crogHoBa-Bepfu'oBa (rgog) aanl'I qerxuft al{afHo3 ororo poAa' yKa3blBae Ha HaJurtl}reB nepeaHefi qacru pocrpoB c[I{HHoit 6oposalr, xapexrepuofi llg npeac'r'aBarenefi ceueficrna Duvaliidae. Conobelus Stolley, 1g1g. Tun poaa _ Belemnites conopltortts Oppel, 1858' TuToH t{exocnosaKuL,. He6onr,urue poc'pbr cy6uumlHqpuqecKoh urnt cy6xoHl{qecKot"lSopvbl c oxpynnr,rM ceqeHr{eM H anrggofr cnrugoft 6oposaoft. ArsBeora 3aHl{Nraer oKono nonoBr{Hbrpocrpa. Kennoeeft _ norepue, B e r r i a s i b e l r r s[ ) e l a t t r e , 1 9 5 2 , T n n poAa B e l e n t r t i t e se x t i n c t o r i u sR a s _ pail, 1829, 6eppuac-BanaHXr{H @pauuuu. He6onrrure cy6xoH[qecKr{e pocrpb, c orrsHyrofi npunepuuuHofi qacrbro, AnHHnoficnusHoft 6opoeaoft r{ oKpyrnbu nonepeq_ HblM ceqeHl{eM' Ansneona 3aHuMaer oKono L/3 anu'"br pocrpa. TnroH-naJraH)Krrg. (astellanibelus ('ombomorel, r972. Tun p o A d - B e l e m r t i t e . so r b i g / t r - a , u ; i Duval-Jouve, 1841, Heoxol't opanuuu. Cna6o Bepere'oBr.rrrHbre KoporK,e pocrpbl
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poAa - Belemnites latus Blain1878. Tun Duualia Bayle et Zeiller, cnaBneuHtte c 6oxoa pocrpblt cllnbuo ville, L8': 2, HeoKolvt Qpaxuuu.He6onsrune cropoHe Bepunggft. AnuIt cnnnHofi cMelqeHBofi r cropoHoft 6prourxofi BbrnyKnofi c 8IIT. Beona KopoTKag, cnI{HHAg 6oposaa KopoTKag. TUToH poAaP s e u d . o d u r o i i oN a e f , L 9 2 2 ( P o l y g o n a l i a A k . A l i - Z a d e , 1 9 6 5 ) . T u n ne6onuPoctpur Qpanqrn. HeoKoM 1827, polygonalis Blainville, Belemnites qacrl{ cerreHueN414 ArtHHuue, BepereHoBuanbte c pov6u'recxl{M B aflbBeoJrtlpuoft For€PlrB. xoft curnurofi 6oposAofi. Turox pseudobelus Blainville, L827, Tun poaa - Pseudobelus bipartitus BlanorepuB Qpaxuuu. Tosrue cy6rurnrnrapn'recxoft 0opinville, L827, BanagxrH Mbt pocrpbr, clKarble c 6oro8, c Koporxoft cnunxoft u 6onee aJruI{HbINtu6oKogrtuu 6oposaav[, Haqt{Balouu{Mucq oT rtepeAHel'o. Kouua pocrpa. AnbBeona KopoTKas' Banarxnrr - TlrePItB. Ceveilcrso Betemnopsidae Naef. 192? (nom. transl. ex Belemnopsinae Pocrpu c 6prcurxoft 6opoaaoft, xaStolley, 1919). N".ilg22t-(Hastatidae qrtrarouefics or nepeAgero Kouua I{ conpoBo)KAatoueficg cnaftxoft' Ha 6oxoarx B cropoHax B anbBeonspuofi qacru pocrpa napHbte noJlocbl, aanee cxolulrunecq ueHr'saHl{Maer nIlHHs Ocesas ogtry 6opoeay, upocrrexnBarotrqrrocs ao BepuIIrHbI. panbHoe floflo]KeHue. poaa - Belemnites bessirtus B e l e m n o p s i sB a y l e e t Z e i l l e r , 1 8 ? 8 . T n n d'Orbigny , L842, 6ar Oparurn. Pocrpur cy6uruIrH.qprqecKre nnu cna6o BepersgAer or nepeAHeHoBlllgble, C KopoTrofi ant'eeoflofi. Eprcuruan 6oposAa AnI'lHHa9' qacTr{. TllToHr Aaneu ro pocTpa ' n i b i t i t e sAo efrc) npuBepusxsofi poAa - Belemnites hastatus Blainville, trtonito.t, t'gO8. Tun L827, rennoeefi - orc$opa Qpaxunr. Poctpbl yanrHeHHble, BepeTeHOeUAHOfiQorMbr, c xoporxofi arrbBeonofi. Epnuuan 6opoeAa 3axoat{T B nocreanbBeoncpHyn qagTb pocTpa. Hr,aeerCs cuafixa Ha 6prcuruOft ctopOue. EafiOc - anT. V MenoBbx npeacTaBrreneft poaa Hibolites 6prcuLHan6oposaa 3aveqaHns. KOpOqe H BIOJIHe B93MO111HO,qTo OHr, 3acIryxHBalOT BblAeneHHc.a ocO6utft poA' Mornu. Aeropu rdga3a OTcyTcTBtrs MaTep1ana no Hibolites 3To cnenaTb He poaa - Rhopaloteuthis somalierr'sis S o m a l i b e l u sJ e i e t z k y , L g 7 2 . T u n KHMepu4s Bocroqxofi AQpnrn. He6onsrune, cnerKa BepereHonuqHoft Spath, lgg5, c xopouo pa3BuToft 6ptotuxofi $Opr'rtl pogTpbt C oBaJtbHbIM nonepeqgbry ceqeg11eM' qacrlr. Kruepnax. B aflbaeonnpnofi 6oposaofi' p o A a - P s e u d o h i b o l i t e sn a t h o r ' s t i P s e u d o h i b o l i t e sB l i i t h g e n , 1 9 3 6 . T n n Blfthgen, 1936, HeoxoN,rUnnu6bpreHa. Pocrpur cna6o BepereHoBltaHble nmr cyG UrIII{HApIdqecK[e, c xopouo BblpaxeuHstvg 6oxoBblMu 6oposqalru. EprcruHaa 6opocaa I{ cnafixa He orMeqaloTcs. Heoxbv. poaa - Curtohibolites CurtohibotrtesStoianova-Vergilova, 1963. Tnn Bonnapuu. He6ont1963rnorepl{B-6appev trubatchensis Stojanova-Vergilova, pasnuroii 6pruutHoft 6oposaol't xopouro c pocrpg cy6ullmrgqpuqecxofi u11ae @pr"rut, y3 Am{HI]I pocTpa. BananxuH - 6appev' n anbBeonofi, eaHnvarcuteft ot L/2 no Belemnites minaret Raspail, Mesohibolites Stolley, 1919. Tu n p oaa pocl'pbl c nny6oxoft antF Cna6oaepereHoBLIAHL,Ie Qpanuun. arrr 1829, .6AppeNl aeonofi li=criafiKofi, nFoTgrnBaputeficg 3HaqureJlbHo AaJrbuie HaqaJul anbBeonbt. EprcurHaa 6oposna 3axoAI{T B nocneanbBeonspHJrKr qac'rb. Eapper.l - 8IIT. poaa - Llucrohibolites schaoriM u c r o h i b o l i t e sN a z a r i c h v i l i , 1 9 6 8 . T u n ensis Hetchinashvili, 1952, BepxH[fi 6apper'r lpysuu. Poctpt,l cpeA]Iuer cy6unmffiAp11qecKge, 3aKagql{Barcullecs KoporKutvt utunoBllaHbrNtorpocrKolvt. Eprcu-ruas 6opoaaa HaquHaercs y ycrbs u ucqesaer B cpeuHefi qacrl{ pocl'pa. I-lonepevnoe cerrenue oKpynnoe IIJII{ oBaJlr'Hofi @pvs6 BepxHufi 6appeM - oIlT. p o A a - i t e o h { b o l i t e sd u v a l i a e l o r n i s ParohibolitesStolley, 1919. Tnn Stolley, 1911, anr ceaepHofi qacru OPl. Poctpr,t BepereHoBll!.Hble'c)harbre c 6oposaarutHHa 6oxoebrx cro6oxoe, c KoporKoii 6prcurxofi 6opoeaoft, c aBot"rHblNrl{pouax. Aur - xuxuuf, ceHoMaH. 118
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N ' , e o h i b o r i r e ss t o l r e y , 1 9 1 1 ( N e o h i b o r i t o i d , e sA k . A l i - z a d e , I 964). Tr n p o A a - r o n p e q n o x e u u r cH . K . fopHa (rgogr Beremnit""lrotdistrombeck, 1861' HuxHufi anr ceBepuofi vacru oPf. Pocrpr,r se6onsul'e, cy6unr*H'pr*racxoii unu Bepereuosuanoft {opvsr, c KopoTKofi 6prcurnofi 6opoeaofi, nosrr He 3a_ xoasueft lanee BepuI{Ht'I aJlbBeoJtbt. 3aArurg npa'rua cnafiru oT Bepul'br anb'eorrbr r{.aer' ay.oo6pasHo r{sr'6aecb, x 6prcruHoniy *p"o. A;;;;;* xoporxas, sacr,o o6paeyetcs nceBaoanbBeona. Anr Hpxtrufi ceHor{aH. fl p u v e q a H I{ e' Tlrnoeofi BI{A poaa N eohibolites, flpeanoxerHsrfi lllronneev ( Stolley, 191g) _ nrU^"it"" ,"^iranalicularus Blainville, g neftcrsrrrerrbnocTtr npr{HaAnexllT K Br,rEeneHHoMyB 1911 r. UJronneeN{ poAy Mesoteuthis r noelro. My saMensercq Br{norv,B.eualdi Strombeck (lopn, fga"gi:-pocrpu c 6proruuofi 60_ posAoft, HaquHabruef,cg oT [epeaHero Kogua pocrpa lr conpoBoxAapueficg cnafi_ xoft' Ha crmssofi cropo'e raK)Ke [Meercs cna6ee Bbrpaxe'Han 6opos[a, HaglrHalouagcs oT nepe[Heno KoHua Il Toxe corpoBott(Aarcuascg cnafixoft. Oceaag rurHus cMeueHa Nh6prourHoftcropoae. Eafioc _ TrrroHo 't
Dicoerites Boehm,1906
ir"a;."iir*
s,on"y,
tg27). Tnn poAa _ Be_ lemnites dicoerus Rothpretz, 1g92, *"'nor"fi HHaoHes,.". p*rpbr yaJu'e'Hbre BepereHoBlrA'bre uaz cy6unnr'ApnqecKrfe. 'aftoc - orc@pa. Conodicoelites Stevens, tdosa. f lrr.;;" a _ D i c o e l i t e s h e e u w e n s i sB o e h m , 1912' BepxHss 'opa z'ao'esuu. Pocrp", xoporKlre, cy6xoHr.Ie'xofi umr cy6uunnnaprqecKofi _ TrfroHr @prvrrr. Eafioc"p""*TeJrbtro t lg24r-locrpbr c 6oxogrrvn 6oposaaMIr B nepenxeft qacrr{. Anr _ ,."arprr*r. v''rv r E rvr(a'r-n^'1 'll€IJA r"^ r ' Cn"rar"r[Ka AaHa nno pa6ore f.Cru_
BeHCa
( SteVenS,
1965).
Peratobelus Whitehouse, Lg24. Tnn p o A e _ B e l e m n i t e . so r y s Tenison_ woods' 1884' aepxnrft anr AscrpaJir{ri' cy6unrucrAprqecxxe ,,Jtrr cy6xorurvecrcre pocrpbl c IlByMs 6prourxo-6o*oBblMl{ 6oposaavr{ B nepenneft qacTno Bep.nrrf, anr. Dimitobelus whitehouse, Lg24 ( T'etrabeiis whitehouse, Lg24, cheirobelus' Whitehouse' Lg24). Tun B e l e m n i t e sc a n h a m i T a t e r l g g o , PoAa BeprF Hrfi anr6 Bocro'r'ofi Aacrpanru. BepereHoBr{rrr'e pocrpbr c AByMe naparrr foxoBbtx 6oposa B fleperneft .{acr[. Anu6 _ MaacTpt{xT. CeNreficre" pocrpu c 6prcurxofl uenaro B ne_ pealr6fr-;acTilr'r co cnn'Ho-6orosrrr'rn 6opoeaaMu, .[epexo'surlrMu B c,,,,H'o-oo*ot Bbre yltnouenttg. C,enorrau _ MSBCTpru(T. Crcrevaruxa lana no pa6ore .[.fL Hafiarna - p o o , (1965). Poa Actinocamar Miller, 1g2g. Trn _ B e l e m n i t e st ) e r u s M i l l e r , 1823t caHToH Am'rau' llrrurHaprvecxre nnr cna6o Bepere'oBrrrurbre pocrpbr c oKpynJrblM ceqe'n'eM' Ha nosepx'ocTlt pocrpa sa6ruoAarorcs npoaorrrr'bre urrp'x,, I{ [onepeql{bre Mopurr{HKn. Anuseona tto L/ LO pocrpa, np11 paspJrureg[Ir ee creHoK o6paeyercs NteJINag[ceBAoanbaeoJra. Arurgbr CenolraH _ CSHToII. Ilonpoa'Actinocamax-Millerr1g2g. Tun no ApoAan _ .Actinocamax verus Miller, 182g, caurox Artrnun,
- Betemnites pte,,"n;ff;l ulJi'"ili:;;:T;;:"'f##.t64. r'u noApoA, rlonpoa ,orlr:r::.oror'N"1din, 1964. Tun noApoAa _ Actinocamax gros_ souvrei Janet, 1ggO, ,"p*"nfi carrron Ceaeproft eaxunn Poa Gonioteuthis Bayle, 1g7g. Trn Or^" _ Belemnites quadratusBlein_ ville, !827, s}rxuufi xaMrrax Oparuun. llrnrrrnp6rec1rre rnn cna6oBepe,regoBfi_ Hble pocrpbl c ropotrofi 6prcurnoft ulenblo r pasBrrofi ncesaoalbBeonoft. BepxHocrl{' noMIrMo oTnegaTl(oB Ha noc@yAoB, npoAorrbrrbrx lxTprrxoB E crpyex, o6rrrr Ho xoporuo Bbrpalr(eHa gepH[crocrrr. Cagron _ Hrrg1uft raMnaH. I-loapoaco"r?r:yrlrs
bayre, 1g?g. f;" n'ooooa& - Beremnites quadra_ tus Blai'nville, 1g27, ,ror# KaNrraH Opauuun.
,",,Tff : ir","i'; nT,TJ:lt ?p l.eG;: i;;;;
r poAa - Actinocam ax tund gil9
p o A a - B e l e m n i t e sn a n t n i l l a t u s P o a B e l e n t n e l l o c a n t aNxa j d i n , 1 9 6 4 . T u n Pocrpur ul{ill{HApl{uecKoft, raHlOxNofi lllseuuu. Nilsson , LBZ'I, uuxnHft ceHon ueroBIIAHoil
llnu
BeperegoBnAHofi
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6OpOeAxu. EoKosl're 60XOfi CrOpOH, BcerAa xOpOIxOBblpaxeHbl cnt{HHO-6oxOBsre He Bcerqa' Y @pv c coxpaHsroTcn cocyAoB posAKH H otneqaTKlt KpoBeHocgblx nepexgqgqas B lUeflb' 6prcurUan KOpOTKaq Bbrpa)Kega xgpoulg nqeBAgangaeonOfi HuxxUit KaN{naH' cnaftKofi' nerny6oxyro 6prcrunyrcr.6oposAy, KoTopas conpsxega co ' p o A a B e l e m n i t e smucronatus T u n Pon Belentnitella d'Orbigny, L842. cy6xonFlecnmt cy6uHnuHApnqecKoft Pocrput ceHoH-oPf. BepxHuft Link, 18O?, ngnepet{HblM CeqeHI{eM' fOft @pVbt, C g1UfiaMI{ Ha SaAHeM KOHUeT C OKpyrnbIM uTpl{xlto BproruI{ npgAonbHr'le Ha nOnepXHOCTIt Ha6mOAaIoTCg oTneqaTKI{ CocyAOB Maacrpltxr' CaHtoH Hag uenb Koporre anbBeonbl. p oAa - Belenrtites lanceolatus Schlopon Betr*rrtto Tun iowak, 1913. pocTpbl co cnnax(eHHoii noBepeteHoBl{llHble BepxHr{fi ceHoH OPl. theim, 1813, KB\{Bepxnufi anbBeonbl' Haqana BepXHOCTbIo U 6prcurnOii ulenblo' AocTllrarculeft n6H -
MaaCTpHXT' p oAa - Fusiteuthis polonica KonTun 1962. Fusiteutltis Kongiel, pocTpbl co Crfla)KeuHofi noBepxHocTblo I'I fJonurua. BepereuoBllEtllble giel, 1962, vaactpux'r' BepxHnft cna6o BblpaxeHHoft urenblo' Pon
CTPATI,ITPAOI{qECKOE 3II A qETII,IE PAHHE- I4 CPEIHEIOPCKUX FEJTEMHTITOB
Boepacruble rIpeAeJIbIpaclpocrpagegus BltAoB 6enercruroB noaceNreficrs Nannobelinae ra Passaloteuthinae u ceMeftcraa Hastitidae aBropaMr{ paccrvrarpr{BaJrr{cb B 797o^ r. oA'axo, HatruHas c 196g F.r HB ceBepe cu6upu, B ocHoBHoM, T'H' HansHseeofi u T.v, Krpuuofi [poltsBeAeHbr aonorrHt{TenbHue oonsurre c6opsr paHHe- I{ cpeaHelopcKltx 6eneunuroB lrs xoporuo crparr$ruupoBaHgbrx paspesoB, JrBsoannbre c HaxoaKaMr Apyrnx rpyrrn @yurr. Enaroaaps cKasauuoMy oKa3aJrocb BqSMOX(HbIMCyuecrBeHHo yrotrHtlrb BeprltKanbHoe pacnpocrpaHeHue orAeJrbHbrx BHAoB u KoNlnneKcoB 6enel'lHnroB. lloerouy Huxe paccl,rorpeHo crparunpa{rvecKoe sHaqeHl{e Bcex npyffI panBe- H cpeAHelopcKl{x 6enevuuroB, BKJrroqag Nannobelinae, Passaloteuthinae,-Hastitidae u onucaHHoe a B T o p a M r {B 1 9 6 4 u 1 g 6 6 n n . ceveficrso Cylindroteuthidae. B ocuoay cTparl{npa{u'recr:oro pacrrne'eHnrr paHHe- H cpeaHercpcKr{x oTnox*uuft cu6upr u AanbHeno Bocroxa aBTopaMH, KaK yxe yKasbrBalrocb Bo BBeAeHr{r{, TlonoxeHa crparlfpaQrvecxaq cxeMa, paspa6oraHHarr Ha Hoeocu6upcror"r coB€rl6lrl'{ B L972 r' (caxc, AanHc }t ap.r Lgi2). Hexo'roprre cnopHbre nonoxeHsrr eroft cxeMbl' BbtsBJunouuecg npr npnnoxeH[I{ ee K KoHKperHbrM paopegaM, oco6o oroBapl{Balorcrr trpu Aaargefiurev rgfloxeur{I{. floqtu ace c6opsr 6enerurHt{ToBnocneAur.rx Jrer HaAex
B ornoxegusx nerraHncKono' crHeMropcKono u nnunc6axcKoro spycoB BM€cT€ C AMMOHIlTaMtt AOCTOBep[bIe HaxoAKtr pocrpoB 6eneN,rgnros Heu3BecrHbr. A'c' u A.A. .llaruc 6rtrm Hafiaenst Ha ovonoHcKo'r MaccrBe Qpanvoxousr 6eaeuHIfroB B cnogx c paHHeCI{HeMIOpcKxMutr llo3lrelrrnc6axcxuMt{ aMMOHUTaMI{,HO crrcr€MaTuqecKlno rIpI{HaAIe)KHocrb no Spanr"roKoHaM o[peAeJrl{Tb HeJrb3s, K ToIr'$-;1s He rrcKtlrcqeEor qro Qpanr'roxolrbl Monnlr npnlraAnexarb 6eepocrpoagu Qopr"rav. Ha nepr@pnu cx6npcrofi nnar@pr,abr Ha np"ruo npuToKax AHa6apa n xa p'cunefi (Burnoficxaa auaarHa) B cJrosx, oxaparTepr{soBaHnu* ceoftcrBeuHbrMu ToJIbKo nronrc6axy ABycrBopKaM[ lMyophoria cf. lingonensis (Dum.), M"laeaigata
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l i te s , Clas t ot eut h i s Orthobclus. flpn aroru ycrangDrrero npeo6Jraaalouee oxaqeure Blr[oB c f,AJIEE€nr HbrMn pocrpaMl{, BeAun{Mtr rrpe[nono)Knrerrblro cgo6oAlro[rlaBaroqnfi o6pa,e rotguf,. K nPqAoBBnnu c rcoporKnMr pocrpaMrrr np[ypoqennbre, Kax Moxno np€Anolararbr peAxo. Bce BagBattutte. g4lbr rrepexoAsr a ngr,rcllil nrnr o6cranoBtra!{, Bcrpetlflorcs FBP, a oAIIH - Brachybelus (Arcobelus) dolosus - AoxIIBa€T Ao cepeAtBbl aanega. Cnenyer rtoaqeprcIyrb, trro a @rurs:r oTrpHTolo Mops xa Ovorprcf,oM Mach cI|Be n a llpraepxosucKoM rrporlr6e pocrprr 6ereurrlrrroB B nruorc6axe coBeplrr€BHo orcyTcTBylor. flesuAll,touy, o6nacrn orKpbrloro Mopc a Aprrn'recKoM 6aoceltne B KoHrIe nanuc6axa eue tre 6uml sacerlegbl 6enevrltraMu. B Hnsax Toapa B sone Dactylioceras tenuicostatum KoMrrrerc 6enelcuroa xa nepuQepur Cr6upcroft unar@pr'mr a npn6perclofi soxe noKa He JrcTaHoBJreH. B o6nacrb orKpblToro Mops 6eneunurbr roxe eue rrorrru rre rrponttxaltrr. Tonsro eA[Hutrrbte HaxoaKt{ pocrpoB caenauur a pafioxe OvonoucKoro MaccEBa. 3necs safinex Catateuthis aff. westhaiensis BnA, He orMetrennrfr na nepu@pnr Cr 6upcxofi nnar@puu. 3na'rurenrxo 6onee 6onatslMr cragoBsrcs. KoMnnexcrr 6eneuurrroa B goue Harpoceras falcifer. B npn6pexsbrx @uuax na nepu@puu Cu6rpcxofi nnar@pubr nor nBnserctr psl HoBbLKBIraoB poaoB Acrocoelites, Catateuthis, Brachybelus, 1o9Bnrrrorcs HoBbre panee He orMerraBunecs poqbr Mesoteuthis (c 6 BsAaMtl, Nannobelus, Dactyloteuthis. O6rqee KorrrrqesDBoBrrtroB Q4l Bospactaer 6onee qeM BTpoe. Focnogc'rBo B KoMnneKcax 6ener'nruroB neper(oArr cyr Acrocoelites x Mesorcuthis" QTrlacrrr Catateuthis. Kar n s 6onee HHsKrrx rpprroouTax, npeo6naAartl B KoMnnerce BuAbI c y[nuHeEHbIMf, pocrpaMu ( npeanonoxrrenbHo cso6oAHonnasaroruse).1 B $auunx orxpbrroro Mops a paf,one Ouonoscroro MaccrBa B orttortentgr( @uu Harpoceras falcifer pocrpbr 6enevxnroB Bce eue HeMuonorurcnennrr. Haf,aeubr (nau6onee xapaxrepnbr 8 auAoe, us Koroprx 3 nprxaafle)mr x Acrocoelites A.trisulcosus u A.subtenuis), u 3 - x Catateuthis" Iloqnr B[Boe (no 44) aoepacraer KorrrqecrBo .BrraoB 6enevnrroB B npn6pencxrrx Sauusx B 3oHe Dactylioceras commune (Buruoficxag BtraArxa, sanaAxsfr cxnou flpraepxoqucKoro nporu6a, Enuceficxe.Ilexcrsfi nporr6). Bnepabre s[ecb o6ua(Lenobelus lepyxl{Baercs eHAeMIrqHasrplrrra 6enepcrnros Pseudodicoelitinae nensis, Pseudodicoelites primoris), rrplrnaane]Kauas K HoBoMy naacevefictay Duvaliaceae. Bnepnue nosBrqrorcs B ceBepHbrx Mops.x Hastitidae (Paruhastr'tes medius ). Epaaaa, Bce 9TI{ HoBbre rpynnbr [peacraBnexbr peAxnMu HaxotrKaMu. Xapaxtepnofi oco6exuocrbro KoMnneKca soHbr Dactylioceras commune, orrrcqapulefi grot xoMnneKc or Hux(eJlexaruux, HaAo crlnTarb uttpoKoe pagBltrlte BItaoB c KOpOTKXMLpOClpaMI{; BeAIUUTC, KaK Mbr npeAnOJrAnAeM,IlpltAorrnbd O6pas xItc]HU. Taxoaur Clastoteuthis, N annobelus, roporKopocrpoBbre Brrnbl Mesoteuthis, Brac hy b e lus, D ac ty lot e ut hi s. 3Ha'rurensuo o6orauaercs B sone Dactylioceras commune lr roMtrlrerc 6eneuHl{ToB a @uusx orKpblToro rvrops (Or.aononcKufi r.,racc[a, flpreepxosncK[fi nporr6]. 3aecu o6uapyxen* 29 auaoa, B ocxoBHov @pvu c yAJrrHeHBbrMtrpocrpaM[, npxcnoco6nessrre x ceo6oauoMy nraBannro. Ftpeo6naAarcT npelcraBrreJu Acrocoelites ( A.tri sulcosr/s,A.subtmuis, usBecrnufi roasro s erofi goue 'A.pergrocilis u Bt{abl co c)KarblMt{ B cmouro-6pbuHoM Ha[paBJrexrrrr pocrpaMf, - A. he'C. idonea donensis u A.omolonensis), Catateuthis (C.invisa, ). Hafigexu B arofi (H.kinasovi). Holcobelus poaa It nepBble soue rlpegcraBlrrenn -Br'resre c reM E $aunnx orKpblToro Nrops B sone Dactylioceras commune eue Boace Her Hastitidae u Pseudodicoelitinae. B nepxnefi soue Htrxnero roapa - B soHe Zugodactylites braunianus - Kor,rrh nexcbt 6enelrHuroB ocrarorcc oqexb cxoAnbtMn c raroBbrMtt soubr Dactylioceras commune. B npu6pexHblx @urnx, na nepn{epnr Cn6upcrofi nnadopMu o6uee xo(46} coxpansercs [orrru HelrsMeuubrM. Oanaxo ue. Jlt{qecrgo BnAoB 6enevxnros KoTopble Buabl (Acrocoelifes triscr'ssa, A.janenschi, Catateuthis longa) He rre126
pexoAsr H3 3oHbr Dactylioceras commune B oony Zugodactylites braunianus. Ecrt Buabl' KoTOpbIe orpaHnqeHbr B cBoeM pacnpocTpaHeHuD aroft eoHoft (clastoteuthis atctica, Parahastites horgoensis), s TaKXe BuepBbre uosBns,ouuecs B aassofi ooHe r{ noartrrMaroutlecs BBepx tto paspesy B cpeanrorc topy ( parahastites matchaensis' Masoteuthis inomata, Lenobelus viligaensr's,).rlpeacraaulenuHa.stitidae u Pseudodicoetinae B o*o)Kerrrsx so'b( 2d;J;;-rylres braunianus xors eue II peAKIt' Ho Bcrpeqalorcs yxe qaue I{38€cr ust 7 BI{aoB, npma11ne)Karulrx gTr{M npyrmaM. r B o6nactl{ oTKpblTono rvlopfl KoMrrJreKc 6eneprgl{roB soHbr Zugodactylites braunianus (34 n'na) cy':ecrBe'Ho ;r;;*;;" or KoMnrre*ca eonrr Dactylioceras commune' flosBn'Ilorcq Jlulub "" eagnurrHbre HoBbre Br{!u ( Mesoteuthis inornata, M' aff' pergrandis )' flo-npexueMy _ _ _ _ _ - r oTcyrcrByror y rvr .v ruJvr rpeAcraBr{Tenu rrpEAU.rarrr{TeJ Hastitidae u pseudodicoelitinae BrtaeaeHre BepxlleroapcKl'Ix Korv,rrnexcoa 6eaei{Hr.troB ua ceaepe cccp HaraJrK[Baercs na 6oasurl{e Tpyl'ocrlt, Bepxnuft Toap oxapar(Tepr{soBaH aMMoHnTa n,rr u AeJIuTcs Ha aBe soHbl ToJIbI(o a pafioHe Or'ronoucxono Nraccuea. Ha nepr@pa u Ca6upcroft unar$opr'Ibl HaxoAoK Bepxueroapcxt{x aMMotrtrroB BoBce xer. floeroMy 3aecb no.[HeroapcKue KoM[neKclr 6eneuultroB B HacTosulee BpeMs HaAe)KHoycTAHOBI{Tb
HE'Ib3g.
Tpynxoctlr c aarHpoerorl paccMaTprrBaeMbDc oTnoxeHfift ycpy6nsrorce €r,,€ pacxoxaeHusMll B onpeaeneHHH Boopacra aMMoHHTots B paspesax Burnog, Jlenur u ux n p I { T o K o B 'A ' A ' r A ' c ' a a n u c s t ( r g o s ) o n r e c n u p s e u d o r i o c e r a sa l i e n u m A . D a8is BI{A' xapaKTepxeyrcuuft Bepxrue nopr{soHTbr Mopcxoft ropu ua Buruoe, x BepxaN'tHuxHeno Toapa - K 3oH€ Zugodactylites braunianus.ocnoaasueM K eTcLMy gB!{nllcb HaxoaKt{ Ha OvonoucxoM Macc[Be aMMoHI{ToB, o[pepeJrgerurrx A.A.Aa_ rl{c KaK Ps.alienum, BMecre c Zugodactylites btaunianus (d,Orb.). Oauaxo a -Zugod,actylites, Burnoficxofi snaarHe Ps-alienum cJroeB c BM€c""rp"""*o,-u"rr. Te c xapaKTep'brM aru{ aaJre'a KOlrnIrgKcoM aBycrBop ox: Qxytoma jacksoni P o m p ' , o ' i a c h s o n i h e l i m i a r e n s i sB o d y l . , o . | - i r i n o , V e l i k z h . , c a m p t o n e c t e s( B o reionectes) sp., pseudomytiloides marchaensjs (petr..), Arctotis marchaensis velikzh', A"similis velikzilap. floerovy T.H. Krpuua, 4.c. Benr{Kxa'r{Ha I r H.M. ,0'xu'opuaee (tgzq) u T.H. Krpana H.f. Kpurunoasu (tgl+) cqu_ Taror cnor c ps. alienum HH)KHeaaneHcKIrMH. " Bepxnrft Toap yBepeuHo u aaxe c pas6uaxofi Ha ABe 3oHbr Bbraensercs roJrExo a fuuu$( oTKpblToro Mops a parioue ovonoucxor-o Maccr{Ba. K coxane'nro B grotvt nocneaHev pafioae KoMnJIeKcul 6enevsl{ToB B BepxHeM Toape HaM l{38€crHbt eue HeaocraroqHo. B soHe Collina mucronata Bce eue o6ununrr He[oAHtrMapuuecs Bbltxe erofi 3oHbllcrocoelites co cxarbrMr{ c 6oxon pocrpaNrr (A.tr;su/cosus' A' subtenuis, A. polaris), coxpaHqercir prra BHA.B Catateuthis, Brachybelus, Mesoteuthis, cnoficraenHbrx HlrxneMy roapy. C)aroepeveHHo nosBns_ Iorcs I{ HoBbIe B[Abl' xapaKTepHble ronbKo rlns Bepxreno Toapa ( Passaloteuthis ignota) unr nepexoAr'uue r{ B aane' ( ilresoteuthis pergrartdis, Horcoberus umarensis/. Her eure Pseudodicoelitinaeu llastitidae. B eoHe Pseudolioceras rosenkrantzi .o*pu*""Tcs Konr{rrecTBonpeacTaBuTenefi caoftcrBeHHono $auunr"r oTKpbrroro Nropfl Hux.,,eroapcxono KoMnneKca u '* s B n s l o r c s n e p B b r eP s e u d o d i c o e l i t i na e ( L e n o b e l u s t,iliga)nsisl u Ilastitidae (parah a stite s marchaensi s ). Ha nepu$epuu cu6upcxoii nnar@prurbr B paspe3ax MenKoBoaHux @uufi crpatn'pasuvecKr{ Bbmre Fopr{ooHTa c Zugodacttiur)" brauniarttts, ,.". BepxoB HH)KHenoToapa' npucyrclByer Kolvlnnexc 6eneNrHI{ToB,olnuguulfi or "o,rue pa'HeroapcKoro' Koltnnexc gToT Moxer HMerb flooA'eroapcxrli Bospacr, Moxer or'ocl{?bF cg x paHHeMy aaneHy' Moxer oxBarblBarb KaK nosanuft Toap, TaK u paxunfi aareH. Ans nero xapaKTepHo coxpaHeHl{e Jrr{ulbearHuqHbrx BHaoB Acrocoelites, clastot e u t h i s , n o s B n e H n eH o B b I x B u a o B N a n r t o b e l u s parahastites (N.nordt,ihensis), (P'notatus)' o6napyxHBalorcrr npeAcraBaTenu poaa Hastites p.settd"odicoeliu tes ex gr' bidgieui' 3'a'{utenbHo Bo3pacraer no cpaBHeH,ro c Hr{x(Hr{MToapoM
127
u Pseudodicoelitinae u oatroBpeMeaxo y6omaer KoKoJrrrqecrBo BrraoB Hastitidae luqecrBo KoporropocrpoBbrx BltAoB P assaloteu thidae. Mops Bnyrp[ flpneepxocHcKor\o nponl6a roap BblAeJrserB rsunex orrpbtroro es aETaToqHo ycrroBHoo BosNro)Knorcnou N{exay nruHc6axclgrMl{ u @yxucTuqecKn oxapar(Tepf,soBarrubrMtr Hrrxxeaale$cKuMu ornoxenusMu saKrrloqalcrr arcnbro aeprcnrft u K EepxaM HUxHerO Aoapa, A, BOSMoXHor 9aHItSlbIUX CTTHOCSTCS TOap, BO€}MO)|(HO, cTrrrlHo l|tm Aaxe [eJrrrKoM - yxe K aafleHy. Ilpeo6nanaroT B sToM KoMnJIeKce PSeUdodicoelitinae n Hastitidae, Hapsay c HlrMr{ Bcrpeqarorcs peaKre Nannobelinae, Passaloteuthinae, 6onee qacrbre Mesoteuthis n Acrocoelites. Paroreaanexcntft KoMrrJreKc 6enevxuroB B npn6pexHrx @urrx, aarupyer'artfi uo ArctoHaxoAt(aM aMMOH!TT'oBE oTrracTu ADycrBopor (Oxytoma jacksoni Pomp., tis marchdensis Velikzh., A. similis Velikzh. r Ap.) cosepxsr eule AoBorbno MHONOqTCIIEHHbIE
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Bvecre Passdloteuthis, Clastoteuthis, Nannobelus, Brachybelus, Mesoteuthis. 6onee pe[Kue c HrrMu np[cyrcrByror Muonoqlrcneurrbre Parahastites, Hastites, Holcobelus, Pseudodicoelites, B[epBbte B paHHeM aaneue nosBJullorcn Sachsit{ro racaercn poAa Sachbelus, Rhabdobetus (?), Pseudodicoelites hibolitoides. sibelu's, To nMerorcs pasposueHgbre rpe6youuae noarBep)KAexrr{ yKa3aHl{rr H8 H8xoxaeHne eArHl{qHbtx pocTpoB aToFo poAa B ropHSoHTax, HeoxapaKTepl{3oBaHHbtx IIr}KHeaaJIeHcKuMlraMMoHuTar'au, 6rrrs MoxeT eue B Toape, B uenolr MenKoBo!nufi panneaareHcKlrfi ror'maerc 6eanee ToapcKro(' ll saKJlroqaer 28 snaoe. B @uusx orKpbrroro Mops B paHHeM aarrene nocnoacrByror Pseudodicoeli tinae n Hastitidae, B ToNr trscne HoBbre BuAbr: Pseudodicoelites hibolitoides, Hapsa BuaoB Sachsibelus. Hexoropue BI,rabI stites clavatiformis, H.vesicularis, onpaxnqeubr B cBoeM pacnpocrpaHentru roJIbKo Hr{xHuM aaneHoM: Pseudodicoelites clavatoides, Hastites grandis, Sachsibelus gnarus. B noa.IrHeHHbIXKonl{qecru Bax Bcrpeqarorcs rrpeacraBuTeflI{ Nannobelus, Brachybelus, Mesoleuthis, Parahasldles, flpuqeM HcxtttorlurenbHo re BuAbr, Koropbre nepe[rJru lrg roapa. Sror KoMIIJrexc, aa)Ke rrpeBocxoa.grunfi rro KoJrt{trecrBy BrrAoB (eO1 pauueroapcxufi, B ocHoBF npOrU6e. B pafiOXe OproHorvt, HaM nsBeCTeH nO pa3pesaN{ B I-IpUAepXO9HCKOM JroHcxono MaccnB€l B KoMnnerce 6eneMHuroB soubr Pseudolioceras m'clintocki 6onsue roapcKr.rx sJreMeHToB(Nannobelus, Brachybelus, Catateuthis, Mesoteuthis, Holcobelus, Parahastifes), r"reneeBenuK yAensurrfi aec Pseudodicoelitinae (e ocuoexov Lenobelus viligaensis, Pseudodicoelites hibolitoides u Hastites). Sachsibelus. Bosprororo Tal(oe pa3nltqlre o6rscngerCoaepuenHo orcyrcrB)'ror cgr
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Meu[Baercs Muono o6urarenefi upu6pexubrx o6craHosor, xl{Burrx y 6eperoa 6stnullu( Toraa ocTpoBoB. floeaHeaaneucKl{e KoMnnercrr 6enelrnuToB HaNr ugBecfirbr no 6onee onpaHrrreHtroMy tucny paspesoB n orqacrrr noToMy cpaBxxrensHo 6eqsur (16 euaoe). Veyqeubr ourr roJrbKo Ha nepu{nprn Cr6upcroft unar@pvu. B npu6pe)KHbrx @ullax nomrocrblo rrcqesaror Nannobelinae, Passaloteuthis, Parahastites, Rhabdobelus, psl BnAoB Hastites, Sachsibelus, Lenobelus u Pseudodicoelites, erruHt{rrHbrMr{ BHAaMu npegcraBregbr Mesoteuthis (M.inornata), Holcobelus (H,grauis, H. hinasoui), Eornruof, yaercnsrf, Bec B KoMrneKce npuo6peralor Pseudodicoelitinae u Hastitinae. Oauur orKpbr?oro Mopc s flpraepxocucKoM rrponrroe oxapaKTepl{soBaHbl npesMyuIecrBeHHo Pseudodicoelitinae u Hastitinae, Hapsay c KoropbrMl{ Bcrpeqarorcrr B uenou pa3nnqurr eauHurrtrbrcOrthobelus gigantoides, Holcobelus hinasovi. KoMrrneKcos 6eneMuuroB e @uunx MeJrKoBoAHblKu orKpbrroro Mops B nosaneNr aaJIeHe HeBennKu. Parare6afioccxnfi Bospacr caN{brx MonoAbrx cJroeB c Pseudodicoelites, Hastites u Sachsibelus a pa6ote aBTopoB 1970 n. 6urn nocraBneH noA coMHeHue. O!'. HaKo sa trocneaxxe noAbr B psae pafiouoa CenepHofi Cn6upu 6urru,r saftaeur,t BM€cT€ c 6enelcrnraMu I{ panue6afioccKle aMMou}rrbr. B Hueax Htrxxeno 6afioca B cnosx c Tugurites fastigatus KoMnneKc 6eneuxuToB craxoBl{Tcg eute 6onee o6eaxenubrM no cpaBHeHulo c aaneHoM. B npu6pexHulx o6crauoaKax Ha nepn@prn Cu6upcrofi nnar@pMbr BcrperraJrlrcb Pseudodicoelites 128
platyuentriostr's' Ps. hibolitoides, uatiformis' H' motortschunensis.
sachsibelus mirns, s. noaicius, Hastites claa"yrpra nJr"*o.o nporu6a a @urax oT*pbrroI-o lvlops npucyrcrBylor no cyuecrBy Te )Ke BI{abI, TaKlrM o6paeour, B gro BpeMg oKoHqaren'Ho crl'tpalorcrl pagntrql{fl MexAy KoMnJIeKcavu 6eneMHI{ToB a npH6pextHbIx I'I y[aneHHblx or 6epera qacrslx Mopa. B 6oaee BbrcoKr{x nopl{go'Tax H}r)KHe_ ro 6afioca B cJrosx c htormannites u My'tiloceramtts rucifer n"pr6"puu cu6rrpcrofi nnarfupr"lu o6uranu rIo-nPe)KHeMy "u re x(e Br{Abr pseudod,icoelites u sac/rsdbelus, 6onee peAKr,rMH craHoBrrrcs H astites (H.cravatiformis). HaquHas c BepxoB Hl'rxHero 6aftoca r,r Ao cepeAl{Hbr Hl.tx'eno 6ara MbI Be oHBa eNI B ceeepuoi'i cr't6upra r ua AanbHeM Bocroxe HaxoAoK aMMoHnToB (ecnu se '-"or"pi""r_ ctrl'IraTb He narcu'x rotrHono onpe'erreHrs Bos_pacra phyll;;;;atina), Ho I{HbIM'craHoBI'ITcs KoMnnexc 6eneNtHI'tToB. B orgocrtoo * BepxgeMy 6afiocy cJlofix c Mytiloceramus porrectus Hs Bcex oTMeqaBurl{xcrl B 6onee HItgKItx nopu3oHTax BlraoB CoxpaHsrcTcsrronbro Masot euthis inornatau tro AaHHbrMB.n. Itrua_ gOg) (f coBa na Ceaepo-Bocroxe CCCp Holcobelus hiiaiiui. Ha q.aery pseudo_ dicoelitinae u Hastitinae, rocnoAcr'oBaBrur{M B aane'e r{ paHHeM 6afioce, npru(oAsT npeAcraBaT€nl{ HoBoro sHAeMutrHono poAa Paramegateuthis: p.parabaiosicus' P' ishmensis" P, nesc-ia" Sror *otnra*" 6eaevnutoB cyuecrBoBaJl B paftouax Moprr, HenocpeacrBeHno npnnenapur{x K Cn6upcxoft naar@pMe, Bgyrpu flpuaepxogHcKoro nponu6a, B Mopsx cenepo-BocroKa u ,ll,ansuero Bocroxa cccp. KoporxopocTpoBble P'parabaiosicus onpa'nqesbr, B cBoeM pacxpocrparre'nu MeJrKoBoaHbIMI{ saur'gvu e6a'su 6eperoa rcpcKofo MaTepHKa, a o6naa'Rlrr'e nan6onee A,JrLrHHbrMr.r pocTpaMr{ p.nes ci a yKasbrBabTce c ceeepo_BocToKA cccp. He ua6rnollaercfl I{sMeHeHI{fie ergoBoM cocTarre Paramegateuthis, B r{r{saJ( 6arcxoro qpyca, B cronx c Mvtiroceromus hystatymensrs. H" ;;;;;;J*i"ilu"Te Jlrub HaxoaKI{ Mesoteuthis n Holcobelus. B aepxax *n*"""o 6ara s so'e Boreibcephalites pseudoborealis nogsnserc,, Hossrft eu.,,paramegateuthis_ p.monifesta. CyurecreeHHo MeHsercrr KoMrrnexc 6eaevurroB B cpeABeM 6are _ B 3oH€ Cra_ nocephalites vulgaris. Hcqeearcr xapa*TepHbre ,.Jrs Bepxseno 6aftoca _ Hrrx'ef.ro 6ara Pararnegateuthis parabajosicusroaMeurapuu{ecs eue 6onee ropor*opocrpr BbIMr.rP,pressa. flocnelrrtle, KaK r{ p.parabajoslcus, uprypoqenbr r{crJlloq[T€nbno x MenKoBo.aHbrMdaunsu. Bnepnsre a Cu6zpu norrBnsrorcrr B eroft soge [pe[cra_ Bl{reru{cylindroteuthidae: pachyteuthis (pachyteuthis) optima p, (p,) parens, P.(P.) bodylevshii, Cylind"roteuthis lCytindroteuthis) spathi, C. (C.) oweni oueni. B nepxxev 6are B 3oHe Arctocepiralites rpeAcTaBrrrerru ,cylindroteu"ilgun" KoMrrrreKca
6eaev'rroBro Konrqecr'y ""%HffiJ::;f:: llifi: T"""'IrHbI po,inii";;;;"li.,h;;;i'i*i'""ut",Hff,""Ti,
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/essa" OaHonperraerrHona nepn$e puu Cu6upcxofi nnar$opr'rtr o6onauaercJr cocraB, Paramegateuthis 3a cqer HoBOno B,n!' r p.timanensis, vraea'Ee ,,a HaxoAKr{ Pachyteuthis (Pachyteuthis) panderiana B BepxHeM 6are, cgena'Hoe ,,a oc'o_ Banr{r{ npeABaprTeJrbHbrx orrpeaenegafi a craTbe B.H. Caxc" o op. (Carc, Aanuc I{ AP'r L97 2) ' npl{ AeranrHofi o6pa6otx" t"t"p""* He uoaTBepAlnocb. B Bepxtrev 6are upa6pexxbte MenKoBoAHbIe oaurr pac[pocTpa'srorcs rr Ha B'yrpe'Hle sacru rlescxoro nponr6a. Teu se Me*ee ra)Ke e Axa6apcKoM pafioHe 6rrmr o6Hapyxe'bl pocrpbl Pseudodicoelites sp., nrButuMovy o6nrarenefi oT.r(pbrrono Mops, norrrll He 3axoll{Btut{x B npa6pexnbre BoAbr. c Haqana KeJrrroBes roc,,o'crBo B 6enevnaroBoM KoMrrnexce nepexoarr r cylindroteuthidae, cpeAu KoTopbrx nosBnsrorcs HoBbre poAbr rr noApoA u: Lagoniberus (Holcobe loide s Acroteuthis (Microbe lus)" Cy lindiotrutn;"--ltrirtoteuthis ), (communicobelus)' ), C. BI{.q'bIrorpa'uqe'Hbre B cBoeM pacnpocTpaae'rx rorlb.o HU)KHr{MKeffroBeeu -Ectr pachyteuthis ( pachyteuthis f ) tschemyschewil unn ser.Hrm_ Barou'e cBoe c)ruecTBoBa'r{e B oro BpeMs _ Cyhjndroteutiis (Cyt;na-teuthis) conf essa. B nnxnefi @He HrDKHero KeJrrroBes eule BcTpeqarorcs, xors x pearorpa_ ranegateuthis (P. ishmensis, p. manif esta, p. timensii, p.nescias. B sosy Cadoce_ ras elatmae sror poa ]rx(e He [epexoAlrr.
9 4 9
129
O cTpaTl{rpa$lrveCXOv BCe CXaCaHHOenO3BOnseTCAenaTb CneAyroull{e BbIBOAbI pacnpocrpaHeHl{uKoN,{nneKcoBpaHHe-HcpeAHeIopcKI,IX6eIlet'Ittuton. HuxHeno ToaBepxu BepxHero nnr{Hc6axa u soHa I)act,vlioceras tenuicostatum liTVs' ecrl nPeApa. ('la';tttteuthis' , a . s s al o t e u t l t i s , B r a ( l t y b c l u s I I . c r a B r { T e n u C a t a i e t t t l t i s ,O r t l t o b e l t t s P B 3oHe BOBCe' HeT B @uranx oTKpbrToro MOpg B Bepxax nnugc6axa 6eneuuHToB C a t a t e u t h i s ' D a c t y l i o c e r a s t e n u i c o s t a t t l m e A u H H r { H b IHea x o I l K } I B npu6pexHblx @uunx 6onarutfi KoMnHraxHuir Toap. 3ona Harpoceras falcifer. i s , r v c t t rot b e l u s u D a c t y l o t e u l t lesoteuth P o a r r M e s o t e t t t l t i s ^ neKc c npeo6nanaHlreM c npeo6natraHgeM l/us,nORnngloTcg BnepBble. B $aUunx oTKpLIToI-o Mops KoMnneKC l+lesoteuthis. Acrocoelites u catatetilhis, ecTb TaKxe oqexl] 6oratstft KoMII3oHa [Jacty I i oceras commu ne. B npu6pexgblx $aUusx H Apyrgx KopoTKopocTpoBblx neKC c rOcnogcTsbv Narlttobe/tt.s, Clastoteuthis L e n o b e l t t s ,P s e t t d o d i c o e l i t e s " P a r a h a s t i t e s ' B s a n p e A c r a B H T e n u n e p a r r e sopv. Acrocoelites u CataUuqx oTKpblToto Mopq 6onarulft KOMnneKc c upeo6naqalll{eM t e u t l t i s , n o s B J r r r r o r c rr re p B b r e l l o l c o b e l u s , H e r P s e u d o d i c o e l i t i n a e u I l a s t i t i d a e B n p u 6 p e ) K H b@l xu u a x 6 o r a t u f i braunianus. 3oua Zigodactylites BI{AOB Lenobelus, KOJII{qeCTBO BO3paCTaeT KOMnneKc, 6ruextrft K npetrbrayueMy, psettdodicoelites, parahastiles. B Saunax orKpbrror'o Mopq KoMnneKCH€ OTJrlf{aeTcrr oT npeablAyueFo. @3oxaCollinamucronata.B{auusxoTKpblroroMopgKoMnIeKc 6nusox K npetrbrAyueMy, Ho c HoBbrMlrBr{aauu Passaloteuthis,Mesoteuthis,Holcobelus. r o s e n k r a n t z i . B @ u u n x o r K p b l T o n oM o p s B l e p B b l en o 3oua pseudolioceras pocrpaMu' sgnrrKyrcs Lenobelus,Paralustites,ucveeatgt Acrocoelites co cxarblMu c 6oxos Beplnuft toap - suxxufi aaneH. B npu6pexHblx @uusx 6orarutft KoMnJIeKc' 6nnsxuft K HHXHeroapcKr{M,Ho c HoBbrMI{BI4aaMu Nannobelus, Parahastites u HoBbrM poaoM Hastites. B @uunx orKpblTol'o Mopn FocnoacrBytor Hastites, Pseud o d i c o el i t e s , L e n o b el u s . B upu6pe)KHblx mtclintocki. H u x n u f i a a n e n ,3 o s a P s e u d o l i o c e r a s HolcobeHastites, Lenobelus, BItaoB pqAoM HoBbIx $auunx 6orarurfi KoMrrneKcc lus u.HoBbMr{ poAaMr{ Sachsibelus, Rhabdobelus? He nepexoanr H3 Toapa Dactyloteuthis. B @uuax orKpbrroro Mopn 6orarufi KoMnneKc c npeo6naaaHl{eM trl{cne c BlepBble noHastites, Sachsibelus, Lenobelus, Pseudodicoelites, B ToM
B upu6Pexuux @tuguren si s. . E g p x g u af ia n e E3 o s a T u g u r i t e s uHsx o6eqnenuulfixovnneKc c npeo6naqasuevo6uHx c Sauunvu orKpblTol'o MoPs Pseudodic oelit e s ,H a s ti te s , Sa c h s i b e l u s . E c rs raK )K e Orthobe l us, hl esoteuthi s, H olcobelus, Leno be l u s . H cJloll c Normannif astigatus H n x q r r f 6t a f t o c .C n o u c T u g u r i t e s c Pseudodicoelites, r o v n n e x c o 6 e a H e n n t t f i Cunrso tes u lr*tlocera.s. u f i l e s o t e u t h i s " p e A K I { M H H o l c o b e l u s Hastites, Sachsibelus, Bepxu Hllxgeno 6afioca - sepxHrfi 6afioc. Kor'anrleKc c P aramegateutn npu6pexublx @uunx u c P.nescia a @uunx orKpblrono hi; i;iib;i;;nus *r Holcobelus. peAKt{MI{ Mesoteuthis Mopc, A l s t a t y . a , e n s d s . K o r " t n t 1 e K ccb l Hlrxsnfi6ar, CJroI{ c illytiloceramus H e r Mesoteuthis u Holcobelus' poro^tsotruthit parabajosicus n P.nescia r p s e u d o b o r e a l i s . K o v n n e x cc P a r a m e g a t e u a l i t e s 3ona Boreio""ph tr P.manifesta. this parabajosicus Kounnexcc Parames. tes C p e a n u f i - 6 a r .3 o H a C r a n o c e p h a l i _vulgari p. pressa u repBbrMu Pachyteuthis u Cylindroteuthis" satm-mffista, Kovunerc c Poramegaelega!s. B e p x n u f i6 a T . 3 o u a A r c t o c e p h a l i t e s C y l i n d r o t a u t l r d ss n p . s p p . , teuthk timinensis, P.pressa, Pachyteuthis K o v nnexc c Pachyteuk o c h i . A r c t i c o c e r a s Huxgufi renaoeefi.3osa Lr Acro(Holcobeloides) Lagonibelus nepBbIMI{ spp., ,ht" .ppJ}'lrndroteuthis H ap.). ( P . t i m a n e n s i s u r o c n e t r H l { M uP a t a m e g a t e u t h i s t e u t h i s ( M i c r o b e l u s) C y l i n d r oteuB u a a M I { ) K e c r e M u K o v n n e x c elatmae. 3osa Cadocera€ P a r a m e g a t e u t h i s . thidae, Ho 6ee
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poSa Moxer 6bITb ctryqaftrblM I{ nogToMy Her yBepeHHocTI{, oT'octrcg-Jlf, AarF uurft pocrp x Paramegateuthis (rev 6onee, qro Hesc'o, NaK upeAcraB[TeJrb paramegateutltis vor us ApxruqecKono 6acceftsa nonacrE B Mope Ha *,ne pyccxofi pan'r'rusr). Pocrp baiosicus MoxHo onpe'eJr["b u KaK Nannoberus, MOr oH rrp[_ HaAnexaTs r 'osofi oco6u Megateuthis (npr ycnoB'a BToplrrr'ono ,,ctres'o'e'us rrpr{Bepu[HnbD( 6opoea ). B panuev 6are Ha ceBepe cu6rpu nosBrrserce €ue oAllu rrn1 paramegateutiis - P'manifest',no crerleHl{ y''t{He'Hocrr pocrpa rrpoMexcyroqrrfr Me)KAyp.nescia u P'ishmensis' B cpeaaepr 6ate o6uapyxraaercq eure oAIrH B,.A - p.pressa, xapa*Tepus)nouuftc'I pesK,'M coKpaule'reM pocrpa I{ Bq3Mo)KHo aeayrunf, cBoe Har rrano oT P'parabaiosicus' cessr*arb BI{A pressa c. 6eneugo[AesMr{, IrMeKrurfMIt crrrrb'o peAyur{poBa'Hbre pocrpbr ( Belemnoteuthidae), ,u, ;;;;""x ocxosa'rft He BI{A!{M. lonJrg3n-Mo npeArronoxeEue, qro u HeKoTopbre Apyn[e onrcaHHble B rrr_ Teparype BHA'I Belemnoteuthidae MoryT 6rtrr' re'errqec*r cBssa'br c pas*rqHbIMI{ rpylmaMu Passaloteuthaceae, I{ ruc o6reqrsegue B oAlro ceveficrso gBrrf,eTcs I{cKyccTBeHHbrM. B noqnrreu 6me llosBfincs cpaBHlfFenbHo KoporKop@TpoaHfi p.timanensr.s, B pamrnfi xenaoaeli nepexolsr 4 srga (P.isheminsis, p.timanensis, p.manifesta a P'nescia)r HO B Ken,,oBee o'r.r yxe cra'oBsrcs pear*nau u Bbrue 30HbI Arcticoceras kochi, Paramegateuthis soo6ue He [oflrrrMarorcs, Becb orqr poa ueJlf,xoM BbrMrrpaer x KoHUy BpeMeH[ Arcticoceras kochi. Poq Holcobelu's, KaK yxe y*asblBa'ocb, nosB,,ncc a MopsJ( Eopeanuroro ,,o*ca B cepearHe paHHeno Toapa I{ ronbxo B aaJrere nepecerurrcs B npeaenrr Te T[qecKono nosca. lleparrfi l{sBecrublft narra npeAcraB[r\errb Holcobelus _ Il. hina_ souio6naaaa pocrpoM' c)I(arbIM e cmo,xo-6p*,u,,roM Ha[paBJre,.EE ,r noarorvfy BeposrHee Bceno 6rrn cessa' ne'err{trec*u c Acrocoerites ,,g rp][mbr unisur _ catus' BosMox'o A'hedonensts. Aaaercrcrft H.gr:avis c aepere'oBr{aHbrM pocrpoM Bnomre uon 6rtrs noToMKoM erof,-xe JIErruu pi""r"r. Haqaao sarraru{oeaponefr. CKI{M cpeAHebpcKlM Holcobelus, B rrepB}rto oqepeAb H.munieri, cropee Bceno AaJI nosBusruuftcq B rrosAneM Toape H,umarensis co cxqTbrv c Goroa cy6xo_ HI{qecKI{Mpocrpou' cav H'imarensis Mor [por{cxoAr{Tb urMf,tryg H.hinasovi, c HI'M ro &opve pocrpa rP€Acrasrrreaefi Acrocoelites (AJriscissus ;t;:i.*blx Poa Holcobelus paHbue HepeAKo o6teanxgnr c poloM Cylindrotetthis. Anmpu eule B 1964 n. ''o*aSaJ'nr' ,,ro re'eruqecrofi cBgslf Me)KAJr artrMlr ADy!\,rg po'aMr 6rtts He Moxer. rlepaue cylindrotertr,iJr"-roLa"rq B o,rrurrse or Holcobelus pocTpaM[ co cua60 paseurofi nprBeputfiuroft 6prounrofi 60posaof, f,ru Aaxe ,ruuerrr HbIMrr ee' Taxoart 6afioccrue Pachyteuthis liachyteuthis) inden. (carc u Hanrxeeaa, 1966) , cyrind,roteuthis themis,6arcrre c . ( c . )"p.iou. spath;, p.(p.) ?a_ rens v !p. B 1964 n. aBTopbr crrlrTaJr[ Bcnea ga B.A. f]crovecoBbtM nan6onee Beposr'blM rlpeaKoM Cylindroteuthidae ana b,eviformis voiu, io Harrgrrr coapd MeHHbrM npeacTaBnexlrsM oTsocsuutr* poAy B.rach"tbelr". bor.Ko, ecrlr y{ecrb cy6urmnraprqecK)ro pocrpa Oopr"ry Cylind;;;thidae *""_I"*bq c"qarruc pas_ Blrrl{g' To Kaxercs 6onee ocHoBareJIbHbIM ". BuAerb B groM ceN.refictae [oToMKoB Passafoteuthinae,cKopee Bcero pona Dactyloteuthis, aorqrB;;;; sarraagoeBpc. neficrnx Mopqx Ao aane',a. cneayet [pt{ groM rroM'rrrb, ..ro " Hetrompble ,,peAF craBl{r€mt Dacty loteuthis ,nMerl{ Ha pocrpe,csoficr""rrryro Cyiiiorotun,hinae, rpb Bepuruxrryro 6prcrur1m 6opoeay. Kar r rpeAcraBJrgnocb aetopav paHbue, nepBbrMr nQb BI{AuMoMy' [osBt{Jrucv Pachyteuthis -ue -cylindroteuthidae, ir.1r.;'"p. Iou. inden.1l or Hrx oTAeruJrrcb cylindroteuthis' eue co cpaBHlrr€Jlr'rro KopoTrcnMnpocrpaMn lc.themis, c.(c.) spathi)' Yxe s rosarreM 6are vu HaxoAnM cyrindrotiuthis"-""rroo yAruneuHbrMn pocrpaMr [c.(c,) confessa]*o-p"o,'rro"*.fi ,ope Aamr Hasarro ,,epc. ceruraueftcs B Eaporry rpymle spicularii - obelisca - porrecta. oco6ono BHI{MaErs 3acJlyxl{Baer pseudodicobli Borrpoc o t i n a e' rlogsae'ne B Toape srono 6eccnopno Srnoneunn ann Eopearrb'oro rlosca no'ceMeftcrBa s'AMenyer co60ro [osBJre'ue "ro.rrtr'oro rf HoBOno HAA.eMeficrga 6€b
f35
qTo Pseudodicoelitinae 6rp - Duvaliaceae. EcrecrseHHo AonycTllTb' rreMHoraAeft noqca' Mopfi Teruqecxono 3ACenr{BlxHX ,rr{ ftpeft€}4r+ B€€x ApyEI{X -Duvalia<-eaer u Cnlsenca (Ste(lloger' 1952) Poxe yKa3aHuq Hvetorcg Harrr{Harrc 6afioca. e aaneHe I{ Aa)Ke B vens, 1973) na HaxoAKIr B 3aUaAHOfiEnpone Belemnopsis nuTepaType aBTr e Belentnopsis Toape. HO rax KaK onl{caggq ToapraoJIeIICKI4x t{ro xe. KacaeTcg [poBepKe' B gTlt Hy)KtraloTcq yKasaHl{g DaM HerrsBecTHbr, TO K&KI4x-To npeAFseudodicoelitinae, To oHId, HaAo llpeAnonararb' npol{3oulm{ or u IlastiPassaloteuthidae Koropbtx Cylindroteuthaceae, ceMeficrBa ";";;;efi lope. pauueft tidae rocnoAcrBoBaJrl{ B rAe )Ke [MeHIIo I{cKaTb HeBce xe ocTaloTcrl HetrcHocTl{ B Bollpoce o ToM' ro o6uefi dopue pocTcya]ITL, np"AKOB Pseudodicoelitinae.Ecnu Trocpe[cTr.*oo B IIepBJnooeellastitidae' x 6nusKu orleHb pa, To Pseudodicoelitinae necoMHeHHo npe4nonaran B'A'fycpeAb K pOAy Hastites. HvenHO TaKI{e I'eHeTI'Ir{eCKI{eCBq3It MH.HIII. Bcex (igOA), KoTopbrft poA Lenobelus, o6teArHqB.ul.ft no ero rovecoo .p""ua"aicoelitinae, qTo cquTaTb' Haao g Ilastitidae. ceveficrgo Aaxe noMeluan sce xompr'1ft (Jeletzky, 1966), TaKoro )Ke MHeHr{s npr{trep)K'BaeTc' EneUrufi neHeruqecKl'I cBnsblBaer (=Belemnopseina Jeletzky) Duvoliaceae HagceueficrBo npeAnofloAo6Hure B3I'JIgqbI BbIcKa3bIBaIII{B L967 r' u aBTopbt' c llastitidae. q e p e 3 D u v aliinae' c H I { x u P s e u d o d i c o e l i t i n a e ' c J r a n a B u r { ec B r r s b H a s t i t e s K a K B e l e m n o p s i d a e ' 1 4 TaK I{ c Dicoelitidae npI{3HaKoM' otleHb II3M€HOanaxo MbI SHaeM' rrro Soprtaa pocrpoB tlBrgercg tIl{BbIMI{npuroAHbtMAngpa3AeneHI{'IJII4IxbHI{SIII,IXTaKcoHoB-H€BbluepoAa.
Nannobelinae u 6onr'urrscTBe B n o A c e v " n " t t u * N { e g a t e u t h i n a e ,P a s s a l o t e u t h i n a e , pa3nuqarcuuecs no ooprure' cneAyer Apyrux Mbr HaxoAIrM pocrpbl, or{eHb CI{nbHo nepBylo oqepeAb ycnoBI{'MI{ )KI{3H' B qro 6opMa pocrpa onpeAensnacb oyl"to, KoHBepneHTHbIMrlpu3Haxov' K nploKa3blBa[acb )KI{BorHoro }r nooroMy saqacr}rlo BepeTeHoBI{aIIbIe pocTpbl H as Mepy Mox'o yKasaTr, Ha BecbMa cxoAHble no topve ' Pseudodicoelites' Hit i t e s , S a c h s i b e l u s , P a r a h a s t i t e s ,H o l c o h e l u s , L e n o b e l u s , I',I aaxe HaIF ceNaeficrsaM pa3HbIM noAceueficTBaM' npugagne)Kauue K bolites, cepteftclBav. ycTaHoBJIeHUgsunoreueTllqecKHx EeccnopHo 6onee c)rulecTBeHHoe sHaqeHI'Ie ans O6yCnOB[eHHbIe' KaK pocrpon' IIoBepxHoCTI{ Ha Caseefi A9IDKHbII{MeTb 6OpOeaur 'racT€fi Tena xI{BOTBOno' MgrKI{x CTpoeHuq oco6euHocTqMu IIaM IIpeAcTaBnSIeTcSr, y pocrpoB poAa Hastites [Merorcrr TonbKo cna6rte 6oxoerte nonocbl' __v pocrpoB Ho Bce ecTb' 6oroerle IEI6o 6prougble' rm6o Apll1.ux poAoB Hastitidae 6oposart Y nocPseudodicoelitinae' pocTpoB y 6opoeAbr Ha oEu coBepueHHo He noxo)Ktr qactU nepeAHeft neAgru(, KaK u y Bcex Duvaliaceae, xa6r1oAaloTcq 6Opoeau.B Cylindroteuthaceae' V popocTpa, He uMeroruue ce6e aganoroB y llpeAcTaeuteneft cTopoHe pocTpa' 6plotuttoft tta 6opoena [pt{cyTcTByeT O" Lenobelus, KpoMe Toro' U IIOSAHeIopCKI{X 6OpOSAa, eCnU Or6pOCU'rb Taxaa natrtlnarcttlaficg o" t"pttnrl. CypaHnelopcxux Bcex I{3 ecrl) Oxyteuthidae' n pagueMenotoo Cytintroi"u't'iau" po'a l{Meere'' eroro B cocrase Holcobelus. poAa y roJrbKo iindrot.uthaceae pocTpoM' HaTIoMI{HaIouIItMpocTpbl i *"tg1n, BI{A - H,gravis c BepeTeHoBUAHbIM i Pseudodicoelitinae. grauis Henbsrr cqrrrarb HenocpeAcrBeHHbIMnpeAKoM Pseudor Teu He Megee H. -SI,IA-uieecreg TonbKo I'I3 aaJleqa tl cxoAen tro $opue pocrpa , {'roro*o eror di"o"litinae. ' nceBAoAl{uoeJrI{TI{H' c noeA'eroapcKr{Mrr I{ aaneHcKuvra npeacraBlrrenqMl{ MbI sgaeM JIlIlxb Pseudodicoelitinaer nepBble panHeM Toape, KofAa 11ofIBJUIloTcs ,B e cnuHHo-SppuHoM pocTpoM cxaTbtM c H.hinasoui poAa Holcobelus oA!trr Br{A nonBflsloutl{Ml{cg n nepxuefi HarrpaBnesru. oror. npndHax c6nuxbef H . hinasoui c Lenobelus uiligaensis braunianus so'e HrDKHenoroapa - 3oHe Zugodactylites I{3BecTHbIM B paHHeM eAI'IHcTBeHsbIM c 6oposart yTpaTbI 6prcurHofi i u ,rpl, ycnoBul{ Tpynno cKa3aTb' Ps 'primoris' poqa Pseudodicoelites Toape TIpeACTaBuTeneM Holcobe lus pocrpbl H66konsxo Mox'o He[ocpeAcroenHo cBrr3arb renerl{qecKl{ H3B€CTpOCTpbI nepBO1.O ll HanpaBJteHUIr' .,kinasout, CXatrte B Cm{HHO-6prcruUOM BpeMtr Dactyy)Ke Bo uonouBuerOcs L.lenensis-, popa Lenobelus rinono HaM BIIAa ' He gCXrUoqeHotqTo lioceras commune, ToHK11e'cTpeItoBHAHbIe'Cx€Tbte C 6OXOS' 136
npeAKaMI{ L. lenensis 6srru, Apyr-re npeAcraBureJr}r pona H olcobe lus, 6mlsKne K H, umarensis (eror BuA rroKa r{aBecr€H TonbKo u3 BepxHero Toapa HH)KHenO aarreHa). B.A. lycroMecoB (t962, L972) npuaaer oco6o Ba)Ksoe sHaqeHr{e B cncreMar[Ke 6oxogb[M nonocaM Ha pocrpax. 3ro nono)KeHl{e Aocrarorlno y6eanrenuHo IIM ne Morl{Bl{pyercg l{. BogMoxHo poJrb 6oxoertx nonoc allrr ycraxoBreglrrt regeTI|qecKItx csg3efi Mex[y pocrpaMu B.A, lycroMecoBbrM nepeoqeuuBaercs. Teu H-e MeHee I'I B pacflono)KeHl{It napnbu( 6oxosrx noJroc y Megateuthinae u Pseudocicoelitina€ Mu lre yBulI{M cJnqesrBegxofi pasur{ubr, KoTopas 6rr upennrcrBoBana npeA[oJIoxeHI{IO O UaflI{q]rll Me)KAy Ha3BaHHbrMr{noAceveficraav}r neHeTu,recxoft CBg3IT.
Boarrutrrto ponb rtpu BoccTanoBneuuu {unoreHe3a Aon)KeH l{rparb ogToreHeg pocrpoB. HccneaosaHns pocrpoB Pseudodicoelitinae u, oco6epHo, poAa Lenobelus noKa3blBarorr tITo Ha HaqaJrbublx craAr{sx pagBHTI{s xl{Bornbre o6naqa.ru pocrpaM[r orHobntensno 6onee yKopoqeHnbrMr{ qeM y R3pocnbrx oco6efi, cy6uunnHp pnrrecxoft uJtrt cnenra cy6xonuqecxoft pocrpbr Ha Haqanr,Hbrx Sopurr. V llastitidae craal{sx pasBI{TI{s cl{nbHo yalrDrHeHr{rre,cy6u[nrnAplrqecKne uJrl,l cnepKa Beper€HoBll[HbIe. Iloetouty I{ Ha ocHoBe oHToneHe3a c 6dnuuruv octroB€rHlreM Mo)KHo rrpoBoAt{Tb SunoreHerr{trecKr{e cBesn Me)KAy Pseudodicbelitinae n lffegateuthinae, o6JraAaBuI{MI{ HA HaqarlbHblx craal{sr( pocrpaMu cy6xoHu.recxoft sopvror, c MaJrbrMr{ snaqegr{svn [Ia. flepexons K paccJ\,torpeHulo SunoreHeruqecKux caseeft Bnyrplr uoaceveficrBa Pseudodicoelitinae (pnc. 43l, cneqyer cKasaTr,, vro Qunorexerr{qecKas cxeMa, npeanoJloxeHHafl B.A. fycroMecoBblM B 1966 F.r H€ B[oJrHe onpagAbrgaercsr.
Puc. 43. Cxerraa npeAnoJranaeMoro pasBr{THg pOAOB r{ BHAoB noAceueficrBa Pseudodicoe li tinae
il$ t?
, rd 0
HasBarrlrsrfi nccneAoBarenb rrpeAnonanar, qro pasBlrrr4e poAa Lenobelu.s, B Koroprtfi oH BKJIbqan r Pseudodicoelites, uro oT pocTpoB c Amugoft 6prourxoft 60_ poeaofi ( L.lenensis)'repes pocrpbl c ropoTKofi rpraepurrHofi 6proruHofi 6oposnoft ( L.minaeuae) K pocrpaM, Jn{ueHHbrMBoBce 6opoea s saqgeft qacru ( pseudodicaelites bidgievi). B aeficrs[TenbHocru yxe B paHneM roape Baprrgy c Lenobelus, I|MeBuuMu pocrpbt c AJII{HHrrvu 5prouubrMrd6opoeAaruu ( L,lenensis, L.viligaensis), cyilrecrBogarrr{r Pseudodicoelites (ps.primoris), pocrpbr KoTopbD( 6uml mruexbr 6pnuuofi 6opogArL B nogAHeM Toape napaJurenbrro norrBJr*. rorcrl BtrAbr Lenobelus c pocrpaM[, r{MeBrrrrfMrrAJuIH}ryb 6proffiyro'6oposny (t.s;_ biricus), u c pocrpaMn c roporxofi 6prcuHoft 6opoeaot ( t,rogt, L.minaevae)ra TaKx(e BI{abI Fseudodicoelites, y Koropbrx 6proruxaa 6oposau co*p.Hsrnacb ronEF Ko B uepenneft sacru pocrpa ( p". bidgieui, ps.platyaentriosui)i.'""
t37
MoxHo llonycrr{Tb, rrro Bce HasBaHHbIeBIUbI Lenobelus (L. sibiricus, L.vagt, L,m,inaeuae),fipousounn oT L.lenensis. Bce oTI{ Bl{AbI Ha llaqaJlbltblK cTaAtrgr( uMenu pocrpbr oar{HaKoBbreno Sopr're. Jlrrut pocrpbl L.minaeaae tr l.pagg orrJrr{lraru{cb 6onee cna6uv pasBl{Tueu 6plorrryoft 6oposAu. B AalbHeitueM Arr${epenr uuarr14s una 11o $ttnr4u pasflrlqHofi crenerm e3Ayrl{s pocrpa - cna6oro y L.lenenu L.minaevae ,r cn;1lgoro y L.uagt srs, yuepeHgono I L.Sibiricus t{ro racaercs HaquHas c IoHbIx craAr|fir pocrpbt cxaF L.uiligaensisr}rueBuleno, Tbre B cm{Hxo-6ppuruoM HarrpaBrrexrr{, To oH BogMo)KHo nplfiraAnexa[ K caMr BerBlt. Taxoe npeAnono)Ketrne rrpttBoAtrr r BbIBoAy o crosrelrlrttoft nenetr.Iecxofi 11poucxo)Kqen]Iu poAa Lenobelus, qTO OTnIoAb HeJIbss t{CKJIoqATb' nOJrI{Or{Jrr{qecKoM 6ener*xrroBr oCoco6enno ecntl 1n{ecrb ucrcyccrBexrrocrb B uenoM xnaccu$nxau[f, HoBaHHofi TonbKo Ha pocrpax. B poae Pseudodicoelites ro)Ke rpyArro c yBepennocrblo rlpoBo[rlrb renerr&nosrxeroapqecKue cBnslr MexAy rrorrBr{BrurMcrr B paEBeM Toape Ps,primorisr Kar yxe yl€3btPs.platyuentriosus). crstMr{-aaneHcKr{Mr{Br,taaMr{(Ps.bidgievi, B€lrrocb, y rrepBoro r.r3 HasBaEHbrK BUAOB Ha pOCTpe HeT 6pnrUnOfi 6opoeau, y nosgrreroapcKr{x Br{AoB ecrr, B rrepeAHefi .ractu pocrpa Ha 6ppuluofi cropone 6oposAa r{rrr{ ynnoureuue. lloerouy He I{cKJIroqeHo,rrro Ps.bidgieui u Ps. platyventriosr.rs neHeruqecKu cBrrsagbr He c Ps.primoris, a c Lenobelus lenensis. Ps.primoris upu raKoM aonyuenl{rr npeAcraBJrsJl 6sI oco6lm reuer[trecKylo BerBb. noserBrluecrr B aaJrene caMbre no3Anue Br{Abr Pseudodicoelites (ps.hibotitoides, Ps. clauatoides, Ps. gusf omesovi) c pocrpaMu, JltrurenubtMu6pnrunofi 6opoeAbr, cKopee Bceno rrBJrrnorcrrnpsMbrMl{ noToMKaMunoS[HeToapctuxPseudod,icoelites, B nepByro or{epeAb Ps.platyuentriosus. Y pocrpoB sroro BrAa, aoxlrBueno ao paxuero 6afioca, B npouecce eBoJrrour.rxHcqesno yn[ouleHl{e xa 6ptouruofi ctopoHe. AansHef,uas Ar{@penrrnauuc noura no Jrr{Hur{cna6ono R3ayrus pocrpa ( p". hiboIitoides), 6onee crJrbuono ero BsAyn{q ( Ps. clouatoides)uwr uao6opor npuo6peTeHDn pocrpoM cy6unnunAplrqecKoft {opr.,rsr ( Ps . gustomesovi). I-locneAHHfiaonpoc, KoTopbrft Henuss o6ofiru npu paccMorpeul{u Pseudodicoelitinae, oTo - Bonpoc o feHerr{qecKr{x cBrr3sx ux c Duvaliinae u ApynI{Mn ceveficrBaMr{ Duvaliaceae. Bpna nI{ Moxno Aonycrl{Tb He[ocpeacrBeHHoe rlpoucxoxregue Duvaliinae or no3AHeroapcKHx aaneHcKr{x r{ panHe6aftoccKrx curlbHo cueul{aJrtlslrpoBaHHbD(BIIAoB Lenobelus u Pseudodicoelites, o6aaAaaulrx BecI'Ma yAJrnHeHHbIMexAy reM, nepBbre Duvaliinae, noqBJuuourecs a 6are r Mr{ ToHKr{MHpocrpavr. KeJrJroBee (Rhopaloteuthis, Conobe lus), o6naaarm cpaBrrnTenbHo KoporxrrMlr cy6ur{nl{Hnpr.rqecKuMur.r cy6xoHnr{ecKuMu pocrpaMl{. flo"arorry rrpeAr
[a a Aoxolguas y poAa Belemnopsis ao ''p,nBepurxnoft sacrr{. Taxas cKyJrbnrypa pocTpa To)t€ Monna sBurbcs pe3ynbraToM peAyKrrn[ cmlruroft 6opostrr Ha pocrpax poAa Lenobelus' Moxso [PeAllonan{rs la negeTuqecKyrocBssb ceveftcra Dicoelitidae u Belemnopsidae c poaoM pseud.odicoelites, c TeM1neno Br{AaMr{,Koropbre uMeJrr{B nepeA'eft.racru pocrpa 6proru'yro 60posay, xoT,, u ocna6neH*yro (ps.bid_ g i e v i , P s . p l a t y u e n t r i o s u sj . flprneaenHsle coo6paxenl{s He l{crJlroqaror BosMoxuocrl{ nporcxo)KAe',ns Dicoelitidae u l3elemnopsidae oT KaKIrxrnr6o rpyrm cylinclroteuthaceae, 6r*s Moxer Aa)Ke ot o6qero c Duvaliinae [peAKa. Moxcro Aorrycrr{Tb BcneA 3a l.g. KprunoJrblreM (1958) neuetuvecKlm cB'sb pona Belemnopsis, a 3a Hr{M a Apyr[x rpyrm Duvaliaceae c poqoM Holcobelus. pocrprr Belemnop.srs,KaK x pocrpbr Holcobelus, I{MetoJ o6rurft rrpnsuaK - AJuTHH}rb 6proruryro 6opoeay. B gfiJr.rss 16_ xono aonyueHr{s Pseudodicoelitinae or(a3aJrr{cs 6rr r{soJrr{poBanxbrM6oxosnrv ot_ BerBJreHI{eM Belemnoidea, ne ocTaBI{Brur{M noroMKoB flocne cpeareft ropbr. rlocKonbKy uMeercrr pra o6ruux npI{sHaKOB B CTpOeHrr{ pocrpoB, o6rea,nuflroufix . ir I r. r. rseuooolcoelttlnae c ApyrI{MI{ npeAcTaBI{TeJUrMr{ Duvaliaceae, nogo6Hoe AonyueHI{e npeacraBJrsercsl Meuee BeporrrHbrv, Cxopee Moxgo fionaraTbr HTo p.seudodi_ coelitinae s 6afioce rrJru Aa)Ke aaJreHe Aaru{ Haqano BceM Apynr{M f)uvaliaceae, nepecennBuJl{Mcg B Moprr Tern.recxono norlca.
OE YCJIOBI{flX ?IIUSHN MEG.A.TEUTIIINAE II PSEUDODICOELITINAE
Belemnoidea sBnsrorcs BbrMepuefi.npynnofi xuBc/rHblx, B l{cronaeMoM cocrosHl{u cJlfraeB @Taefrcs JrHltll' qacTb BHlFIpeHHeno cl(€.crn H11xB rrogaBngrorUeM 6ongursctBe nqTa - pocrpbr, no rcnppbrM TplIAHo laxe B@c-TaHoBlrTI' o5tuufi o6nnr xl{BctrHor\o Ir reu 6onee cyAlrrb o6 ycnoanf,x eno xrrgun. Orne.rarru MctKnx qaclefi Terla 6eneltHo"' fl@roMy MoxHo nl{urb AocreroqHo }rBeHlef, BcTpeqarorcs B e[fiHurrxbx cnfrasn. Berru rroABrlxpelrgo cqrrt&Tb, rrro rro[odno coBpeMeHHbIM xaJrbMapau 6elevglrrbl aBTopoB o6 ycnonusr( xuoHs n uufi o6pae )KugHr{. 6onee noApoSHo rrpe[craBnexnc gKggronuil 6ener,runroB, Bo MHor\oM npeAnoJroxlrrenbubler [pI{BeAeHbt B pa6one 197Or. B @HoBHoM cornacy€rrcs c Marepnan no Megateuthinae n Pseudodicoelitinae pagee cAenagnbrMr{ BbrBoAar.an.Oopr.,rbl c roporruMt{ pocrpaMl{ ( Acrocoelites ianen s c h i , M e s o t e u t h i s b r e v i r o s l r i s , M . s u b r o s t r i fo r m i s , M . t i u n g e n s i s , P a r a m e g a t e u t h i s parabajosicus, P.timanensis n Ap.) UenI{KOMl{n}t noqTn UenrKoM npH}PoqeHbt K npu6pexxblM MenKoBoAHbM o6craxosKaM Ha nepu$epun PpcKono MaTepuKa u Mox(er pa6o6ulr AarKe Benu nprrogxblfi o6pae xusrtu noAo5uo onucarrHbru B rpeArlgyuefi MbI He B soHe npu6pexxof, a Moptl. rny6nHbt onpeaenulr Nannobelinae. Vaepexno te cocTotrttnn, HO, eCJII{ CyAHTb nO [pIdSHaKaM paSMbIBOB H ganuql{rc cpaBHI{Ten-bFo Kpynugsepxl{srblx ocqAKoB, eu nny6nuu 6ulnu ne 6onee 5O-1OO v. noxaeanu' qro H no xuMuqecKoMy Hccnetoaagus .|'|.E. Kqrgnoeofi H Ap. (1973) cocraBypocTpblNannobe|inaeornnqalgl€gnoBb|[ueHHbIMcqqepxaHHeMx(eJle3a' TnTaHa, MapnaHua, KpeMHr{g u anroMl{Hns - npOqyKToB Bbltt@a c 6nnsnexauefi c}run. y Nannobe[inae no cpaBxenlrro c [nI{HHopocrpoBbIMI{ Passaloterrthinae 6onee G/t{erJII{BOBbrpaxeHbr TeMHbte u cBgfJIbIe KOHUeHTpHqecKuecJIoI{ B pocTpo(r oTpaxapuHe cesogHbre HsMeHeHns ycnonufi pocaa, 6onee pegro nposBnarorufiecs eSnnsu cyuru. Bce sro cnlaKrrr cBHAsrenbcrBoM Toro, qro KoporKopocrpooule $opt"ttl 6enevnHToB Hatlro xqnrrJrlr Hau6onee 6naronpnnrxyto oSctaHosKy ans ce6g e npn6pexxofi ooHe. Kacaercs [peAnojroxexr{s o roM, rrro raKue $opvul, ocTaBascb KaK H Bce 6enevHr{Tbr ceo6onHorulaBaroluuMn, Belrr{ fip}rAoHHblfi o6pae xugHLIr To 3To octasrcs nHub flpeAnonox(eHr{eM. B nonrey rono, qro ycnoBnrr xHgHH KoporKopocrpoBblx H anHHHopocrponulx @pv 6eneusmoB paoJrr{traJrr{cbMe)xAy co6otor roBopsr Haptuy c pa3nuq11cMn e Sauuansuofi rrpnypoqeuxocru H yKaganxbte pagJllrtlus B xHMHqecKoM cocTaBe pocrpoB lr reMneparypuoM pexrMe cpeabr o6uraxuq (noapo6xee cv.Hroxe). notftH UenI{KOMnpHMXOrue AJrHHHopOCTpoBbre Qopr"rsrOKagblBalotCc UenHKOM HnH ypoqeubr x o6craxoBKaM orKpbrroro Mopc, nae BenH, HaAo AyMarb ceo6oaHolln8garctl1gfto6pae xrrsnlr. B ux qncne MoxHo HagBarb ^lcrocoelite.s ltcrgrccili.s, rl..strbt e n u i s , . { . t r i s r r / c o s u s , ! + l e . s o t e u t h i sp e r g r a t t d i s , n o q r u B c e B } t J l b tH o l c o b c l u s u P s e u d o d i c o e l i t i n a e, P a r a n t e g a l e u t n i sn e s c i a , C y l i n d r o t c u t h i s c o n f r s s a . H a n p u v e p e lnrlHHopocrpoBbrx Passaloteuthinae, xl{MrqecKuft coctaB pocrpoB Kc/ropblx H3francg, BUAHO,IITO nO cpaBHeHr{lo C KopoTKopocTpoBrn'ar Qopr.aaM}ryAnI{HeHHble pocTpbl o6eA8eHbr npuMecgMu, IIpHHocHBurrrMHctrc c)nxH, MeHee qsrKo BbIpaKeHbI B HHx cnon. fny6nxrr e o6nacrr orKpblToro Mopfl 6ulan, cyaH cegoHHbte KoHrreHTpr{qecKr{e cocraBy ocaAKoB, He MeHee 1OO-2OO r'r ( e nnnxlrcroMy no nfieur.,ryuecrBeHno pafioxe Ovonogcxofo MaccHBa Bo3MoxHo H MeHbue), Ho Mbl He 3HaeM, cnycKaJlllcb nn 6enevHuTbt B HIDKHHecnou Boabl. t40
HapqAy c BIuaMH, o6naAasuII{MI{ yAnr{Heuurrvu cy6xoHr{lrecrr,rMu r{ cy6uraruqpnqecKI{MH poqrpaMHr e o6nacTl{ crrrpblToro Mopc x}rJrr{ 6enevnurbr c BepeTegoBtuHbrMu I'I Aaxe 6ynaaosrAlrblMl{ pocrpaMr ( Pseudodicoelitinae, Hastitidae)- Bax 6prcuurov Ha[paBJresnr pocTp' caoficrBe'Hbr BrrAaM, o6nraaurM B ,,pn'or'br yc_ JIoBIlsx' Pocrp, cxatufi c 6oroo, xan 6laro Aeficrsmenblro, AoJrxeH o6nenq&Tb xnBqrHoMy ynpaBflsrb cBoI{M TenoM npl| rurasarruu B rcpusopranuroft 11rrgg*ocT]r. Oqnaro norITH Bce nepeqilcnentrble Bbrue BuAbr co cil(erbrMlr B clluuno6ppgo,ou HATIpABJIeHI{gpocTpaMl{ IlefluroM I|JIrt noilITl{ [eJrnroM SaceJlsrrn oSnac-ru crTrpbrr,ono Mops. Aaxe ropqrKopoctpoaufi A,omolonezsis He Bcnpeqetr Ha Merrro'e'bsx rra ' nepr@prn PpcKoro MareprraBosl\doxno, pocTpbr, c)r(arbre s cnruo,o-6p,orusoM Harrpa'aeHHHt cJIJD|(uJIl|ongpof, 6oroarn"r turaBnrrau, cnoco6cDBoBABIur{M uorp]rrKegr4o Hrrrr BCtuIbrTI{IO xt{Borgono. flpencraane'l{e o ToMr irro He*orppbre ,,p,,.nrar,., paorrnqa,oufiecs B Tex ,,JI'. t{IIbIx rpyflnaJ( pOCrpOBt CBsgaIIbl C nOJIOBbIMupaantrqrrsMu, OqtraeTqs 6eqqOrA3a,TeJrbHbIM' Bogil'toxHor trro [o'oBble ,,pr{o,,arfi BoBce He .,'paxalrr{cb na Qopue f,o}qaeMbtx HaMIl p@TpoB. C ap]tror cTopotrbl pocflpbl caMuoB I{ caMoK Morrru Eac,roJltKo cI{JrbHo crntlf,tlaTrcs Ap}tr or [p]rrar ryro MbI (vrlloCtrM IDr r pagHbIM BlrIaM ltJrtr Aa)Ke poAaM. llo raclosUteno BpeMeHu He BbrsBJresbr Tarlre npusnal(r{ Ha pog,rp{x, roropbre Moxxo 6uno 6rl csecrrr K no[oBoMy AuMopOHsMy. n*rotr,ty rrpru(qnrcs BoqAepxt{B&Tbcs crr ycranoBneHlts noJroBono nrvop$noMa y 6elevuurog. Ha Hasa,lblrblx qraarp( pocTpbl Megateuthinae u Ha caMbrx HaqaJrbnbrn ..roA,,m pocrpbl Pseudodicoelitinae 6unn cpaB'm€m'o ropsrKr{Mu cy6rox^recK[Mr{ r{ cKopee Bcerc' no asaJloruu co Bgp@ttblMl{ ropotropoerpoBblMr o6nranu a {opuaun, npn6pexHsx o6cranosKax. ero rloArBepx(qaercs n oco6ennocrf,Mrr ,(f,M'qec'org cocraBa loHbrx pocrpoB, orrurlarouerocs, Kar rroKasang JI.E.KosnoBa f, Ap. (fgZC), or BSpOC'ISX OCO6efi noBbrrrreaHbrM coAep)KaHreM nprHocf,Mbrx c cyu'I npnr,recef,
141
(xeneao, Mapraxeu, Turau rr Ap.). Pocrpul Pseudodicoelitinae oqenb 6ucrpo, Aaxe npH MaJIbIx AnaMeTpO(, STaHOBHnI{Cbcy6Uunru{Ap}rqeCKHMUyAnr{HeHHbIMI{I{ )l(HBcvrubre, cyArr no r{Merour{Mcs Ho(oAKaM TaKTDCpocTpoB, nepecensn}rcb e o6nacrr orKpbrrono Mops. Tonaro npu npn6nr.Dxeul,ruK pagMepaM Bgpocnono xlrBcrruorro, pocTpbr y orAenbHbrx Br{Aoa npuo6peraJrn 6ynaaoeuaxyro nru.r pesKo BepereHoBI&HyIo 0opt"ry. Taxosl.r I.enobelus uagt, L.minaeuae, L. sibiricus, Pseudodicoelites clauatoides, Ps. bidgieai, a raKxe Holcobelus gzauisue Megateuthinae. Eonruoft I{HTepec [peAsraBJrrrror B@cTaBneur{e TeMrrepar)rpubrx ;rcnoaufi, B KoTopblx xnnu om{cBlaaevue 6eneMHrrrbr, r{ rgMeHeHI{s grl{x ycnoeufi Bo BpeMeHu u npocrpaxcTBe. Meroaura nanegreMnep&TypHbrx o[peAenennfi, BbrnoJrHgeMbrxI{a.ocHoBe ycTaxoBneH'rr coAep)Kaur{tr B poc"rpalc rrDKenor-o lrgcrrolra o18, MarHug vr cTpoHul{s, npnBoanrcq B cneu}raJrb}rrx pa6orax (lopu u Ap.r 1954; Eepnrx n Xa6aroa, 1966; EoysH, 1969; Caxc, AHurnHa r{ Ap.r L972) n vu na neft qnecb ocTanaBnuBarbcs He 6yaeur. K HacroflqeMy BpeMeHI{ npoBeAegbl y)Ke B Aograroiltro 6onguroM KoJrl{qeqtBe onpeAeneHr{s cpeAgerqAoBbrx nanecfi€Mnepar]rp vopcxofi Belbr no pocTpaM pagtrer.r cpeAHeropcrru( 6eaeuxuroB c reppnropuu Cn6upu (EepnrH r{ Ap.r 197O; Caxc, Asnrnxa ll [p.r t97 2). O6uee KoJrntreqrBo raKru( onpe[enennfi qocrtafaer 4OO. ,[na onpeAeneuuq ucnoJlbsoBalrbl B ocHoBHoM pocTpbl Nannobelinae u Passaloteuthinae no roapy u pocTpbl Cylindroteuthidae no 6ary. flpn neyreuuu narr€c[€Mrrepar]T aaJIeHa6ttnu npuBneqeHblpocrpbl Pseudodicoelites, Lenobelus Ir Ilasf ites, c6,urFnruro( s o6nacrr{ orlxpbrrono Mops.- _ flo coorsouexr{ro HoronoB KrcnopoAa iol879161 orpeaenexurr rpoBeAunncb Ha orpaurqeHrtoM KoJrr{trecrBe pocrpoB (9 - pagHss npa, 3 - cpetise rcpa). Pocrpu, Bssrble LIs MenroBq[Hbrx 0auufipanHefi rcpu (rogeu rurnrc6ana - Toap) e 6onunrgqtBe noKasanu qpeoMepHo BbrcoKue reMneparlpbr (35-5Oo). B pa6ore 1970 r. aBTOpbI 6unu cKJroHHbr ryraBHylo npuqruy TaKI{x oTKnoHeHI{fi oa BoSMoxHbrx pea'Ibp HbIX TeMnepAfyp MopcKruc BoA ncKaTb B r{SMeHeHHsx coJreHocrl{ B MeJTKoBQ[HbD[ gaJr}rBbr.OAnaro, or}r I{oMeH€Bq[oeM$(, KaKI{Mr{ 6rnu Bunpficruft u Xaranrcrnfi Hul tre Mornu 6urg BenI{KH, n@KoJrbKy B gaJruBax xr{Jrl{ TaKI{e creuoraruxxble QopMbI, KaK roJrooononr{e (peaxue aMMonrrrbr r{ oqenb o6unsnrre n pasxoo6paexue 6enevnurrt). fl@:roMy 6onee BeposrHo ocHoBrryp [pFrr{Hy ngMeHeHufi coorHoueHHs rlgcnonoB KllcnopoAa B pocrpal( BllAerb B 9nr{reHe[IGtecKID( npoueccat( - a o5t'{enHbIx pearuurD( c rpynroBblMu BqaMr{. Ilocnenuue, Kqnarl{ cxao&Tb, HMeJru B HuxHelopcKxx crroR:( rr@ne cyrnoxenus ro[us MoulxocTbrl IIe Menee 1-2 na ocaAKoB cpenxefi H BepxHefi roprr u Mera reMneper)rpbr He HruKe 3O-5Oo (ncneactBr{e AeficrBurr reorrepMr{qecKoro fpaaneHTa). Bqbr ec.r€c-rBeHHo cBo6qHee Ilyrmoeue Irl{prynl{poBanu B MenroBqnnux 6onee KpymrosepHr{qtbrx [opqaJ(, Hexenu B nnr{Har(, ornoxuBluruccg B @KpbIToM Mope. H aeficrsrreJrbno, pegro gnnblureHuble gHaqenl{g naneoreMneparp Aarru poqrpbr, [pu]poqenxbre K MenKoBqHbrM Saunev ( AcrocoeIites, Passaloteuthis, Catateuthis, Nannobelus). onpeaeneHurr rrarrecrreMnep&Tlp rro oTHoruesulo caAg B tronl{lresfBe oKorro 2oo flerouy Aa.nu e o6uev se6onsurofi pae6poc enaqexufi (Eepnrx r{ Ap.r 1970). ponb BTopHtrHbIx l{gMenegnfi s colepxaHl{r{ Mannr{tr B pocrpax oKac}brBaer,tcg Heguaql{T€rrbHofi u vrr BnpaBe noJrbgoBarrcs cpeauuMr{ gHaqenugMr{ Ans ycTalroBnennc naJleoreMfiepaTyp. Eue MeHbIu]rMIr, gBrrsx]rcg oTKnolreHvc. B naJreoTeMnep&T]T$(, orpetrensBuuxcc tro cqlep)r(aurro B pocTpal( crpoHuus (Caxc, Annrrna H Ap. , L97 21. Tarux orpeuenenufi erlnoflgexo 185. Oqeus Huexufi xo${ruuern Bapuaunr{ cogep)KaHIrs cTponul{s B pocTpo( qAuoro Bospaera H Hs q[Horo rurr r{s 6nngKrlx Meg.roHa)cor(AeHuft HapaAy c usMeHeHusMlr B coflepxasr{H srpoHrlr{s B pocTpa)c pasHono Bo.3paqra Id ug pasHblx pafioHoe, AaeT @HoBaHr{e crrHocrrTbcs K Tar(r{M o[peAeneHU'IM C AOBepUeM. HeoSxoguMo onoBoplrtbcs, tITo Mbr He MoxeM c yBeperlnocrbn olrexlrrb BooMoxHbIe usMeHeHHc Ha IIpoTf,xeHHu K)pcKoro nepHoAa B c@TaBe x coAepxaHrfrr pacTBopeHHbIx coneft B MopcKru( BQ[ax. Me)t<.qy reM, Tarr{e HgMeHeHr{n 6eccuopHg ^crnpaxaJrucb Ha coAepxaHr{I{ B Bolle, a cneAoBsrenb}ro H B poeTpa)( Klrcnopola 0.6, ManHI{s H erpoHuus. 3To oScrosreJrbcrBo gaqtaBnseT c ocTopoxHocTbK) noqxoAtrrb 142
Pnc. 44. He\,{eneHus cpeAgeroAo_ BbIx TeMneparyp Bolllr a r.aopnx Ce_ aepnoft Cn6upu B TeqeHHe panHe_ I{ cpeAHercpcxorl onox (cyvvapHrre Kpr,rBbreno AaHHbrM onpeAeaenlrfi e pocrpax 6enev'uroe
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HrDKe a6conprubrM ogaqegllqx [aJrecrreMrrepm]? MopcK[rx BoA. Henb_ 3g HcICJIptIaTb TOrO, qTo B pesynbTaTe [poBeAeHHbrx onpeAeneHufi Mbr non] IaeM ''e a6conlorHbler a rlIIIrL ctr'ocl{Terrbl{bre BenI{qI{Hbr ,,aJreoTeMflep€,"yp Mopc*ru( Bq[. Ho, ao-uepBbDt, arlt orHocI{TeJIbHbIe gHageHug Bpgn nu cI{JrbHo orJrr{rraJrl{cb or I{crI{H_ Hbf,rt ut Bo-BToPbrxt coorHolueHl{s reMrlepaTyp, noKagaHubre Ha puc. 44, ao BpeMeHr{ U HA IUIOUIAAU, H€IAO AYMATb, OKASbIBAPTCg PEAJIbHbIMI,I. Ang xorrua naunc6al(a r{MeeTcs^n}uub 7 onpeaenenrft no ornoruegr{K) Cai!N4g, AaBIure SHarleHlrs noptuKa 22-230 Ans Xaranrsrono saJrr{Ba r{ oKoJro 27ans gaJrlrBa. Maloe Bnapf,crono Korufqec,rBo [€urgbrx He nogBoJrgsT c yBepeugocrbK, )"rBeprqarbr trro no,I] rerlHble pesyJrbrarbr He gtsnsroTcs SaBbrrueHHbrMr{, no xpaftteft Mepe gne Bunrcficxoro ttt'n"". Buecre c TeM HeJrbos He yqr{TbrBarb I{ Tono, qTo Ko'eu nn nc6axa og'aMe'oBancsr BecbMa BooMox'o r{MeHHo B cBsg' c flo.€ilr€HI{eM MopcKID( BoAr IIPoH[rIcIoBeHueM B cn6rpcrue Mops ue 3anqanofi Eaponrr 6eTIEMHITTOB. flnn uepaoft nonoal{Hbl pa'H"lg Mbr pacnorraraeM rraJrecrreMnepaT}rpaMu, Jp.pa onpe[enelrHblMu no orrloueHl{K) Ca/Mg TonbKo ang Bnnrcficrono oarruBa. 7 onpeaeaenxf, llo*agannn SHaqeHI{s nopqtrxa 2r-22o. Bropae norroBl{Ha pa*Hero Toapa BpeMs Mar(cl{M€uasoro pacuBera 6enevuntoB B iu6upnn o*"p"ro.pr{soBarra uar6onrluI{M Ko'r[qec'rBoM onPeaenennfi naneorevqeparyp: rro MarnuIo - gg, no erporrurlp
1 26' Ho onpeaenesufi no or'orue'Hro 01 8/tti bt;;il;;;rr" rpercras*errbHbrM JII{IIIb qHo - tto pocTpy catateuthis ex gr. subinauditane Xmasr.croro ga'rr{Ba c TeMnepmypofi 2grgo. onpeaeneHns ctr''oue'',,s caNsIIaJII{ e[r{Ha*oBbre s'aqe'r,rs [€uecrreMnep€rr]p Anc Xera'ncKono n BunpficKono saJrr{BoB _ tropq'x" ,l_';;i;'r{ HecKoJrbKo MeHb_ IIIH€ - o*oJro 21r5ouo 2 o6paeuSM IIrs Mopc s o6aacru ovono'c*oro MaccuBa. onpeaengnl{cb rpH.?ToM pocTpbr Passaloteuit,in"", Nannobelinae, B Me'bureM KonlrtrecrBe Acrocoelites u Mesoteuthis, onpe4ene[rs [aJrectreMnep€rryP rIO erpoHtlrilo lloKasanr{ srraqesns uo 6o o6paerIsM [ns Bunpficroro oaJrr{Ba 24o u no bS oOp."rIaM Ans Xararrrerono s€urr{Ba 22,30' flpu eror'a oSou paftonax narreoreMrrepatrrrrpbl no I c Kopcrr*r{Mu @pvaM posrpaMrr ( Nannobelinae) o*aoanrcb Ha 2-2rsb HHxe, qeM ro Anr{rr'op@TpoBbrM Catateuthis. flpn.lrxbr erorp Hescubr, pauee Carc, Anuxrsa H Ap.r (Lg72) nuc_ Ka3€ulfi [peAnoJroxeHHe o roM, q19 Nannobelinae xu,rll y AHA, rle TeMrrepmy?a BQqbI 6stna Huxet a catateuthis o6nra.rru B BepxHr{J( [porpeBaeMbrx co,,ErIeM cJrorD(
BOAbf.
Ter'rneparJrpbl BoAbr a o6nacTu orKpbrroro Mops n Hpraepxosr{cgoM nporn6e u ua ouonolrc*oM MaccuBe 6amu Bo BTopoft nonoer'e palr'ero Toapa, KaK no*agbrBaIOr o4penene'us no cTpoHuHKr, Irr.DKe u coc'raBnsJrr{ o*oJro 2';"',1n;^1;;;; uau). B noqnHeM Toaper KonAa Boopocfla cTenenb sueMr{qnocru cn6rpc6}rJ( 6eneuHITI0B' TeMnepar]rpbl BoAbI B Mopflx Cu6upn cr€uru HecxorrbKo Hr{xe. flo onuoure'Hro
143
saJrr{Be yc.rauaBrluBalcrtcs reMrrePer]pbl rtopqAra L9-22o Ca/Mg a XaaHr€KoM (uo 11 o6paeuau). flo l{soronaM KUCnOpq[a eArurcrBeHsoe o[peAerles[€.rIaJIeof€Mnepan)Fbr B pocrpe Hastites sp. cocraButlo 18r7o. Onpeaenenl{sono Ca/Mge O6nacrn OvOnOUCTOnOMacC[Ba IIOKaganU SHAqeHus OKOnO2L-22-. Ha npmnr<eHr{r{ aafleHcKoro BeKa noc-reneHHo BbIMIIpapr npeAcTaBnTeII[r ToapcMopcKru( BQ[. rofi 6ene\anHrosoft $aynu E q[Iro^Bperuellro snnaKtf IIaJIeqI€M[epar]tpbI a/or 6 gartu x&TarrreKono aaJIeEe parrueM B Arts Or Onpeneneuus no orHorueur{p Hastites s.p.), nnn o6naclu ortcpblT'oro Mr oarruBa reM.nepar)rpy 16170 (a pocrpe sp). 1615o (a pocrpe Pseudodicoelites ps B flpuaepxotrrcgoM nporu6e r"rrrlpr"y-py esrb ua 24o^ roapcKlo( Ca/NlL Huxe Onpeqeneur{s no orruourexuro "-5*"t a parrHeM aaJrexe H Ilo 1615" B rIG?[o6paeurr c naJreoreMnep€nlpavn 15r4-17u HeM aaJrene. Hapq[y c sruMu uu$pavn uMeKvrcs n 6onee BblcoKlre SHarreEng tlafleolCpennue BeII1nqI{HbIAII' Men*oBqsofi o6nacT}t B Xananr: TeMnepeTyp Ao 22-2go. aarrene (no 18 onpeaenenusv) u 1?r5o B roqAcKoM sannBe o*--;od ";;.r HeM aaJrese (4 onpeaenexun). .['ns o6nacru srlcpblTono Mops a flpraepxoflIlcKoM BO[bl MaccnBa B paHHeM aaJleue reMnepery?bl nponu6e r{ B npeAeJro( Ouonoscxoro (7 o[peAenenufi). 6unu B cpeAHeM oKorlo 18o Onpeaenegus no c-rpoutl{rl Ans aaneua [poBoAt{Jrncb nl{ulb nng o5nacwr CrrKpblrloKaoafl$ no 30 o[peToro Mopg e llpueepxortrrcKoM nporu6e (p.Kenuvnp).^OHn qeM B ToM xe paloHe nnrce 19o, ua 2r5" AeneHHrrM cpenueloAoByro reMneparlpy pocrpbl Pseudodicoelites u u 6ynasoBI{AHbIe BeperreuoBltrsbre B Toape. IIpu erolr Hastites Aanlr sgaqegr.rrr nareoreMnep&T)? xa 10 Huxer tIeM cx&Tble B clluHlro6prcuruou Ha[paBJreuur{ pocTpbr Lenobelus uiligaensrs. BoeMoxnas npl|rll{Ha sToro pocrpaMu orrlcK&Tbcc u 6ynaaoBIUHbIMu cnoco6nocrr Sopv c BepereHoBrrAHbrMH ua 6onee sgaqrrt€nbgbre rny6uxu or rroBepxxocru. 3 paxueaanegcKrD( [peAcraBl{garl-uBar'c58,fl no coTenc Nannobelinae He Me;1KoBqlgbtx orno]KeHufi Xatanrcxono na 2-3o Bblue BqbI TeMII€pATy?ar( IIpI{ pocTpax, x-tulH B AepxaHr{ro c:rpoHllrs qeM B OTKpbITOMMOpe. B rosue paHHeFo 6af,oca BbrMuparoT notrtn ECe BIdlbI I{ poAbl TOAFaJIeflCKOnO KoMrureKca u Ha cMeHy HM llpru(qnr HoBas cHcTeMarl{qecKas npyn6eneunrroBoro K coxaneHr{ro, IreAocrarotlH@Tb MaTepI{aJIa Ao cl{}c nop He Paramegateuthis. na nosBofluna [poBecTr{ B HyxHoM KoJMqecrBe naneoreMneparypHble onpe[efleHl{s B pOCTp&\ 6afioccrnx 6eneuuuroB. V Hac HMelcncg nnurr- 2 onpeneneHl{s orlroueHutl tla/Ilg B pocrp&x 6afioccrnx Paramegateuthis sp. ne XaraxrcKoro oa[I{Ba' AaBurI{e sHaqeHns naJlesreMneparyp 18r3 u 1913-, T.e. TaKse xe KaK B aaneHe. B cpeaHev 6are B Mopcr( Cenepuofi Cu6npu HapgAy c Paramegateuthis nosBnflrgrrcs nepBble (,ylindroteuthidae. Enuncreel{Hoe uMeloueecs o[peAeJIeHI{e naJreo3ElJrIrTeMneparlrpbr B pocrpe Pachyteuthis optima ug MenKoBqxoro Xsrasrcxofo (la/llgsnatreHne 15r4o, T.e. Ha 3-5o Hrore qeM B aaJI€Ba AaJro no o/rgou"rro He Lr 6aftoce. prctp" Cy' Ang noenneno 6ara Mbr r{MeeM olno o[peAenexne no 018/016 " lindroteuthi.s spathi na llpuaepxosucKorp nporl{6a, noxaoaBulee narreoreMrlepm}Ty raKxe o1rpeAenenus no orHoureul{p []a,/]ilg gaauue B 2 poctpa;c Hueprcg l4r5o. C..spttrhiue flpuaepxofircKoro nporr6a nareoTeMnep€rTlpy-i5r5o, B pocTpa)( C' spathi u Pachl,teuthis optinta Hs MenKoBoAHoro XataHr-cxoto saJrI{Ba rlaneoTeMneparypy 17,6-17 rgo. llo ctponUglo naJleoTeMnepaTypbl OTIpeAens[I{cb s 29 pocTpg)( nOsAHe6aTcKru( Bonbl oKaSarlocb Pacltl'tetttltis spp. '3*o ns XaranrcKoro 3€urHBa.cpenHna TeMnep€rrypa roro xe Xaranr€Korc r" 2-3o HHxe rreM naneoreMneparlpbl paexofi 18rgo. SannBa B aaneHe. Ang uaqaaa KennoBeficxoro BeKa MbI pacnoJraraeM oAHIIM oIIpeAeneHHeM no Cll6 ue llpreepxorurcKoro npofu6a. flaneoreMneparypa oxaoaJracb paoHofi 018/ 1gr7o, Ha 50 BbtuJe qeM B nogAHeM 6ane (a auavurengxofi qacru 3To trolrxHo o6ueneHr{eM flpnaepxos}rcKoro nporn6a K Kouuy rlq?Axero 6ara). flao6rscus6cg neoreMneparypbr, onpeaeneHubre no clrHouJeHHIoOa/\lg B pocrpa( paxxeKennoeeilcsl). l{3 MenKogoaxono Xaragr.croro SaJrI{Ba(18r1-19r1o), KHx Irararnt,gatcutlti.s Toxe y1asbrBaror Ha 3aMeruoe noBblrueHne reMnepar)tp BoAbl a cx6upcKHjx MoptD( c 144
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ocxoerr na.neo6noreorpa{r.rec*orp pafioxrpo'a*r{s pa'He- H cpeA'eKrpcKrDr Mopefi ocaeuregbt aBTopaMr{ B cnettualrrsoft pa6ore (Carc r Ap.r fgZf ). pafiorupo_ BaHI{e IIPOBeAeHOC ]rysroM pacnpocTparreHr{s Bcex saxxef;unx rpynn ncronaeuofi Sayxrr, uo n nepnyp oqepeAb 6aeupoeaJrocb sa cno6onno nJr{raarourx oFr.as,,sMax (ar"tlorumr, 6erer'onrrnr). Jlaa parueft rcprr rrplrcHaercs HaJrugf,e naJposoor\eonpafqecKux o6nacreft-Eopeanunofi aoxpyn [oJrroca, BKJtroqarouefi rvopn Cenepo^3anamofi Eeponu, n Teruqecrofi B Tponuqecrofi u cy6rponu.recrofi gonax. B Eopearnnol o6naqm Bblren'rlorcs [poBtrHuuu: Eopearuso-ArnanrlFrecras a Eapone s Aprrrrrecxa.s n cu6npu, Karaae u l.pexaanAmr..fus clpeoref, bpHr rraquxas c 6a^fioca, ycrruBuraccc KoHTpacrHocru $ayn no6yxaaer Hasa€ulxrre o6aacru rrepeBeqru B p€un noscos c o6nacTsMr{ (e pa6ore 1gZ1 n. [poBtrrun*r.,ru) BH]FEplr Bnx. B EopeanuroM noflce paoJlrrrraKtrcs o6lacru ApmrvecKas n Bopearnno-TrocoorearrcKaq. Mopn Ceaepo3anannofi Eaponar B cpeAnefi npe BbrrrrJrr{oa [peAenu Eopeanbnorp trosca. Kapru reorpa$mecxono pacrrpocrpasentrs Ea seMHoM [rape 6eaeMnnros n B f,x qxcne Megateuthinae u Pseudodicoetitinae, B pdurnefi rcpe r Baqare cpennel ppbr onldnxxoaagbr aBTopaMr{ B lgZO r. lr qAecb 6uno 6sr neqenecoo6paono ltx noBTOpgTb!
eaponeficrfix.rr,ropnx Megateuthinae nosBnfforca erqe B paHHeM neilace Welemnites (Acrocoelites) pro"Zo* Schwegle, Hrrroreno nErranra pxnol q6.'f, ". oPr, Mesoteuthis laeuis (Simpson) o Lpoo" Hrurarero nefiaca AnrrmJ. B.H. C[aBrrIfbIM (1963) I{3 BepxoB cuxeMlopa Kapnar JmasbrBarqncs passalotei;i" p"" dohastites. C sa'Iara nnrurc6axa a 3ananrofi Eapone o6nranr yr(e q€!,rbrpe pqa Megateuthinael Acrocoelites, M e soteuthis, Pseud,ohastites u Salpincoteuthis. Mag"r"uihin"l PacnpocTpaHspr.'cs u ua BocroK, oBr ecrb a Kapnararc f Mesoteuthis pseud,oclongata ue c6opoa A.C. Aannca, Pseudohasfites (Cnaarm, 1963)l,re Eonraplrn ( p"ruAohastites). flo lansrna |.fl. Kpunrnorrbtra (1965) u K.III. fiyqrO&.e (lgid6i;. Kasraoe s Ml'rHc6ar(e BcTpeqatorcs Mosoteuthis pseud,oelongdia (paxiliosa Hlngr6uaoe), Pseudohastites charmouthensis. Ha tor ; Eaponu=a nnsnc6axe 6eleruHrrrbr saKenlrrrl n s Ceaepuyro AQpnry. K xoHuy nnntc6axctcono BeKa Acrocoelites, nar6onee Bepor11go BoKp]rr Ceaep-. Hofi Esponbr s AenurcoBMesrno c Passaloteuthinae n Nannobelinae rpouf,[arrr E Mops' oMblBaK[lue Eepaerarcrnfi MsDep]rK c ceBepa rr ceBepo-BoqDora" [Ipn o1roy Bce usBeqrsbre HaM Hal(qAKr{ pocTpoB npr{ypoqegbr K MenroBqubrM npr6pexruu 0aunsM Ha nepn$epun 6uaruef, sa Mec-Te Cn6npcrof, nnatsopMbr cyun, a o6nagrrt c/rrpbrrorQ Mops Ha ceBepe Cn6rrpn 6enewnrbl B ruurucfaxe etqe He claxqrrrx. Toapcrnft Ber B 3anaquofi Eapone osnaMenoBarrcs pac[BqnoM M"grtuui|in"L, a rMeHHo Acrocoelites, Mesoteuthis u Sotpingoteuthii, Bo rropofr norroBrxe ro"p. Acrocoelites n Mesoteulhis coctaBlcru ocnoB[oe fltrpo 6enermnrorrx lrodmetrr@B B sartalxe r{ toxrroeBponeficnr:c Mopsx, our nocnqcrBypr B cosDaBe 6eae nrroB s Ha K)rGBOsroKe Eaponrr (a Eonnapuu, B Kapnarax, a Kprruy n Ha Kaaraoe). floEr rIeAHIoIo o6o6uauuyn pa6my no paunelopcrof, $ayne Kanraga Hanf,cara K.III. Hyu)6unse (1966). Erc rpuaqqesu orlrca'rs n noo6paxen.s pcna BxnoB Mcsoteuthis ua BepruIeno roapa KagKasa. Eonturag tractt poqrpoB, otrpegerreggux rar lfa. soteuthis, a raK)Ke pocTp, oTueoexrrrfr r poqy fulpingoteuthis, npruagIexrc Acrocli_
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Da(tyloteuthis, K lrlesoteuthi.s x1s (yrHocsrcs pocTpbl, onpeAe[eHHble KaK HastaHl|eM BeletroA Toapa AonSacca olincan ,i3 HII)KHero (19O8i A.A. E;ncttK p1'ramidalis' Ilesoteutlris x 6nusxyro rnttitrs att. cctrrtpr{'s.st/.s Sopvy, tll{cne }legateuthinaerpacnpocTpaVe eeponeyrcKr.rxMopefi 6enevHI,ITbI, LI B ToM H Ha Iono-B@ToK ao TypA{puxll Hsngcb B Toape Ha ror Ao CHuunuH H CeBepuofi ''Megateutlris cf giganyKasblBaercn nefiaca Bepxueno ur{r.rH Hpaxa. B Hpane Ho 1 8 9 O ) . ( Weithofer, tea' B Haqane roapcKoro BeKa e Cu6upn Bo BpeMl Dactyliot:cras tenuicostatunt 6enevHutbr lr B r{x rrucne Acrocoelires 3acensnl{ B ocHoBHoM Me[KoBoAHbIe )^racrpeAKI4 e' O6nacrl{ oTKKH B6nilsu 6eperoe EapaeuaacKoro MaTeplIKa u 6b'flu oqeHl' CCCP. Her AaHHbIx o cyueirabBaHul{ 6enevHnroe pbrroro Mops na Ceeepo-Bocroxe B gro BpeMs D MoptD( AansHero Bocrora. I KaK nogasaxo na puc.45, rovnqeKcbl 6enevBo apevn llarpoceras falcifer' u pa3HUTOBB MenKoBoAHbrx SOHa)( cn6Upcrofo Mops cAeflaflucr' 6Onee oonaluvu ne Enironut nepecenHBuluecs HrpaTb nOo6pasHblMu, cyulecTBesHylo ponb B HID( cTanl{ rocuoAcrCCCP qrKpbrroro Ceaepo$ocrore xa Mopn Mesoteuthis. B o6cranoBKa)c Bypurr{M KoMrroHeHToMB KoMIUreKce 6ener.aru{ToB6unu Acrocoelites. Bo BpeMs DacErue 6onee pasHoo6paexuruu c-raxoBrrcc KoMnrexcu 6eneMHr{ToB I{ a XaraHr€KoM o6ctanoBKa)c npu6pexxux B venrosoflHbrx tylioceras commune. tes.
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148
- BoHa Harpocerasfalcifer, Tt
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BunpftcroM SaJIHBa( HapqAy c nG-npexHeMy MHoroqHcneHHbrMHAt'rocoelites H Ilesoteuthis nonBnmsrcs flepBble Pseudodicoelitinae; Pseudodicoelites primoris u Lenobelus lenensis. B o6ctaHoBKax orKpbrroro Moprr ua Cenepo-Bocroxe CCCp Pseudodicoelitinae elue Herr, Ho saro BMeqre c npeo6narapurnMrl Ac.rot.oelites u 6onee peAKHMr.rMesoteuthis HafiaeHbrnepBbre Holcobelu.s (H. kinasot,i). flpu paccMcrrpeHr{r{Ha puc.45 xpuaulx r.r3MeHeHufieo BpeMeHH.B}uloBoro H pGAoBoro cocTaBa 6eneruuuroB r{ gnaqeHufi raa$$rauueHTa HsMeHeHH"t ,.r*"g He o6paTnTb BHUMaHT{eHa cyueqrBeHHoe cxo[cTBo Mex[y xoaoM yKaSaHHbrx KptrBbrx H KpI{BbIx tr€ilecrreMnepar]'p BoAbr, noKagangbrx Ha puc,44. Cranoenrcrr Aocraroqgo oqeBllllHoft ceces pacllBera 6eneunmoB B cu6upcrnx Mopfix B Toape H rrocren€HHofi aenpallaulrl'I n nepecrpofixu KoMrureKcoB Ha npo/rrDxeHurdcpeAHefi rcprr c Kone6annguu reMneparlpuono pexlrMa. Bo sper'an Zugodactylites braunianus pacrpocrpaneHue r,r cocraB 6ene\aHuroe MeHglcrrcg cpaBHIrDeJIbHoHeeHaqnTenbno. B soue MenKoBoAnfi y6ulnaer poJrb Acrocoelites H SaMerHo BogpacTaer, xcrrrr Bce eue HeBerrr{KoKornqeeTBo Pseudodicoelitinae, ubcBntrorcfr HoBbre HJ( Br{Abr. Ornoxennn Hagana nosAHeno roapa - soua Collina mucronata - yBepeuuo ycraHaBnl{Balorcfl ToJrbKo s o6nacTu OtuonoucKono MaccI{Ba, n $aunruc c/rKpbrroro Mopg. 3aecs coxpansercn B o6utev raxofi xe KaK r{ B Koxrre paunera Toapa KoMruregc 6enevxuroB c Acrocoelites, Mesoteuthis, Holcobelus, flocaeAsnfi poa oco6esuo xapaKrepes Ang vopefi Ha Cenepo-Bocrore CCCP. Bo apevs Pseudoliceras rosenkrantzi e o6nacrx Ovonbucxoro MaccnBa yr(e saMsrxo coKpauaKlrcs panueToapcKl{e oJIeMeHTbr, ucrlegalot Acrocoelites c,g cx&TbrM c 6oxos poqrpaMr{, garo nosBnsrorcs Pseudod icoe lit inae. Bo aropofi noloante panHeno.Toapa, Bo BpeMrr crrfloxeur{s crroeB c Dactylioceras, un:a B tto.3AHeMToape 6enewurbr H B ru( ql{cJre Acrocoelites, a Bo,.3MoxHo H Mesoteuthis, yxe 6eccnopuo paccenfiorcc B Mop$c Aansuero Bocroxa, B'nosAHeM Toape (6ener.aHnroBbre crou c Haugia aff . iaponica) oHu nponr{Karor ll s flnosup. B roape xe 6enevnurbr uruporo pactrposrpaxflsrcs B [pegenax Ceeepsofi Ar.aept{K}I - s Ceeepgofi u 3anaggofi Kanaae n Ha Anscxe. Megateuthinae crrcKrAa He onpeAensnl{cbr lro no yqrHoMycoo6utennp IO. Eneuroro Acrocoelites B KoMIr[eKce 6e-. neMHHToB Bsrpeqarcrrcc. Kpor',re Tor.o CI. Eneurufi (Jeletzky, 1966) rrpgBo[r,rr oqeBrulHo us BepxHeno roapa unu aaJreHa Ha)(olKr{ Lenobelus n Hibolites (cropee Bcero Pseudodicoelites). Cneayel fipeAnonararb, 'rlo Ha ceBepe Ceaepuofi Aruepurol, KaK H Ha Aaflbgerr,r Bocroxe B Toape 6rln pacnpocrparreH Tor xe KoMrrJrerc 6eneusutoB, qTo n s Ce.eepnofi Cn6upu. Kounaerc gror oacerrsJr Mops Apxtuvecxoft upoar{Hur{u. C oqHofi eropoHbl B HeM Haptuy c pqAoM o6unx c Enponofi sulos Acrocoelites 14 Mesoteuthis eue coxpalrsJll{cb enerraernu nnunc6a:ccrofi $ayurr 3anaauofi Eaponu, c [pyrofi npuc)rrgrBoBaJru tr B rrq3AHeM Toape ogaqprenbgo ycfinucb egAeMr{tltrbre pq[br ( Holcobelus, Parahastites) ) n aalr<e sgAeMnqrroe Harceveftcrao ( Duvaliaceae). Bo nropofi nonoauHe panHeropcrofi enoxu 6enevsurbr npogr{Karor r{ B pxnyro qacrb loxaofi Avepuru (Apnenrnra, t{unu). orcroga yrcasbrBaKrrcs Passaloteuthinae u Nannobelinae, xapaxTeplrbte AJrs eapoueficroro runrc6axa. t{To Kacasrcs Acrocoelites, Tor cyas no pa6ore Crrseuca (Stevens, 1965), B Hr{xgefi rope IOxno[ Arraepuru u)r Hert. Kar pr<e orrMerr{nn aBTopbr a ]-9?O r., Hau6o[ee Bepo*rnufi Iryrb MuDpaurl}r 6eneMgrroB B roxgytcl qacTb IOxHofi Auepuxra ue 3anaAnofi Eaponu npoxoAnn rlepeo Ceeepxyro A0puxy u Arnanrury. 3ro Monno 6rna npn ycJroBrl{r tITo B nepaofi noroBl{He lopcKoro flepr{ena s toxgofi qac-ru Arnanrnxu6sa cpaBHI{TenbHo Heuupoxuft u uenny6orufi solxrlfi 6accefui, xmopufi He Mor npentmgrBoBarb paccetleHulo csoficTseunblx MenroBoAHbrM yqaqrraM Mopq 6eneMusrog. HatrnHas c a€urenar B Mopfl( 3auaasofi Enponrr nosBJrrrKrrrr ,r"pror" Holcobe/zs, aepornrbre ncrroMrr{ ceBepocn6upcrux roapcruJ( Holcobelus. pqa BrtloB Hotcobelus [prcyrsrByer u B aarese Kagrasa (Kprnuronru, lg7g). Hapgnyc rrgMu -or'oueune asuux
cJrMMbI BblMepunx I{ BHoBb rrosBr{Brur{xcs BI{A.B K KoJrr{gecrBy nepexoBrraoB B IIporIeHTa)L
149
Pnc. 46. Cxeua reorpa$uqecKono pacnpocrpaueHng 6eaeunnloB B no3AHeaaJI€HcKoe BpeMs flonoxenre Marepr{KoB AaHo B coorBercrBun c xaptoft A.Cuutar A.Epafingna 1973 ); cyura noxasana KpanoM; npepblBucrbtMs r f.J\ruapn ( Smith and oth., o6nacrefir ToqeqHbIM rI]HKTITJrr{HlrgMn laHbr rpanuubr naaeo6noreorpar[uqectctx poM - rpaH[ubt naJreosooreonpa$urlecKrx rpoauxuufi. L-2 - Bopeaaunas naJleoo6lacrb (1 - Aprcrnvecxas npoBxHurnt 2 - Eopeantxo-Tnsooreorpa$uvecrag
acrocoelires, salpingoteuthis, npqoJrxafln cyueerBoBaTb tlesoteuthis, pexe Toape nosBuJrlrca s 3anaElofi EepopaxbuJe B eurc trmr AaKe Boelro)Ktro oAgoBpeMeruro ne nepBHe Duvaliaceae - pon Belemnopsis (Roger, 1952; Steven.s, 1973). B nqaaxeaanegcKoe BpeMs n Eapone paccens€t$cg n 6uclpo npno6psra€rr nnaBesrByKxrr)ro poJrb pa[ tlegateuthis, npeacraErtenH Kcrropono AoqtHrnn xan6ontlutu cpeln lJelemnoidea paovepoB (pocrpu HMetor anuuy go 45 cv). Hapf,Ay c Hr{Mtf o6urann Holcobelus u cTaHoBsru[ece pre peAKruu Jlesoteuthis u Acrocoelites (puc.46). B Aprcrnuecrofi npoBHnUsH B paHHeM aarreHe fipqnonxaru cyltlecrBoBarb orleHb 6oranue Br1gaMH.tcoMrureKcur 6eneuxuToB c Bogpocuefi no cpaBHex}rp c roaPoM qrenegb1g eHAeMHqHosrH.l Pseudodicoelitinae, Sachsihelus). flpu eror"r KoMIuIeKcbft ppcKorro u npn6pexxblx odcMenKoBcflHblx B M&TepuKa Ha nepusepnn o6rr1asuue @o6extro Muono roapcrl{rc Qopv H B ToM qncne $opv, 6nngTaHoBKax cqepxaJru Kor"tnneKcbt ort(pbtrono Mops 6unn Ktrr( nnu rox1qe6:1geggbrx ganaagoeBponeficKrv. gIIIleMnoMa. ctenexbKt Bbrcoxofi H orfluqaJrncb 6onee ceoeo6pagnbrMrr B nosareM aaJrexe cn6npcrue KoMrurexcsr 6eneMxnroB cyuecrBeEHo oSeAHclorcs (cu. pnc.45). B KoMrurerco( Aaxe B MenKoBoIlHbIx oScraHoeKar( nelocnqcrBo pexo1nT K sHAeMHKaM ( Pseudodicoelitinae, Sac'/tsibelus, eHAeMI{qHbleBuAbt Ifas-
150
rooreaficxas npoBHHtIHs); 3 - Or6rpcraa naaeofoioprrcrriecrac o6racrr;4-S-TeTnrlecras nareosooreorpa$nsecr€fl o6aacrs ( 4 _ ClenugevnoMopcxas flpoBrrr_ ttrll8r 5 - Hnno-TuxooxeancKas npoBnnryrg); 6 - Hn[o-Eeponeficras naaeo$no_ pgcTIFIecKas o6racrs. EyxaaMH noKagaHo pacqpocrpaHeHHe oTAeJrbHbrx poAoB 6eaeurarros: O - Orthobelus, Ms - Mesoteu-this,-Hm'- flomaloteuthis, A'- Acrococlitcs, l,ll- Megateuthis, Hl - nobobelus, Hs - Hasti-tes, R - Rhabdobelus, S - Sachsibelrs, L - Lenobelus, Ps - Pseudodicoelites, B - Belemnopsis.
tites ). Bce oro noBoplrr od )rxy[uegrfl ycnoaufi xfioHn, cKopee Bcero o nonrxe.9 Hxn reMnep&T]rp Be[br B Mope (cv. npeg.brnyruyp rnaay). 3acnytr
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B nosane6afi47. Cxeua neoFpaou.recroro pacnpocTpaHeHrls 6eneuunroB occKoe BpeMg A'Cr'4rta' A'Epafiheflonoxenue MaTepuKoB AaHo B cooTBeTcTBtlI{ c KapTofi ADofiHbIMu xpanoM, noKa'3aHa cyura 19?3); ua u l.Apr{BpH (smith and oth., TOIIKEME nOgC-oB' na;po6gqpeorpa$rvegKgx npaggqbl npepbrBrrcTbtMr{ rrri1u916u AaIbI llnOpao6nacrefi. npepbrBrcrbrMu JruHusMrr - rpanrrrlbr naJreo6[oreorpaib[qecxnx Mn noKasanbr: 1^3 - Bopeaaurorft nosc: ( 1 - AprrnvecKag naleosooneorpa0nrrecKarr o6nacrs; 2 - EopeanbHo-TuxooKeancKas naJreosooneonpaSurecxar o6-
pnc.
loxxyrl trac:rb IOxnOil AVepnnr flpenCranryteJll{ 3TOrc xe KOMIureKca IIpOHuKaJI}r B r { n n x , Holcobelus admirandus (Steinmann, a lMegateuthis gigantea ( M'6ricke, 189s) a Aprexrtnre (Stevens' 1965)]' 1 8 8 1 ) - e E o n u n u u , B e l e m n o p . s i s ,D i c o e l i t e s HerlotrHble (6ee aeprulEbl) om{can 1898) B Aprenrmre TopxKE:{sr (Tornquist, palrHeno Aorr€pa u B.corespinazitensisHs pocTpbr nqA HasBarrusMt{ Belemnit)s Belemdobaensis rs cpeAHero Aorr€pa. Cpequ'usoSparxeHllfi B.espinazitensis eGIb p a r a b o i o s i cus' g r ' ' e x P a r a m e g a t e u t h i s nopsis (raOn.1O, $nr.?) u- aoewoxno crtrelg65 s cxarufi (ra6n.1O, B.cordobaensis $6.8), (ra6n.1O, $nr.4, 6). Pocrp 6pnruxov HanpaBneuru, Moxer fipllHaanexarb Acrocoeliles ex gr. omolonensis. cucTeMaTr{qecKoe noJroxeHHe pocrpoB H3-3rl ID( Henorulc'fbl veepeuxo.ycTaHoBuTb poqF' HeBG3MoxHo. Bce x(e aacJr)rxr{Basr BHuMa}Ius Halcox(Aeuue B l0xsofi Avepure Ceee3a [peAeJlar'nr gTHocuTbcs HensBegurgMy poAyr no6a K Moryr poB, KctTopble pa H Aanasero Bocrora CCCP. ParaEcnu pocrpbr B.espinazitensis AeficrBurerlbno qacrrttlHo llprutaAJrexdt K qepeo CeaepHyO rrponuKartu Bceno cKopee Ar,aepuxy lOxnyo e megateuthis, To oHIr Avepnry, xcr1n e Ceeepnofi ArraepuKerroKa Paramegateuthis u He HafiAeHu' TOxe MO)KHSCKasaTb u O Acrocoelites ex gr. omolonensis, flpaaAa, aUA omolonensis Ho ronbKo B nnuxc6anre. Boo6rqe Acrocoelites c poc'regrb u e 3anaguofi Eapone, 152
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nacrb; 3 - Cu6npcKaJr naaeo$nopr{crilqecKas ofrracrs); - Tern.recrfl{fi +6 noac (4 - CperulseMHoMopcKas naaeosooreorpaQnqecKas o6aacrs, 5 - hHaoTnxooxeaHcKas naJleosooreorpaSnqecKas o6nacm, 6 - Hrrao-EaponertcKafl rraaeo{nopncrl{qecKag ofuiacrg). Eyrcnar"r}r noKasano pacnpocrpaHeHue orAenb}rbrx poAoB 6eaevnnron: Ms - Mesoteuthis, Hl - Holcobelus, lul- Megateuthis" pa - par a m e g q t e u t h i s r l ) 2 - B r a c h y b e l a s ,C - C y l i n d r o t e u t h i s , P - p a c h y t e u t h i s , L - L e nobelus, Ps - Pseudodicoelites, B - Belemnopsis, H - Hibolites, D - Dicoelitidae
paMl{r cxarblMu s crnxrHoSplotllHoM Ha[paBnegu]r, B 3anaanofi Enpone nocJre Toapa rrcqegarqr, a Cn6rpu corcparrsKrrcc ao KoHua aanega. r{to xe KacaeTcs Hailaeu_ trbtx- B cPensefi rope lOrcxofi Avepurn Belemnopsis, Megateuthis, Holcobelus, To o[In BeposrHee Bcer9 rIPnIuu cloAa no Eaponu n@o6no paggeKrpcrunvr 6eleMgrmaM rrepes Ceaepxyo AQpury u Arnanrrry. Moxuo Aytu€rrbr trro r cPeag_eft ppe, cropee Bceno r 6aftocy, 6rpr6,qg'rcfi rrsxe[KIr B Aprenrrue coBMeerxo c Belemnopsis u Dicoelites Braihybelus, yr(asbrBaevue CrnBeHcoM (Stevens, 1965). Oru Ha.lcqAKH ,ronrrqno cBsgbrBarb.Bo BpeMeHu c HaJcox[eHl{eM B cpgAHefi rope (aanen-6af,oc) Hogofi 3enaugrn om{cas'ono F.CrnBeHcoM Brachybelus zieteni Stevens (non Werner). Ilp[sqAeuuue $ar
t53
B cpeAne6arPnc. 48. Cxeva neorpasn'recxoro pacnpocTpaHeHns 6eaevrulToB cKoe BpeMs q Kaproft A.Cvnta, Il.Epafiaena flonoxesne Marepr{KoB AaHo B coorBercrBul{ l(panoM' xr{pnblMl{ npepblnoKa3aHa cy*a 19?3); n f.Jlpnopu (Smith and oth., nOgCoB' ToH!{I{MI{ naJreo6noreorpaouqecK[x rpaHl4tlbl BHCTbIMUJII{HHSMI{AaHbI o6aacrefi' IluQpallaJleosooreorpa$Hqecxlx FPaHI{tIbI J1{HI{SMI{ npepbrBgcTbrMu naJreosooreoFpa0n( ApmnvecKas 1 Eopeaaunrrfi nogc Mu rroKa,3aHbr: 1-g o6secKas o6aacG; 2 - EOpeaasno-TuxooKeaHcKas naJleosooreonpa{u'recrac
srorp pqAa B tclxuoM lronyuapru D cpeAuefi lope (e 6afioce ? ) flpencraaurern cnMetrerrbrTarxe sa ManaracKape ( M.cf. gigantea Lemoine, 1gOG) I{, MoxHo IlpenllonaraTb, lrrEpoKo pacllpocrpaHsnHcb B oKpaEtlHblx Mopfl( fonneaHb| HaptIAy c (SteBelemnop.sis. flocnepnue usBecrubr raroKe na 3eune fpeftar'ra a Auraprrule vens, 196?) I-lpueeaegnbte AarIHbIe saqraBllslor H s Hruolresuil (Stevens, 1973). 'cKJroHflTbcg K BblcKagaHHoMy l. CruaeHcoM npeAnoJroxernlo o roM, rfno B KtpcKoM nepr{qAe crrAeJrblrbre qac'ril louAeaxrr 6srnu paqlenenbt etue roJrbro cpaBnrre[I'Ho yoKxMI{ [poJII{BaMu, He npensrcrBoBaBruuMn o6ueHy $ayH. e Oco6orp BrruMaxr{s sacrr1rxuBaer Saro Harcorxlerrus A.H. Hsaxosofi (1959) 6afioce Capacoacrono flosorrxsc poslpa, HagBarruor"o eto Mesoteutltis baiosicus, Ho BHeruHecxoErono c 6opearuruiwu Paromegateuthis, oco6exuo P.parabajosicus. Ecnn Aanuwfi pocrp nefic.reurenbxo rrpunagnexl{T K poly Paramegateuthis,To rlosBgagraBns€rr AonycKar' xorg 6u rpatrrenlre grono aprcDt+recrono pqqa s floaonrGe qactrn paBnuxbl c AprcruPyccrofi s roxuofi Mops cBsob KoBpeMeHH)ruo 6afioccroro qecxuM 6acceftlov (atlArruro qepes flesopcxyrc anannny). Oasaxo HH B 6acceftHe Mops xer. flosIleuopu, Hr{ B Cpeaxeu u BepxxeNa floson}Ge creAoB 6afioccroro 154
aacrb; 3 - Cn6upcKas naaeo$nop[crr{qecrag - Teflr.recrs{fi o6nacrs); +6 nonc (4 - Cperu3eMnoMopcKas naJreosooreonpaQnqecKas o6[acrb, 5 - HntroTnxooreaucKas narleosooreorpaQnq@Kas o6nacrs, 6 - Hnno-Bponef,cxan rraaeo$aopucruqecras o6tacru). EyxsaMH rroKasaHo pacnpocrpauenrre orAelrbxbrx poAoB 6eaevnnros: M - Megateuthis, Pa - paramegateuthis, c - cytindroteuthis, P - Pachyteuthis, Ps - Pseudodicoelites, Rt - Rlropaloteuthis, B - Belemnopsis, H - nibolites, D - Dicoelitidae
ToMy ro nosBneHl{e 6orree [pe.AsraBnreflbnono M&Tepnalra Bo[poc o cBcgo( a 6afioce AprmuqecKono n l0rso-pyccxorp 6acceftroa crreAy€rr @Tagru, crrxpbtrbrM. B EopeanbsoM nosce B HaqaJIe 6afioca [pqAorrxaJr cyqegDBoBarb rotrbKo crJrbso o6enseHnuft aarleucKltfi rouruerc 6eneuurroB (pqrr Pseud.odicoelites, Sachsibelus, Hastites, Holcobelus n lfesoteuthis). Kax arAno Ha pnc.4S, onaqrTertbxo coKpaua€trcs no cpaBHeHHIO C nosnHrrM aaJreHoM rr KoJrrFrecTBo Bl{IOs (C 16 AO g). cruparorcs 6slslune B Toape I{ Haq€ure aarre*a c'noJrb pegK,,Mr{ pagJrrq,,g B co",,.aBe 6eneuxmoB B o6craHosxax npn6pexnbrx r{ orrcpbrrono Mops. tlro xe raca€rrcs neorrpa$mecrru( rpaHI{Ir pacrPoc'rpagegus panxe6af,occrorp 6opeanaxono roMnnerca 6eneMHrrOB, To l{3-ga OnpAHnqeHHOgnHr{MetoEIIrKCs AaHlrblx t[x HaMstlfrb TpynlrOo B cepeguxe 6afiocar TorIHee nocJle crrJlor(eHHs croeB c Normannites u-Arhelloceras, T.e. Jloctle cepeAlrltbl BpeMegI{ Stephanoceras humphriesianum, HaglDyrla€T peernfi nepelroM B cocTaBe roMrurerca 6eneMlnrroB Eopeanurono nogca. flonnoqttp BbIMI{paroDlfastitidae (Hastites y Sachsibelus), uoqrn rrorrrrocrbb pseudodi.."fi_ t!nae' I-lepexonm qepes sTor py6ex nrurb enr{rrr,nrrle ltug"tuui[inae: Mesoteuthis (M'inornata), Holcobelus (H.kinasovi) r{ BrrepBbre rtoqBnsercq nogufi grueMnqrbrfi
r55
pqn Paramegateuthis, sacennBunfi KaK [pu6pexxbre MenxoBoIHbIe o6cranoexl{, TaK I{ Bocrsquofi Cu6upu, CenepoSocroKa x o6crauoBKrr orxpbrrono Mops B [pelenax (puc.4'7). B npr6pexublx $aurmr npeo6naaanu KopctrKoAanuxero Bocrora CCCP pocrpoBbte P.parabaiosicus, a Qauuor crrrpbrTorp Mops tlopvu c y[nrHerrHbIMI{ PoclpaMH - P.nescia. B Ceaepxoft Ar.aepure, Kar( ),xe noBopunocb Bbrure, Ha)re[Ku Paramegateuthis uetroBecrnbr. B 3anaasofi KaHaqe ytcsobrBaKnts roJrbKo o6uapyxuBaeMble qAecb B[epBbre Cylindroteuthidae: Pachyteuthis (Pachyteuthis)sp.nov.''inden.(Carc n Hansueeaa, Hs 1966) us crroeB co Stephanoceras sp. u Cylindroteuthis hemis Crickmay e-DetrHro( ropHsoHToB cpeA$efi ropu. crparnrpa$rxecroe nonoxeEl|e o6e[x gTax HaxqoK tr]rxgaer,rcs B yroqxeun]r, oco6erno ecJru )nresrb, qro B Ceaepxofi Ctr6rpu nepBbre Cylindroteuthidae nafiAenu nuurb B cpegneu 6are. flpu conpeMexgoM cocrpslrrrtr gtranuf, npnxq]rrcs Aorrycrurb, rrno KoM[neKcbr 6ener'rnnroB B Eopeanruo-Trxooxeancrofi n Apxrruecrof, o6nacrsrc Eopeanrxoro rrosca s 6afioce pasJlrqanncb Ha ]ToBBe cerr,reficra [Cylindroteuthidae a nepaof, npoBru[r{u u Passaloteuthidae (Megateuthinae) ao aropofi ]. Taroe rpeAnoJrox(exre yBsgblBas-rcs H c TeM, .rto HoBar gruleMnqnas ane EopeaJrbrrolo ilocca npynna aMMoHIrroB Cardioceratidae (,Arctocephalitinae) e 6afioce n Haqarre 6ara roxe 6rrna B cBoeM pacrrp@'Tpatlenurr.EopeanureTnxooreancxofi orpannleua o6nac.rsp. Cxasasxoe Aaer oclloBaxue BosBqtrrb B pasr o6nacrefi Apmr.recKyro H EopeanuHe TrorooxeancKylo rrpoBrururn, BbrAensBur{ecs pa[ee B.H. CaxcoM r{ rp. (1g7]-) u C.B. Menenrnof, (1973). Bantgo nqAqepKr{rrlrr. rrro pegroe ngMeuenr{e cugreM€}Tr{qecroro c@Taea 6ene}rHIrnoB B cepeAl{He 6aftoca a EopeanbHoM nosce coBnalro Bo BpeMeHH c He MeHee pegKlrM I{sMeHeHueM cocTaBa aMMoxmoa. B Mopsr( Aprcruuecroft npoenxqrru [onHocrblo ucqegaKn paHne6afioccK[e o6uue c TennqecK[rM noccoM peqrl aM]{oHr{ToB ( Normannites ns ceu. Otoitidae) u ceveficTga: Graphoceratidae''\Tugurites) n Stephanoceratidae (Arhelloceras). Bo aropofi noJroBr{He6afioca a Aprcrnrecrofi o6nac:nn eoo6qe Her HaxqoK Ammonitina (ue aMMouou[efi cocpaHsrqrcs roJrbro r@MonoJl|frbl Phylloceratina), a EopeanuseTnxooxeaxcxofi o6nacrn B nosAHeM 6afioce HapqAy c nepBbrMr{ Arctocephalitinae (!ttegaspltaeroceras, Eocephalites) ceveficrs, ceoficrneHHbrx Tetucy. xrnu Mxonue [pegcrasurenl{ Ilpnvruu flepenoMa B pasBlrrun 6opeanbnblx nonoBononru( MoJrnlocKoB B cepeAnHe 6af,oca nora HeccHbI. Hx voxso ucKarb nn6o B HsMeHeHnHTeMneparypHono pexnMa Mops, nu6o B HgMeHeHr{HconeBono c@TaBa eoa. Haao npr{guarb, rrro a 6exroce cpeAu apIcrHqecKIDCABycTBopoK n @pauuun$ep B cepenuHe 6afioca He Snrcnpy€ircrr nq[o5xblfi peexufi nepenoM a QaynncruqecKux KoMnneKcal(, Koropuft, xaoan@s 6ul, AoJrxeu 6rln nponBrrlbcrr npu l{sMeHeHr{rdreMnepar}'p Bo[br, Ho Kcrropufi von He qyBcrBoBarbcs B MeHee greHoraJInLIHHbrx rpynnai $aynul npN He6onbur{x rroMeueHltfit( cotreHocTn. BMecTe c TeM, He1tbgn He cquTarbcc c qrcyrcTBl{eM Ha cerqnnunnf, AeHb AeraJrbgbrx rd ueneHanpaBneHxbtx HccneAoBanufi cpeAHeppcKru( apKTuqecKro( ABycrBopoK H $opauuuu$ep. BaaMox(Ho, TaKne HccneaoBaHurr BbreBrrr H B grl{x rpynfia:( $ayuu nepenoM B pagBr.rrr{r,rB cepelnHe 6afioca. flpuseleggble B rlpeAbtnyurefi rnase n€ueo/reMneparlpgbre otrpeneneuuq (cr,.r.puc44 ) TolKe He o6uapyx(HBalor B cepeA[He 6aftoca SaMerHoro ugMeHeHr{rrreMflepar]pHoro pexHMa a cn6npcKlo( Mopruc. K coxaaeHr.rro no 6afiocy raKr{x ofipegenenuft ssHo noKa HeAocTaToqHo. B 6arcxor'a BeKe eaponeftcrne Mops vr npunerarcuue K Hr{M Mops lOne 3anaanoft Asuu Hacensn KoMnneKc 6enevnuroa, B o6ureu cxoauslfi c 6aftocpoabr Holcobelus, Belemnopsis, Hibolites, Dir-oelites, Conodicot,cKuM. 3ro lites, ltlegateuthis. Bnepeste nocBntuorcs I)u'aliinae - po^ Rhopalotetttltis, B Mopsx ljoxAeaHsl l{Merorcs yKasaHus Ha HaxoAKIr ronlKo Belennopsis (/vlaAaracKap, HxAoneelrg ) . B EopeanbxoM nonce e Aprcrn.recxofi o6nactrr npoaonxaJr cyuecTBoBarb KoMnJleKc BIAtroB Paramegateuthis (puc.48). B panHev 6ate ylKe He o6HapyxnBarorcrr penHlcrbl Toap-aaneHcKor0 KoMnneKca - poAbr ltlesoteuth is s HolcobeltLs. B M€nKoBoA_ uux {aunror npeo6naAanr KaK H B nooAHev 6afioce, Paramegateutltis parabajosicus, 156
a Sauunx ctrl(pbtroro Mops, gacroJlbKo Moxuo cynrrrb no r{MerouleM}rcs Marep[afly, P,nescia. B cpensev 6are r Paramegateuthis rrpucoeAr{Hsrorcs Cylindroteuthidae, trpoHltrouue B Mopn Ceaepuofi Asta{ ua Ceeepxofi Auepuxll. P.parabajosicus oaMeua€trcs caoeodpaensn'a P.pressa C CrIJrbHopeAy[rpoBarrHbrM p@TpoM. B noqageu B roMrrneKce apmnqecKru( 6eneprHmoB Bospaqraer, 6ane ponb Cylindroteuthidae ouu cocraB,nsror )ry(e nonoBr{Hy roMnnerca no KoJrr{qecrBy Bn[oB. Bnnorb Ao nooAlreno. 6ara BKJrprrI{TeJrbHo a Apmuvecroft o6nacru Beponruo e o6nacrn crrKpbrrono Mops coxpausJrr{cb P s eudod,icoelite s. Ha nperorcexr{l{ 6ancroro BeKa Paramegateuthi's BMesre c Cylindroteuthidae paccenflorcs Ha oaIIaA a ApnruuecKoM 6acceihre Brurc'rb Ao 3evnu K v n i l P a r a m e g a t e u t l t i s( P . i s h m e n s i s u p.tina,ensjs). lloAo6uag xe KaprHua ua6npraercs a Aprcrurrecrofi naneosooneonpa6nqecrofi o6nacTI{t rAe e CeeepHofi Cu6npra Parantegatettthis (3 erua) coxpaHsrorcrr B cnnoxegsru( soHbl Arcticoccras kochi, Ho nonuocrbro orcyrcTByror B soHe Cadocerar;i;;r;. Bnonue BogMo)l(Hor qro Parantcgatetttltis B HaqaJre KennoBes >Kunu u e pafioue 3evnu Qpaxua-Hocr.rfia
yKA SATn.nb BIIIIOBbTXIIASBAtIIIil II PSEUDODICOELITINAH ITIEGATEUTIIINAIi
a e q u a l i sM e s o t e u t h i3s l * , 4 9 , t o , 5 l , t 2 , 1 3 4 ,
14r
l . r a n z r r r s i .A r
59,134,l4o
curtus Mesoteuthis 57 c I av atoide s P s eudodi co e I i t e s Bi'*, 96, I 04, I 0j,
106,107,I4l, r42
conoideus Acrocoelites I l*, 23,l32,l?,4 dillbergensis Acrocoelite s ll* , t4,ltz e l l i p t i c a M eg a t e u t h i s 6 l e l o n g a l a M es o t e u t h i s 3 0 , 4 t giganteus Megateuthis I 14, t24,143, lrl, llj gracilis Acrocoelites 10", 14,]3, 132 gracililormis Acrocoelites ll-, 4l gravi s Holcob e lus 74, 7 5, 83, ltr, 1 3 1 6|, 4 1 , t 4 Z g u s t o m es o v i P s e u d o d i c o e I i t e s 8 3 ' , 9 6 , l d t 7 , l0g, l0g, l3g h i b o l i t o i d e s P s e u d , o d i c oe l i t e s 9 7 , 9 6 , l 0 Z , l O 4 , 106,107, l2g, l29,l3g incurvatus Ho lcob e lus 77 ilminstrensis Acrocoelit-es l0*, 14, 15 inornata Mesoteuthis 3l-, 46, 47, 48, 49, lZ3,
126,l}|., t34 islmensisParomego,teuthis 60*,5!" 65,67, I I 4, 1 2 5 , t 2 9t ,3 4 , 1 3 51, 5 7 j onenschi Acrocoelite s IZ', 23,24,Zi.,Zg,lZZ, 127,Ltz,l4O juuenis Acrocoelites I l*, 132 kedonensisAcrocoelites 10, l2' ,25,26, 29,
r23,126,lt4, 135,l4l hinasou i HoI cobe lus 76, 77, 124,! 28,129,136, t4t I a e v i s M e s o t e u t h i s3 l * , l 3 l , I 4 7 laptinshajae Mesotetttftis l0', 33,39,&,
42,
l zt , lj4, l4l g 2 * ,8 3 ,8 48, j ,8 6 ,9 0 ,I 1 7 , Ie n ens is Lenobelus 123,rt7 , l3g longiconusAcrocoelitesl0*, 14, 15, 132 l o n g i r o s t r i sM e s o t e u t f t d3s0 ' , J 3 ,f r , 3 6 , 3 7 , 3 g , 124, r34, l4l
/ n n . g r . s s i m.rsr M e s o t e u t h i s 3 4 , t t , 3 8 . 4 9 m a n i l e s t a P a r n m e g a t e u t hsi 6 0 * , 6 2 ' , 6 3 , 6 4 , 6 t , 6 7, 1 2 5 , l } i - , 1 3 0 , 1 3 5 , I 4 l m a t i s c o n e n s i s , t l e s o t eu t h is 3 l ' m i n a e u q . e L e n o b e I u s B Z , B 3 , g ? . ,g 9 5 , 96, lz?., 4 rt7, l3g, l4l, 14? nunieri Holcobe lus 72, lj5 r r e s c i a P a r a m e g a t e u t hsi 6 A . 6 1 , 6 2 , 6 3 , 6 5 , 6 7 ,
1 25,l 2g, 130,134, 13t,I 40, I 55
o ) t r * c o n u sA c r o c o e l i t e s 9 , 1 0 "
t t4, t72
20,41,42,43,
omolonensis Acrotoelites 10, l.Z',27,2g, 29,
t 2 3 , 1 2 6 , r 3l44,t , t 5 l
p a r o b a j o s i c u s P a r a m e g a t e u t h is 6 0 * , 6 7 , 6 g , 6 9 ,
7 0,7l , r25,l 2g,134,135, J 40,I tl , 152,ItJ
pergracilis Acrocoelites l0',13
13,14, 123,
132,140 pergrandis Mesoteuthis _30*, 32,73,34,t6,lZ3, 1 26,127 ,134,l 4o persulcatus Acrocoeliles l I* ,Zj p l a t i v e n t r i o s u s P s e ud o d i c o e l i t e g 3 * , 9 6 , 9 9 , 1 0 0 , l 0 I , 1 0 4 ,I ? r , I 2 g , l J 7 , l 3 g , t 3 9 polaris Acrocoelites I l-, 17, lB, lg,ZO, 123, L 2 7, l 4 l praecox Belemnites (Acrocoelifes ?) 9,l1l,
t47
pressa p aramegateuthis 60', 69,70,7 I, llt,
1 3 5 ,I 5 6
primoris Pseudodicoelites Bi'*,96,10g, 109,
tto, 123,t36, ltg, t4l
pseudoelongata Mesoteuthis 1l'
47, 49, 62,134,L47
,!,4t,
p y r a m i d a l i s M e s o t e u t h i s3 l ' , ) 9 , 4 2 , 4 1 ,
123,134, t4g
44,
q u e n s t e d t i A c r o c o e l i t e sI l * , 1 7 , 1 8 , 1 ? , 2 r a u i M e s o t e u t h i sn ' , 4 I , . L 3 4 r ec o n d i t u s L e n o b e l u s B Z ' , 8 3 , 1 2 5 r h e n a n aM e s o t e u t h i s 7 9 , 3 I ' , 4 7 , 6 4 , l l 4 , 1 3 4 r o s t r i f o r m i s Mes o t e u t h i ; 2 3 , 5 6 , 5 7 sibiricus Lenobelus 8?* ,9t,Bt, 89, gO,l?t, l3g, L4l,142 spinaelormis Acrocoelite s 12*, lt4 stimulus Acrocoelite s lQ', 36,132 stricfus Acrocoelitas I l' s t r i o l a t a M e s o t e u t h i s 3 0 t , 3 r , J g , 1 i g ,4 0 . 4 2 , L Z Z , 134, L4l
l ) J l r ' r r r r l f i x o fni o g v e p x H y r b t c r p a l t l t u b r , r ' A e / l a e r c R o n n c a l r r e B l r A o R ,3 B e 3 A o q K o f tf i o x a c a H H u r p a u l . u l b tB o n p e A e n i l T e J t b n b l rxa 6 a n u a x B n a o B .
f58
46,
* s c r / i d u s , 4 c ' r o c o eI i t e s I I * srrl,rbrepi.stl e soteuthis 3l s u h c o n o i d c a M e s o te u t h i s 5 4 , 5 6 srrbgracilis Acrocoelites l0+, l!, lJ s t t b r o s t r i I o r m si M e s o t e u t I t si 3 l * , ' 5 4 , J 5 , 5 ( ) , j 7 ,
58,123,140 subrhenana Me soteuthi s 3l' s u b s p i n a ef o r m i s ' A c r o c o e l i t e s l l * subtenuis Acrocoelites l0*, 14, lt, 16, 126,IZ7,
ttz, 140 sublriscisstt s Acrocoe lites lI*, 23. subpyr ami dal i s ill e soteu t hi s 23, 3l*. s u p r a p a l a t i n u sA c r o c o e l i t es 1 0 , l 2 : , 2 6 , 2 9 , 1 3 4 t i m a n e n s is P a r a m eg a t e u t h is 1 9 , 6 0 ' , 6 5 , 6 6 , 6 7 ,
69,129,I 30,I 35,l4O,lr7_ -,
l r u n g e n s i sM e s o t e u t h i s 3 l
52,14,122, lll,
134,t40 t r a b e c u l d A c r o c o e l i t e s! , 1 0 * , 1 4 , 1 3 2
tramayensisAcracoelites12',L)2 lrduthi Holcobelus 76 tripartitusAcrocoelite s 9, 14,17, 21,24 t r i s c d s s u sA c r o c o e l i t e sl 0 , 2 0 , 2 2 , 2 3 , 2 r , 4 1 , J 4 , r 2 2 , 1 2 7 l, 3 2 , l r 5 , l 4 l r t r i s c r s s i / o r m i sA c r o c o e l i t e s . l l , 1 7 . 1 8 .2 0 . 2 l l r i s u l c o s u s A c r o c o e l i t e sI l ' , 1 4 ,! 7 , ! 8 , 2 0 , 2 1 , 22, L?6,127, ltz, l4O turniaceusi s Acrocoelite s 12* umarensi s Ho lcobelus 7 2, 7 r, 76,,115 unisulcdtusAcrocoehles 10, LZ-,26,29r1t4 a a g t L e n o b e l u s8 2 ' , 8 3 , 8 5 ,W , g L , g z , g l . , g : . , 123, 137, l3g,142 v e n t r a l i s A c r o c o e l i t e sl l * . 1 3 2 viligaensis Lenobelu s 82*',8t,W,87,88, I 10, l?1, 127,ln, lt6, lt7, l3g, l4l, 144 voltzii Acrocoelites49 vulgaris Mesoteuthis30, 31, 45,134
JIITTEPATYPA
1964. O cncreuarl{qecKoM nonoxeHr{n Neohibolites semicanaliculatus }r Ann-3ate A.A. o pole Neohibolites. flaaeonr. x)fpH., !, c.78-86. 1965. Hosbtft poa ceveftcrea Belemnitidae. - llaneour.)KypH., 3, c.L42-144. A.A. Anu-.3ate Eepaun T.G, KunplrKoBa E.II., IIoasKoBa V.8., HeftquH,[,.fI., Caxc 8.H., Tefic P,8,, Xa6aros A.B. 1970. Hexoropeie npo6neMbr naJreoreMneparypHoro aHaJrr{sa.- leon. n neoS.,'11, c. 36-43. A.B. 1966. Knunaarrt Eepann T.C., HafiaxH A.fI.r Caxc 8.H., Teftc P.B., Xa6aroa B ropcKoM H MeJroBoM nepr{oAax Ha Ceaepe CCCP no naJreoreMneparypHbrM o[pea€Jl€HHBM.leofl. n reo$., ltQ .. 17-3 1. A . B . 1 9 6 6 . ) O r u u x e a H a r H T [ q e c K [ e o n p e A e n e H n s o T H o r u o H l { eK a J I b EepanH T.C., Xa6aKoa urrq H MarHHs B pocrpax 6eaeunonAefi xar Meroa oueHrs reMneparyp cpe!,bl o6nranIas a Mopsx MeJIoBono nepl{oAa CCCP. - feoxnvuR, 11, c. 1359-1364. EoAstJreBcxnft 8.14. 1960. Hosbre [osAHeropcrc{e 6eaeN{Hntt Cesepnoft Cn6xpH. - B rx.: 'Honste Br{,qbr ApeBHHx pacreHnft u 6ecno.asoHort}rblxCCCP'r.r.2. M.r'HayKa', c. 193-1 95. A.A. 1908. @ayxa aoHeuxofi lopbl. l. Cephalopoda. Tpyau I'eon. KoM., HoB. bopncsr 96 c. cep., E, 'Hayxa', 2O7 c. P. 1969. llaneoreMnepar)?tr.stft aHaan3. /l.r Eoyen oMarep. BopoHeu H.C. 1937. OayHa MopcKono Mesosos Elpenncxoro 6accefiH&. - B xn.: no neoJl. Elpenncxono KaMeHHoyronbHoro 6acceftna'. II., c. 47-74. orBoponeu H.C. 1962. CrparnnpaSun n roJroBonorne MonnlocKn IopcKI{x H H}r)KHeMeJIoBbIX aoxexnfi Jleno-AHa6apcxoro pafioua. M., focreoJlTexHsAar, 236 c. A.E. 1969. Heroropbre paHneMeJroBbre6eaeuHnrsr Pyccxofi naar$opvrl. - TpylaasyHoBa Ebt BCE|EH, uoe. cep., 13Or c.22L-231. F o p n H . K 1 9 6 8 . K c n c r e M a r u K e p a H H e M e n o s s l xB e l e m n o p s i n a e . - I l a a e o u r . ) K y p H . r 3 r c.99-1O1. . flaaeorr{A}oxr{M}rs reppurenHbrx Torlul. - Tpyalr HHHFA, l,Xf, 17Oc. fpar',r6epr H.C. 1973. H . C . , C n u p o H . C . 1 9 6 5 . F l a a e o n n A p o x l r M n sc e B e p a C p e n s e f i C r 6 n p u B r r o 3 A fpav6epn 12O c. HeM naJreq3oe r{ Me3o.3oe. - TpyAU HVlUlA, I42, E!,\ 1956. Bepxuercpcrs{e 6eaer{nnrsr Pyccrofi nnar@pvst. AaropeQepat lycroMecoe KaEA. Ar{cc. Mocx. Foc. ynnsepc., 21 c. B.A. 1956. f ycroMecoa K gxoaonnn BepxHeropcKlx 6eaevsnroe Pyccroft naar@pvrt. Enna. MOI4[, orA. reoJt., 61 (31), errn. 3, c. 113-114. B.A. 196O. Hosbre nooAHeropcKneH BaJraHxxHcrue 6eaeunxmt Esponeficxofi f ycrouecoB qacril CCCP n Ceaepnono 3a1paabs. - B rn.: tHoaue BnAbr ApeBHHx pacrexnft r 6ecnoeBoHorrHbtxCCCP', 'r. 2. M., 'Hayra', c. 190-209. B.A. 1962. O sHa'reHun 6oxossrx 6oposa pocrpa Arc pa3pa6orxu cHCT€M€rfycroMecoe rtrrs{ 6eaeMxuroB. - Ilaneout. x}?rlr =1' c. 31-40. Pycf y c r o M e c o s E L A . 1 9 6 4 . E o p e a n r r o r e n q e A H e r o p c p r e 6 e a e p t r r x r e r( C y l i n d r o t e u t h i n a e ) crofi [email protected]. - TFyrbr fHH AH CCCP, 1O7, c. 91-209. 8..\ fycroMecoe 1966. Hoabte 6eaevnnrsr Hs roapa H aaJrena Cn6xpr, - IlaaeoHt.xy?H.r 1, c. 60-71. B.A. 1973. O xacrurax H xasrnronoao6rurx 6eaeun[Tax. - Enna. MOHil, fycroMecoa or.xL neorr., 48(21, c. 113-120. B.A., YcneHcKas E . A " 1 9 6 8 . O p o g e R h o p a l o t e u t h i s( B e l e m n i t i d a e ) n e r o fycroMecoB c. 65-78. KpbrMcxux npeAcrasilTeJrs)q - Ema. MOHn, oTA. reoJr., 1q(5), A . A . 1 9 6 8 . T o a p c r c r e a M M o H H T b (I D a c t y l i o c e r a t i d a e ) C e a e p a C n 6 u p u . - T p y n s r tlanuc
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fin 'nn.- rpvarr ^";1T"fi.;i.:"[Jj#:"";iil:i:'rg*:y, BHl4fprl: @, ;. un-"o. (roapoo,ri o".""in.p. anno". roriTTffi;ff' "py.) K o s a o n a ' , ' ' E , K n n" p H K o B . - E r . , &flr::?l'j"#*'ti"nu" Hrta"r'i.n., caxc
u.ffig7g. ueHTpuqecxoficaoncrocrr.t B pocrpax o n p ' p o A eK o H _ 6eaeunuroe. _ fe_o-a. a.o6., g, *tH;"fr"#:.t932' c. g8_4g. IOpcr"" 6.r,erxursr l(psrva " n Kasrasa. - Tpynrr taas. FeoJr.-pa3B. Kplt r'anorru l'f' Lg47' Knacc ce.phalopoda.orpsa.Decapoda, noAoTpsAB.turnoia"". Eeaer,rsursr.Araac pyKoBoAsunx $opv n"nonu"rurx $ayn CCCP, T. Br KpstvroJlbu f'fl' rg-og' c. L9S_2O6. n*-t""4"a."".'r.iiu.-nnrto."".pu*o""HHbre. ToJrorr.rH. ocsoBrl ,arreo'Moaarccxr.r_roJroBoHor[e. fI, M., c. t+S_tZZ. Kprtr"rnoJIbu f'f' l-960' Anttoq"*t o[peaeae'Hs MeoooolicxnxnoJroBoHon'x. 89 c. HsA-ao /rfv, KprtvnoJlbu f'fl' 1965' Eeaeunnml xaK rroKasareJr'Bo3pacra B Meoosoe l(apnarcxo"*; ^::";,r:"Jil;
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Kprrr'anoJrbu f.fl., flerposa f.T., [I .{eaHHueaa B.O. 1g$g. CrpanrnpaSnn u MopcKHX vesosoficK'x o?rroxeHlrfi Qayna cesepHofr cn6npa. _ Tpyalr t";;;;lt HHHIN 45, 131 c. Avr'ro""t", r'r so'arb'as jg73' crpar'npagr.ra 6afioc-.6ara ft? cn6rpn. Hoeo''*i'M tneArna c'8" H aasHseBu t'": 1g73. feorpa$xvecKoe pacnpocrpaHeHr{e H 6ereMrrrroB B noo,qHeM aMMoHHToB aaJreHe EopeaarHori oOr""r". _ B xH.: [aaeo6noreonpaQna sepa EBpa.rl Meoo3oe.- 8M., ce." ; . "Hayra, f,- nilg. M o c r a r r e H x o 3 . :. [ . 1 9 6 g . rcpcKHx oraoxenufi Bepxueno flpnavpra. alr Hlnl Co - TpyCCCR n" .:;T::H3 H a e a p H u B H J r " 1H 1g69. p;,a Mttcrottihorite.s T'r'' Nasarishvili, JI[KaHcK€Is roH$epeHu's 6Jen. no\,. - B xH.: pecny6_ MoJIonbIX-JrqeHblx fi acn'paHron rpysuxcxofi ccp, lOO-aernrc nocasueHHas co Ans pox.qerrng -BepineveJroBbre B.H.IIeHun a. T6unucu. HafiauH,[.n. 1964. 6eaevxursl pyccxori naar$oprurr u coflpeaerrb'brx AxrnrroxavaKcbr, nonuoreftrucu u 6enevueaaoKaMarcbr. :ffi"r-' hea-ao Mry, H afiaH H A.n. 1965. BepxHeveaoatle_6eaeMHr.rrbr (cevericrao IJelemnitellidae pavlorr,) .M., cxoft naar@pMbr H conpeAeJrbnrrx pyc_ o6aacrefi. Anropeo. aoxr. o"... 4L .c.
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EIl., lloncKoBa UJ.. L9'I 2. Marrurft n crpoHCaxc 8.H., AHnxuHa f.A., KunpuKoBa Boabr ApeBHr{x Mopcxrrx 6acce*Hoa. uuft e pocrpax 6eneunnroB - r{Hru{xa?opbr reMnepar}? leoa. u reo$., 2, c. 1O3-11O. A.A., Jlanuc A.Gr 3axapoa 8.A., HsaHoBO E.Q.r Caxc &H., EacoB B.A, Ilaruc M.C.r HaasHseBa T.H., lllynurHHa H.H. 1971. MeaetrnHa C.8., MecexHuKoa o6ut. r'ropef, Fopeaasnoro noflca B Iqpe r{ HeoKoMe. - C6. 'flpo6leuu flaaeq:ooreorpaSne penuoH. u reor.' Hoaocn6rpcK, c. t7g-2LL. H.C., PoHKr{Ha 3.3., AnaoHoBa 3.H. 1969. Meeosoftcpre orCaxc 8.H,, fpav6epr snaaxHu. - TFyag HHH|A, 29, 226 c. rroxenus Xarasrcrofi A.C., MeneaHHa C.8., MecexnrrKoB M.C., IIepnaCarc 8.H., .[I,aFlrc A.A., Aarllc MeHT M.A. 1972. CoeeulaHne no 6uocrparurpaQnn Mopcxoro Mesooos Cu6npn u Aaatnero BocroKa. - leor. u reoS., 7, L36-I47. ". Caxc 8.H., HaJrbHsesa T.H. 1964. Bepxuercpcxile u HuxHeMeJrosue 6eneN{runu Ceaepa CCCP. PoaH Cylind.roteuthis u Lagonibelus. JL, Hsq-so AH CCCR 166 c. Bepffreropcxfie H HHxHeMenosHe 6ereMnlrrbr Cenepa Caxc 8.H., HaJrbHseBa T,VI. t966. CCCP. PoaH Pachyteuthis u Acroteurhis. M.r 'Hayxa', 259 c. naaceveftctsa Passaloteuthaceae D noaCaxc 8.H., HaJIbHseBa T.H. 1967. O BhIAeJreHuH orpsAe Llelemnoidea (Cephalopoda, Dibranchia, Decapoda). Jlorn. AH CCCP, 173, lt 2, c.438-44L. T.H. 19676. K cucreuarr{xe lopctorx n MenoBbrx 6ener.,rHnroB.Caxc 8.H., HaasHseBa r"reoooog Cho6ocHoBanr{g Aerarugoft crparnr?a$rn C6. "flpo6aeMbl n€MeoHToflorntrecroro 6npu n .fl,aasneno BocroKa'. fl., tHayxat, c. *28. T.H. 1970. Panne- H cpeAneropcrore 6eaeMHHTbrCeaepa CCCP. Caxc 8.H., HaJrbHseBa Nannobelinae,Passaloteuthinae n Hastitidae. fl.e 'HayKa', 227 c. Cbxc 8.H., PoHKr{Ha 3.3., lllyaunHHa H.H.1Eacog 8.A.1 EouaapeHKo H.M. 1963. CrparnnpaQnn rcpcrofi u r.{eaoaoft cncrev Cenepa CCCR M., Hoq-ao AH CCCR 226 c. B.H. 1933. O ueroropux rcpcnrx oKaMeHen@Tcx c P./1., EoastJreBcxnfi Car.aofiaoan{ ocrpoBa fyxepa (3er.,rac Opanua-Hocu0a). - Tpygbr Apxr. uH-Tar L2, c.29-4L. Cn6upsKoEa /1.8. 1961. Cleaxercpcxan $ayna MoJrJrrocKoBEoasuoro Eaaxana u ee crparnnpa$urecKoe sHarIeHI{e. - B xn.: 'flpo6ler',ru ne$renocuocrn C}eAllefi A,snr'. - Tpyqbt BCE-[.EH, Bbtn. 5, HoB. ceprs, & 223 c. Onncanxe HeKoropbrx BHaoB N,resogoftcrorx oKaMeneJrocrefi ns Cnu6npH.O. La77. Cnsuoe ' o6-sa ecrecrBoncrr.lT. 5. OAecca, 5 c. cxofi u CaparoncKoft ry6epsnft. 3an. Hoaopoccuficr. B.l,l. 1963. CaaanH Tpnacoaue H ppcxr{e orJroxeHns Bocto'rrbtx Kapnar n flannoucxoro cpeanHHono MaccHBa. focneoarexnsaar, 171 c. gen nov. (Belemnitida) oT aonHorpeaF CrocHoBa-BeprnrroBa M. lg6S.Curtohibolites Huro oTJroxenr{s n Erarapnn. Ernrapcra Axaaer,crc Ha Hayrcrre! TpyAoBe Bbepxy FeoaoI'Hsra na Eurapus, cep. n€ureonT., KrL 5, c.2LL-227. CrosHoBs-BeprHJroBa M. 1969. flpeacraonrerr{ Ha ropcrxs po( Rhopoloteuthis a Erbfirapr{s. - Hss. Ha reoJr. }rHcr.r cep. naneoHToJr., rx. 18, c. 97-104. Tyvxoa lopcxne aMMoHr{Tbr n 6enenxsrar Ceaepo-Boqora 14.V. 1954. CCCP. - Marep. rto reon. tr rroflesn. Hcxon. Ceaepo-BocroKa CCCP, Bbr[. 8, c. 96-125. tA.E. 1952. Menosue 6eneunnru flyenn. - Bectn. loc. M]roea $ysnn, XeqnHauBunu r. 15-A c.63-116. lopn 1., JIoyeHuraM f., Snurefin C., Max-KnHBu K. 1954. Orpenenenre nareoTeMneparyp, B r{acrHocrn TeMneparyp BepxHeto uena AHunH, Aaffilt tr lorb-BocroqHbtx uraroD ClIlA. - C6. 'hcoroml B neonorHu', Il., VJl, c.543-572, A b e I O.' 1916. Palloblologie der DibrEnder Cephalopoden aus der Gruppe chlaten, 282 S. Jena, Bayle.E., Zeille r. 18?8. Explication de la carte gdologique de laFranprincipaux ce. Fosslles des temains. Atlas, wol 4. Paris. B e n e c k e E. 1898. Beitriige zur l(enntnis des Jura in Deutsch-Lothringe&geol Abhandl Specialkarte Elsass-Lothringen 9? S. N.F. FL 1. Strasburg, Benecke 1905. Die Versteinenrngen E der Eisenerzformation won Deutsch- Abhandl Lothringen geol und l;uxemburg. Specialkarte Elsass-Lothrigen. 6, Straesburg!" N.F.Ii . 1 - Nouw. Bull. sci. pan bfainwille D. 1825.-'M6moiie sur les B6lemnites. philomat. Soc. p. t7!-1,76, Par{s, B lainwi'll e D. t82Z M6rnolre sur les B6lemnites consid6r6es zoologiquement et g6ologlquemenf. Paris, 136 p. BlUthgen J. 1936. Die Fauria des Oberjura und Stratigraphie und derUn-terkrelde - tfniw. von l(i5nig l(arl L6nd. 95 S. Dissert. Greifswald, B o e h m G. 1906. Geologische Ar-. Mitteilungen aus den Indo-AustreUschen - Neues 1. Neues aus dem Indo-Australischen chipel Archipel. MiJahrb. - BcL 22, S. 3Aa-4L2. neral, GeoL und PaliiontoL, BeiL
162
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13 o e h m G' L912' Beitriige g-"ologie von Niederliind.isch-Indien, 1. Die SUdkUsten d.er Sula_In-sebn-rr1 Taliabu-und Mangoli, Th. 4, (Jnteres Callovi_ en" - palaeontographica, Suppl, 4, Abt. 1, A-bschi.-+, 'ZgS. Bi'ilotr-Trummer E, ag2o, cephalopoa. aiu".nchiata, - Fossilium catalogus, I. Animalia, ps. 11..Berlin, 313 S, c o m b e m o r e I R' L9?L, Figuratior, .de di^gnose d,AcrocoeEtes bobeti -plements Lissajous, Belemnite du" i " oLias superieur - Extn geo_ Francais, bios, wol 4, fasc. 1. Lion, p. 61_69, , -4 mt,emorel R' L973' Les Duwaliidae -' / \ - o P_anrlow (Belemnitida) du cr6tac6 inf6rieur r-rang?i_s-: Do-c.um. Lab. s-ci. I_ryon, N 5?, p. 131_1gS. 9,6ol, Fac. Delattre M' tgsz. caractbres syst6matique de Berriassibelus extinctorius E=iii"n (Raspait) nou geru" r - o(e"r"-nites). - e,rii Museum nat. histoire, natur", ser, z, L 24, N 3, p. 32L_32?. . Deslongchannps E. 7-8i?, r,e jura Normand,. Etudes pal6ontologiques des dirre'rs nil'eaux jurassiques de ie Normandie le description et I'iconographie de tous "ompr"nant Ls fossiles rrert6bres -.-:;;;6bres qucils renferment, Liwr" 2, Monogn 6. paris-Caijn "f D u m o rti e r E. 1a64:1'a?4Etudes pal6ontologiques sur les d6p6ts jurassiques du bassin du Rhdne. Vols 2-_4. pariso aOO p. Duwal-Jouwe J' 1841, B6lemnites als-terrait= cr6tec6es inf6rieurs des e'rrirons de castellane (Bassel-Ab;=i consid6r6es g6ologiquernent.et logiquement, ..,ooavec ta description -_ -LcaO Aes terrains. te! rqrr-. ' pa_ sci natun ris, Bo p.. _.
E r n s t w' a923-a925' zur stratigraphie und Faunenkunde des rtias g im norrcwesilichen Deutschrand. palaeontographic., B; ;!,-uu. stuttgar{, 126 s, Janerrs ch w 1 9 0 2 . D i e J"urensis-schictrien als - Abha'd' Ebass. geol Specialkarte Elsass-L6ttrringen" N"F-.;, ;. Strassburg, 151 S, J a n e t c h' 1-890' Note sur troi,s rro,rlr"tl"= B6lemnites sdnoniennes. - BulL Soc. 96ol France, 3 ser, 19, p. ZIO_ZiL, Jeletzky J' 1965. Taxonomy and phylogeny of fossit coleoidea (-oiura'chiata) (a'ustn), - cieol paper ci".a.] gL*. 65-2, p. ?2-?6. JeIetzky J' c o m p a r a t i w e m o r p ; i ; ; y , p h y l o g e n y *Mor,r="a, -'1966' and classificationof -contiu. fossil - uni.'. coreoidea. paleoritol I(ansas alt- 6, !62p. Jeletzky J" L9?2" Morphologv attd-t.;J'omic status 6r irre Jurassic bele-
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sp;il,
1e35. - pataeontology, wol 15,
r( o r b H' a942. Die Beremniten des jiingerenLias in Nordbayern - z.Dtsctt (ies,, geol Fd. 94, H. 3-4, S. 145-fig. I.( o n g i e I R' 1962" on belemnites from Maastrichtian, campanian and santonian sediments in ttre middle ..,-rr"v Vastauta (a;l;"1;oland). - prace. Museurn ziemi, N S, stn ZA. L a n g W, 1928. The Be,lemnite marls of Charmouth, a series in the Lias of the Dorset Coasl _ Ouart J, GeoL,Soq,London, wol'g+, ;i- ZrN 334, V t96_22?L e rn o i n e M, 19i-5. Re.y, crit" pal6ozool Ann 19, N 4, L i n k H' ago?' Beschreibung der Naturalien-sammlung der uniwersitiit zu Rostock' T' 3' Fossile ueberbleibsel organischer r<'6rper, sogenalnte versteinerungeru Rostock. Lissajous tVl' 19a2' sur une coupe de Lias rnoyen et sup6rieur dans te Mtsonna_is MAcon. Lissajous M. 1-975. Oueloues rFr?1AFdrt6e
BurL soc. li,stegs natun Jt?;:jff:::il";. JVl' 1'925'
t?:.tut"mnites jurassiques--
Lissaious
n6fe*oir"-.ipi'r;;"ii[r"a6=B6remnites jurassiques pr6cdde d'r:n Essai de classification, - Trav rrab 96ol Fac. scl r4ron fasc. 8, m6m. ?, I?3 p. Lissajous M' 192?' de'scription d e q' ruaecl ,q u e s n o u w e l l e s e s p A c e s de B6temnites jurassiques. - Traw' Lab, g,60r. sci. rryon, fasc, 10, m6m, ?,42.p, M a y e r-E f/ m a r r(. 1883. Gnrndz-ge der r(lassififtationr ' v r r saer petemnites. Dtsch. geol Ges, Bd. 35, S. dEo_A+s. _ -?. M i 1 I e r J. IA23, Obsenrations on Belemnites. _ 'Mem.Trans. v ' s r r r . r r q r r F GeoL Soc. Lon_ don, ser. 2, pt 1, p. 4s_62. j., Montf ort D" 1-808' conchyorogie syst6matique et classif,cation m6thodique des coquille^s. T, 1. coquittes uni.Tatrres, cloisonn6es. -;i;';;-Lrieroorith paris. M 6 r i k e w' 1894-1895. Versteinerungen d"= won chile in G' steinmann Beitriige zur Geologie und paliiontologie won sgdamer{ca. ''' Neues Jahrb' MineraL, pakiontol, GeoL una g, stutBeiL-BcL rgad, S. 1_t_Oo. 'rt?T: Die rossilen Tintenrische. Eine pataozoologische Monografie. " ";:f
r63
S . L 8 2 7 , Petrificata Nlleson Gothonrm. 39 S.
Suecana.
Formationis
Cretaceae.
* tondlni
der oberen l(re iiiber die Cephalopoeien J. 1913, IJntersuchungen Nowak ( s e r " B ) . Sci. natur' e raC r a c o w i e s c i . , A c a d . de in Polenn T" 3. Bull. covie p. 335-41'2. jurassiques, t' L. Terrains francaise. d'A. 1842. Pal6ontologie O r b i g n y 1 1 9 P a r i s , Cephalopodes. P. et cles mollusdes coquiiles Orbigny d ' A . L A 4 5 , P a l 6 o n t o l o g i e uniwerselle ques. Paris und des siidFrankreichg A . 1 8 5 6 - L 8 5 8 . D i e J u n a f o r m a t i o n Englands, Oppel Wijrttember$,' Naturkunde l'aterl Ver. Jahresku Deutschlands. westlichen 857 S, Stuttgart, Bd. 10-14, rrol t. 2nd ed, London. of Yorkshire, philllps J, 1835: Geologie Palaeontogn of Eritish Belemniticae. phillips A monograph J" 1865-1g71. p' 13O 23. London, 1?, Soc. Il'Ionogr., 19' ?-9, ?Z phillips of Yorkshire" 3-rd ed. Lrcrrdon. J. 1B?5. Geologie larrdt frrana-Joseph faunr: of Cg'pe Flora J. 19OO The jurassic Pompecky Nansen byF" and its neighbourhood of Cape Flora sketch g"ological *iif, 2, Chris1. rrol. N l?esults, Scientific bxpedition Polar North Norwegian The p. 97-1O5. tiania, of tselemnites cf the genus Rhc'* ka H, Lgs?, On two species ews acz Pug po* Acta Paleontol (in eoEsfr). Poland of the Jurassic from f-aloteuthis 3 8 5 4 O 4 " loru, Warszawa' P. of Polend' - Acta from the jurassic H, Lg6L. Belemnoids a.cr.ewska Pug p' 236 6, Warszawa' polon.' rrol paleontol - Petrefactenkunde DetrtschF. 1846-1849. Die Cephalopoden" euenstedt 5OB S. Bd. 1. Tiibingen' lands. GeoL F. 1,852: Zu den Belemniten. - Neues Jahrb. Mirreral, euenstedt und Paleontol. F, 1858 Der Jura. Tiibingen, A42 S' Quenstadt - Ann. sci..' obsel11/., des Belemnites, naturelle p.=pail ].8,2g. Histoire F.V T.J., p. 277.-33I. OoEthen-Ciebirges' des Nordder:tgchen R o e m e r A. 1g36, Die Versteinenrngen 2LA S. Hannower, de pat'6Trait6 in J. Piveteau" des Dibranchiata R o g e r J, 1gS2. Sous-classe 7Ot-731'. Paris, ontologie. P. auf Timor und Rottiim' A. f8gZ. Oie Pe,rm-Trias-Jura-Formation Rothptetz - Paleontographica, Bd. 39. S' 57-1'06' Cassal, Archipel. indischen 'schlotheim der versteinerungenin E, 1813. Beitriige zur Naturgeschichte Minera- Leonard's fijr die gesamte Taschenbuch Hinsicht, geognostischer a/M. Frankfurt l<>gi;. ? Jahrg. auf ihrern jetzigen Standpunkte' 1820. Die Petrefaktenkunde Schiotheim-p. wersteinerter und fossltrer UberSammlung seiner die Beschreibung durch 43E S' Gotha, erliiutert' der Vorwelt reste des Tier- und Pflanzenreiches - ZbL Schwabens. E. 1939. Belemniten aus den Psilonotenton Schwegler B, N 5, S' 2OC-2O8' Abt. GeoL und Paleontol", Mineral, und Ergebnisse E. 7-949. Vorl6ufige Mitteilung iiber Grundsdtze Schwegler Jahrh" Jura. - Neues des Schw&ibischen der Belemnitenfauna einer Rewision s. 294-3a$. Abt' B, N 10. Stuttgart, Mona'tsh', und PaltiontoL GeoL Minera1., E . L 9 6 1 , - 7 - g ? 1 - .R e r r i s i c n d e r B e l e m n i t e n d e s S c h w a b i s c h e n . T u Schwegler A' Bd 116' 1-7-Br7-2O, 12'4, 3.32, a38, 245 S" Abt -r"e. 1p, ,1-6. - Palaeontognaphica, Lias. 1-sL ed. York. of the Yorkshire S imp s on M. 1855. The Fossils 141p. Lias. 2-nd. eci, York, the Yorkshire of Fossils The i-884. M. p o n m s S i world maps. P h a n e r o z o i c 1 _ 9 7 3 . G . E . D r e w r y J . c . , A , . G . , B r i d e n srni[r, N 1'2. Paleontol, Paper time. - Spec, through and continents Organisms 42 P. London, dem genach Wiirttembergs die Versteinerungen ijber Ubersicht S t a h L jA24. des WfjrttemCorrespondenzblatt der Petrefaktenkunde. Stande genwiirtigen - Tiibingen' 91 SStuttgart Vereins, laridwirtschaftlichen 6ergiscfren der Jura und l(reicleformation won CaG. 1881. Zwr I{enntnis Steinmann BeiL-Bd. - Neues GeoL und Paldontol, (eoUwia), Jahrb. Mineral, racoles S. 239-301. 1. Stuttgart, 2 AufL Leipzig, S' in die Paldontologie. G. 7-gO7. Einfiihn:ng Steinmann 347-355. and cretaceous G. a965. The iurassic Be'lemnitesof NewZealand stevens of thelndo-Paciand Cretaceous of the Jurassic and a rerriew Belemnites - N.Z. GeoL 36, 2A3 p' Paleontol 3t-1[1", wol Sunr. fic Region.
164
c
st
eve ns G' 196?.lJpperJurassic fossils from Ellsworth Land, west Antarctica and notes on Up'per Jurassic Biogeography of the south pacific Region, N. Z, J, GeoL and Geophys. , rzol 10, N 2, p. 1_45_1 Sg. s't e v e n s G. L9?3. Jurassic Belemnites. Atlas of palaeobiogeopgraphy, Ams. terdam-London-N.y,, p. ZS9_2? 4. s t o I I e y E" 3'97-1'. studien an den Belemniten der unteren rtreide Norddeutschlands. 4 Jahresber' NiedersS.chs. GeoL Ven Hannower, S, Lz+-119L, S t o l1 e y E. L91-9. Die Systematik der Belemniten 11, Jahresben Niedersachs. GeoL Ven Flannower, Sg S. s t o I I e y E. 1927. Zur systematik und stratigraphie median gufurchter Belemniten' 2o Jahresber, - GeoL Ver. Flannower, S. 111-136, Neiders6chs. strombeck A. 1861- uber den Gault und insbesond.ere die Gargasmergel (.a.ptien_-d'Orb,) in nordwesflichen Deutschtand. _2. deutsch. -"' geol Ges,, i>Bd. 13. Berlin, S. 20-60. T a t e R' 1880. Description of Belemnite from the Mesozoic _of a new species s.trata of centrar - Trans. Austraria. Ror. soc. s. Austra1, wol 3, Ade_ laide, p. 104-1OS, Tenison-woods J. 1884. on some Mesozoic fossils from central Australia. Proc. Linnean. Soc. N.S,W., p. 235_242. Tornquist A. 1898-1901:,?"1 Dogger.am Espinazito-pass, pa1. Abhandl Dames und l(oken, Bd. B'(N.F. Bd: 4). Jena, S. a3S_2O4, v o Lt z M' 1830. obsenzations sur les B6lemnites. IvI6m. soc. histoire natua Strasbourg, ?O p. weithof er rt' 1890- iiber Jura und r{reide aus nord,wesilichen persien. Sitzungsber. Akad. Wiss., math.-natunrriss. KL, Bd. 9g, Abt. 1, 1gg9. Wi_ €h, S. 756-223. w l n e r E. i-gLz- ijber die Belemniten des schwabischen Lias und die mit " ihnen werr'rrandten Formen des brauen Jura (ro,.."u). - palaeontographica, Bd, 59, Cassel, S, 1,O3-L46. westermann.G. 1-9?3. strength of concawd septa and depth limits of fos- Letha.i.., .rol 6, p. 3g3_4O3, sil cephalopods. whitehouse F.w. L924. Dimitobelidae - a new family of cretaceous Belemnites. - GeoL M-g., wol 6a, N ZZ3, p, 4LC,_476, Young G', Bird J. La22. A geological sunrey of the yorkshire coast. 1-st ed. Whitby, 33 5 p. Young G', Bird J. 1a2a. A geological sura,zey of the yorkshire coast,2-nd, ed. Whitly, 335 p. Zie ten K. 1930, Die Versteinenrngen WUrttembergs, H. 4_5. Stuttgart. 1O2S. zitt e I A. 1895, Gmndzrjge der eikiontologie. - Abt, L. Berrin.
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1-3. Acrocoelites pergracrlls Sachs sp. nov. . . . . . . . . . . . . . o . .o ctp. 12. 1 - ronornn lS 87-1r nrxsn* Toap, soHa Dactylioceras commune, p. JIeeHfi KeAon ( ta - BHA c 6pnrunoft croponbr, 16 - axA c aesoft ctoponrr); 2 - pocrp IrIe gZ_4, rnrxxufi Toap' p. Keaxuep (2a - BuA c 6prcurnofi cropoHbr, 26-auac rpaaofi croporol); 3 - pocrp ]S 87-19, nnxuuft ToaPr eoua Dactylioceras .orrnunu, p. /leBbtfi Keaon (3a - nonepeqHoe ceqen'ne y Bepur{Hbr aJrbBeoJrbr, 96 - nonepeqHo"'""*"o" ;-;;;.punsnof, qacrlr)
Oxr.
4-6.Acrocoelites hedonensis Sachs sp. nov. . . . . . . . . . 4 - naJtoTHn l& 87-28, xnxHnft T'oapr p. Kemrr..rsp (4a - BrA 46 - 'sun c npaaofi croponrr); 5 - pocrp l* 8?-3O, lrnxnnfi (5a - Bra c 6prorunoft cropotrbl, 56 - alla c npaaofi cropoxu); nnxnrfi Toap, sona Dactylioceras commune, p. /lesufi Keaon, BepurrrHbt aJrbB@rrbt, x2.
TAENHUA
. . . . . . . ctp. 25 c 6prounofi c$opotGr, aaren, p. Torlp-ropsx 6 - pocrp l.te gZ^gL, nonepeqHoe ceqexrre y
II
< D r r . L - 3 . A c r o c o i l i t e ss u b t e n u i (sS i m p s o n ) ._.. o.......... o.... ctp. 14 1 - pocrp lt 87-5, rnrxsnfi Toap, soHa Zugodactylites braunianus, p. Jleaarfi Keaon ( ta - BI{A c 6prcrunofi cropoltbl, 16 - auq croponar); 2 - pocrp }\I9 g?-6r nnxsnfi ?oap, sona Dactylioceras commune, "-r."on p. fleagfi k"no" (2a - BnA c 6pguno* cropoHbl' 26 - P,na c npaaoft croponrr); 3 - poctp lrls 87-8, rnrxsnfi mapr sona Dactylioceras commune, [onepeqHoe ceqetcre y Bepu'Hbr arbaeonb.I, x2. Qrr.
4. Acrocoelites trisulcosus(Simpson) . . . . . o . . . . . . . Pocrp ls 87-12, 'orxunfi Tmpr eona Dactyrioceras commune, 6pnurnofi croporrbt, 46 - sn! c aegofi croporor).
P. Bxapf,
crp. 17 (+a - BHA c
(Drr. 5-7. Acrocoelites ianensclli (Ernst) . . . . . . . . . .. . . . . . . . . . . . crp. 2g 5 - poctp l& 8?-25, lutxnufi loap, soHa Dactylioce.u"-.omun", O. Map:ra (Sa - ana c 6prcrunofi cropolrbtr 56 - aua c rpaaoft croponu)j 6 - pocrp t* gfZ+, Hnxnnfi Toap, p.Mapxa (6a - BnA c 6prculno$ cropoHbr, 66 - suA c aegoft croponu); Z_p*rp M a7-27, mxnufi toapr eona Dactylioceras commune, p. Mapxa, nonepeqgoe cetreg11e y Bepu[Hbr aJrbBeorrbt
. Onr.
TAENHUA III Acrocoelites trisulcosus (Simpson) o . . l-$. . . . . . . . . . . . . . . . . crp. 12. poc" ]'is 87-10, 1nrxxrfi Toap, p. Kemrvap (1r - Bra c Opto,uxon croporarrl6-BnA co cnf,Hllofi ctoponrr, le - BnA c npasoft croporrr); 2 - poctp N B7-g, ror**"n soHa Dactylioceras commune' p. reahll Kegon "oap, dpnulrot croponur26-B.nA V:; ""o. c aesofi qoponr'r); gz-lg, pocrp l.Ie 3 aep:crdfi Toap, p. Kenn14ry (sa _ *n.p.*_ Hoe @qeslre y Bepurutlbl albBeoJlblr 36 - nonePeqHoe oeqexEe B npuaepgrnxnOfi vacru).
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cTp' 52 . . . . . . . . . . ' ' ' ' ' 4-7. Mesoteuthis tiungensiSsachs sp. nov. .-. p. Brrloft (4a-anA commune, Dactylioceras g?-g1, HuxH#;Ip, 4 - ronorur Ns "o"" HnxHnlh Toap' 5 - PocTP ls 87-82, c 6puunofi cTopoHbr, 46 - snI-c nesofi cTopoHbl); (Sa - BuA c 6pgunofi cTopollblt 56 - snA c p. Bnnrcf, commune' O*rilioceras "."r nuxuuft Toap, soHa Dactylioceras commune, 6 - pocrp ls 8?-84, aeacif; cropoxrr); - srg c absofi cropollbl); 7 - pocrp cropotlr1r..66 6prcurnof, BnA c p.-g"".O'(6a braunianus, p' Bnarofi' nonepeqHoe ceqeZugodactylites mrxHufi Toap, solra Ne 87-a9, x2' anuneonH, y BepuHHbl HHe TABTIH UA
IV
I crp' 19 " " " " " @un. 1-4. ActocoelitespolatltsSachssp' nov' .. " " " comm-un€, p. Mapxa(la-ena Dactylioceras g7-L4, .o"" n"*""i"ro"p, M noJrqrxu I aepxrrrfiroapt 2 - pocrp }16 8?-16, c 6pnunofi cToponbr, 16 - BI{A c nesoft cTopo'bl); BuA c aesofi cropoHbl); 3 - poctp p. Kennusp (2a - BHA c 6prorunofi cropollblr 26-p. Mapxa (3a - sxA c 6pnuuofi ialcifer, nnxnufi Toap, eorn Harpo..r"" lS 8?-1S, gZ-8S, turxslfi ToaPr eona Dactypocrp-fin 4 cropor6t, 36 - sxA c nesoft cropo*bl); y EeplultHbl aJIbBeoJIbt'46-noCeqeHue (rla nonepeqHoe p. Map)G commune, lioceras qacrn)' nepegnoe cevenl{e B gpuBepunnnofi ' crp' 42 .. " ' ' ' ' " ''' " " " Mesoteuthispyramidatis(Zieten) Our. 5-?. - euA c 6prounoil cropol|bl, (5a p. Kearrvnp Toap' aepxxuft 5 - poctp N9 87-61, eona Zigodactylites mrxsnft sap' e - po"-tp l'te 8?-63, .tlp*itl; 56 - auA .'"*.f cTopollbl, 66 - Bua c aeaofi cro6prcruno* (g" trq eaans Axa6apcrorfi braunianus, ceqeglle " ,orxrnfi aanelrr Ana6apcrorfi saltrto, nolepeqgoe ponu); Z - pocrp l1n 8?-65, y BePuHHbI anbBeoJlbl, x2. TAENHUA
V
' ' ' ' ' ' ' ' ' ' t... . e U f . 1 - 3 . A C r o C O e l i t e s t r i s c i s S u s ( J a n e n S C jh. ). . p'Mapxa Dactyliocerascommun-€' soHa Toap' **"tf 1 - pocTp l\$987-20,
CTp'20 (la-arA
c 6 p n u r n o i l c T o p o H b l ' 1 6 - a u g c r r p a a o f i c r o p o l l H ) ; 2 - p o c r p } s 8 7 - 2 2 , t g i x H s f i T- o aEua p ' c negol (2a - Bsa c 6plorunofi cTopoHbl' 26 eona Harpoceras falciferr- p. Mapxa p' Mapxa (3a - nofalcifer' H"tpoceras Toap' ,.'t*ilfi pocrpttngz-zg, ,cropoxrr); tqac?u]' "o"t 36 - nonepe'rH;e ceqeHre B llprBepususofi nepeqHoe ceqeHlle y B€pururbt aJIbBeoJIbIr 2'7 o''CTp' t''''' s m o l O n e n sSd as C h SS p . n O V . - . . . . . ' ' e n f . 4 - 8 . A c r o c O e l i t eO Kep. flegul braunianus, Zugodacty_lites gz-3+ o*r"*--.pr'""ra 4 - roJrornn ls - poclp N8?-35' cTopoHbIr-46 - BHA; aesofi cTopo'b'); 5 '.o"" AoH (4a - suA c 6pnuxofi (5a - Bn! c (flpxcer*6rpr) Ea11er'r6ypr spinatum, Pl"uro"ur"" aepxnxf; *nrJO.r,, TGP' aeprurnl pocTp l$ 87-32' 6 croponrr); 6prcrunofl cToporrbr, 56 - nnq c nesofi - PocTP 7 cTopoHbl); --srlc npaaofi p. Keanusp (6a - BI|A c 6ploruuoil cTopoHblr 6e -r;un c rpaaol (7a - ang c -6prcrunofi cropoHblt 76 Toap' p. Byop-eercrr l* 8?-33, braunianus' P'ToZugodactylites soga uuxnnfi to"p, 8 - pocrp ]$ 8?-36, cropoxu); xL aJIbB€oJIbIl y BepurIHbI CeqeHue K}P-rcpsxi nonepeqHoe
TAEAI{UA VI crp' 32 ' ' ' ' ' ' ' ' ' ' ' ' ' ' ' ' nov' Onr. 1-3. Mesoteuthispergrand'rsSachgsp' t(ap.Jleoarfi rcsenlirantzi, Pseudolioceras ooua gz-3g, aepxxrft map, 1 - rorrornn ls ED(2 - pocrp ls8?-4o' - suq c resoil cloponr); t6 -opo""r, 6prcuurofi c auA ao' (la I cnopoEbL 26 - sunc rcnofi cropourr)i (21 - Bf,A c 6nlynon Eufi aaneu, Olf.l(*-.pcx 6fibg€oJlHo aeplllf,xbl @qegse rroneperlnoe ] rrurnnn aarEII' p. Jlenrrfi'KeAoE' pocTp lrft 8?-39, cTp'40 ' ' '' '''' vorcnez............. Qgp.4-5. MesoteuthiSlaptinshaiae coT1une' p' TpHr (4a - ana Dactylioceras soga Toapr unxnufi 8?-5?, l1n pocTp 4 Ttoap' .oEe m*,sil 5 - pocrp N 87-58' --su,.c npasofi croponu); c opnrunol sr.opo'bl, 46 x 2' ry6ai [onepeqEoe ceqeEe y E€purf,lrbl alEagoJr$ fuia6apcias Dactyliocer"" "o*rune, ' ' Clp'go ' 'o ''o' . .S. .) .. . . . . . . . @ U r . 6 - 8 . M e s o t e u t h i s s t r i o l a t a ( P h i l l .i p p' Mapxa (Oa. - ann commune' Dactylioceras eoHa t"Lp, o*"ttr 6 - pocrp N 8?-52, N 8?-54'Err@ltoaPt 7 - pocT c 6prcrunoft cTopogbt, 66 - esq c rpagofi cToponbl); - BEI (la- - snA c 6prcurrol cropOnr'76 p. Kegog Jleaufi commune, soHa Dactylioceras commune, Dactylioceras g?-51, soHA Toap, g *rxsd - pocrp ]16 c aenofi x2 p. Trcnr, "[poor); noneperrnoe cerlen[e y BepunHbl SJIbE€oJtbtl
r86
TAENHUA
VII
Onr. 1-2. Mesoteuthis aff. pergrandisSachs sp. nov. . . . . . ., . . . . . . . . crp. 94 pocrp lQ 8?-41r 1aepxuuft Toapr eona Pseudolioceras rosenkrantzi, p..neasir Kenon (la - sHA c 6prouno* cropoHbl' 16 - srq c npanoficroporor);2-pocrp lSgZ-42, tlrxuufi aanen, n.os Yprcnn-Tyr,ryc (2a - BHA c 6prcurnoft c"oponbr, 26 _ sng c aeaoft cropoxrr). Onr. 3. Mesoteuthis laptinshajae Voronez. . . . . . . . . . . . . . . . . . . . . . . C 4 l 4. 0 Pocrp lS 87-55, aeprurlrft Toap, p. Keaur,.rnp (Ba - BnA c 6prorunoft cTopoHbt,36 - sHg c npaaoft croponrr). Qnt. 4-7. Mesoteuthis longirostrrs Sachs sp. nov. . . . . . r, . . . . . . . . . crp. BS 4 - nqnorlln N 87-43, nnxxufi Toap, soHa Zugodactylites braunianus, p. Bnaroft (4a - BHA c 6prcurxofi croponbl, 46 - ana c npinotr croporrlr)j S - pocrp lr19g7-46, rurxnnfi Toap, goHa Dactylioceras commune, p. Tronn ( ba _ BuA c 6pnrunoft croponbr, 56 - sna c aenofi croponsr); 6 - pocrp l& 87-44, sxxnnft Napr etua Dactylioceras comnlune, p. Bnanft (Oa - BnA c 6prorunof, cropolrbt, 66 _ nna c npanoft croponrr)J 7 - pocrp l$ 87-49, tllrxnnft map' soHa Dactylioceras commune, p. Bnnpfi (Za - noflepeqHoe ceqeHre y Bepul{Hbt aJIbBeoJIbI , x23 76 - nonepeqHoe ceqeHne B npgBepunHHofi qacrr, x2).
T A E N HU A V I I I O n r . 1 - 3 . M e s o t e u t h ^ i s , p s e u t d o e lS on a st as p . n o v . o . . . . . . . . . . . . . ag ch crp.44 I - nonornn.lt 87-66, aepxnnfi nnnHc6l:q sona Pleuroceras spinatum, Eamev6;pn (Jl$cev6ypr) (la - slra c 6prcunofi croponbr, 16 - aua c aenoft z -iocrp t$ 87-68, unxnnft Toap' sona Zugodactylites braunianus, p. /lenslfi".oporr"r); Kegog (2a - BnA c l6ptounoft cropoHblr 26 - sua c aesoft croponal); 3 - pocrp Ne 8Z-ZO, lrsxHnft Toap, 3oHa,Zugodactylites braunianus, p. fleBbrfi KeAon, nonepesHoe celrenr{e B [pnBepuunnoft qacTIl. Qnr. 4--7. Mesoteuthis aequalis Vorcnez. . . . . . . . . . r . ., ctp. 4g . o . . . . . . 4 - pocrp lt 87-76, nxxHnft Toap, soHa Dactylioceras commune, p. Bxarcft (+.-rnq c 6prcunoft cropoHbr, 46 - rrng c npaaofi cropourr); S - pocrp IIe g7_7S, rurxuuft Toap, eoga Dactylioceras commune, p. Ehrnnft (5a - BHA c 6proruHoftcropogbr, 56 - exg c npaBofi croponu); 6 - pocrp },ie87-78, nuxHnfi Toap, soHa Dactyiioce."" .orrun., p. Bnarcft (Oa - BtlA c 6pnrunofi croporrbr, 66 - anq co crruHHofi cropor6r, 68 - Br{A c nesofi croponst); 7 - poctp N 87-80, Huxunfi Toap, soHa Dactylioceras commune. p. Trcnn, noneper{Hoe ceqeHlre y BepuHHbI aJlbBeoJrbt, x2.
TAENH UA IX
Onn. 1-4. Paramegateuthis nesciaNalnjaeva sp. nov. . o . o . . . . . . o . . crp. 60 1 - nonoTnn lt 8'7-J 42, nepxuuft 6ar, noo VprcHrsTyrayc (la - Bua c 6prounoft cro-poHblr" 16 - nun co clrHHHoft croponrt, 1o - Br{A c aeeoft croponrr);2-pocrp Ne87-147, aepxuxft 6ar, rr.oo YpronrrTyrayc (2a - BlrA co cnr{HHofi ctoporor, 26 - B}rA c aeaofi ctoponrr); 3 - pocrp lt 87-146, uuxnnft xenaoneft, sona Arcticoceras kochi, Aua6apcran ny6a (3a - BHA co cnuHuoft cropoHbr, 36 - er{q c npaeoi croporcl); 4 -'pocrp ls 87-149, sepxnufi 6ar, n.-oe Yprcnr-Tyrvlyc, nonepeqHoe ceqeHHe y BepurHHbraJrbBeoJlbl.
Our, 5-8. Paramegateuthis manilesta Nalnjaeva sp. nov. . . . . . crp. 62 5 - ronorHn Ne a7-L48, eepxnuft 6ar, n-oa YpnHr-Tyrrayc ( 5a - Br.rAco cnnsuofi cro. poHblr 56 - eua c npanoft ctoponrr); 6 - pocrp IIe 87-150, nuxsnfi 6ar, eoxa Boreiocephalites pseudoborealis, n-oe Yprour-Tyuyc (6a - BHA co crmxsofi cropoHbrr66-nuq c npanoft cmpoHu); ? - pocrp ls 87-151, Huxurift 6ar, sona l3oreioceplralites pseutloborealis, n{B Ypronr-Tyvryc (?a - BHA co cnunuofi croponst, T6 - BHA c negofi cropoHbr); g - pocrp Ne 8?-154, nepxHufi 6an n
187
TAEJI14 UA
X
. . . . . cTp. 78 @nr. 1-3. Cylinilroteuthis (Cytindtoteuthis) confessa Nalniaeva'sp. nov-. cropoHbl, (1a 6prouttofi BI{A c p. 6ar, Selalr: bepxnufi 1 - ronorHn l.I9 8?-157, aepxunfi 6al, p. Seron(2a-srq n.*-n cropoHbr); 2 - pocrp llft 8?-159, 16 - BnA sepxrrft " 3 - poctp lS 87-155' n c npaaoft cropo6u); c 6progrnofi cTopolrbl, i6 " 6ar, p. 3elo{r: nonepeqnoe ceqenl{e y Bepu[llbl aJIbBeoJIbl' Cfp'70 o. ' ' '' ' ' ' '' Q n f . 4 : . 6 . P a r a m e g a t e u t h i s p t e s sNaa l n i a e v a S p . n o v . . . . . (4a - BI|A C 6prcutuOfi cro6ar, non Yprcxr-Twyc 4 - pocrp rr\rnaz-rs8, "p.a*fi 6arr n"oa ypicHr\5 - nonornn lS'97-L}4rcpeAHrfi po'br, 46 - g1r1ac aesofi croporor); - snA c neaofi cropoHbt, 5a - nonepeqHoe 56 crcrpoHbt, 6prcrunoft (5a BxA c Twv" ( 6a - snA aepxnnfi 6at, noo Vpnnr-Tyr'ryc po"tp"h 6 - pocrp 1168?-135, ceqe're c 6ptourrofi cropotlbtr 66 - ang c aesofi cloponrr)' crp.68 va i SP.ngv. ...!....o. O n f . 7 - 1 O . P a r a m e g a t e u t h i s p a r a b a i o s i c u s Naael n (7a - BnA c 6ptounofi cTopoHblt ny6a 6aftoc, Axa6apcxas poctp ls 87-129, ? 6afioc, Ana6apcraa ny6a lS 8?-126, 8 - roaotrn ?6 - sng c nesofi cropot$t); 9 - pocrpNST-13O, (8a - BxA c 6prcrunofi cTopoHbt, 86 - sun c npaoofi croponrr); 96 J strA c aesofi cTopollblf ( BHA c 6prounofi 9a Vprcnr,-Tyrr'ryc nrxnnft 6ar, n-oa flafioc, Axa6apcrcas rY6ar noneperrgoe cerreH[e y 10 - poerp I$ 87-132, cropogg)i BepuHllbt €ulbBeoJrb!.
T A E N H U AX I ' Crp' b4 o " t " o.... s S t o m e S O V. . . . . Q g p . 1 - 4 . P a r a m e g a t e u t h i s i s f t m a n sGdU (la -.BnE c 6prcunofi cropony6a Ana6apcxan 6at, aepxnnfi pocrp la 87-118, 1rur, 16 - BuA co cnunHoft croporor, lB - anACneoofictoponH);2-Pocr?IS87-119' Yprcur-Tyvryc (2a - ElrA co cnnsnofi kochi' n{B ruaxnnfi rennoaefi, eoHa Arcti"o.u;3" ruxnufi reuogefi,soga pocrp lt 87-123, 3 croponbl); nesofi suA c cropoHbr, 26 qDopoHbr' 36 - suq c npaaofi cropoishmae, p.Hxr"ra (3a - BuA co cnnxnofi Arcticaeras noa Yprcnrkochi, nrr); 4 - pocrp I* 8?-1 24, unxnufi xenaoseft, soua Arcticoceras x2' y arbseoJtbl, BepuHHbI no[epeqnoe ceqeHrre TWyc, timAnensiS GUSIOmeSOV, . . . . . . . . . . . . . . . . Cfp' 66 PArumegateuthiS Offf. 5-8. cropr c 6p'uxoft aepxunfi 6ar, Ana6apcxan ry6a (5a -."tA b - pocrp l\& 8?-11O, totr56-B!,rAcocnuHHoftctoponu'5s-Bl{AcaesoftcropotlH);6-PocrP}S87-114' 66 - BrA c npasofi aeixirnft 6ar, Arra6apcrorR eanua ( 6a - BHa co crnrnnoft cropoHblr (7a - BnA co crIHHnySa Aua6apcran 6ar, oepxHnft 7 - pocrp lle 8?-113, ctopo6r); ruaxlilrfi xerutoaefi' 8 - pocrp I'fe 8?-113, nofi cropogst, T6 - Bxa c npaaoft ctoponrt); y BepuHHbt atlbBeonbl ceqeHHe noneperlHoe ry6a, Ana6apcxae kochi, eoHa Arcticoceras
TABflH
UA XII
. . . . . . cTa.' 72 Offf. 1-2. Holcobelus unarettsis TuChkov. . . . . . . . . . . . . . . . 6pnuuofi cTopoHbtt ( BHA c ls p. Tatrtnuqan I - pocrp N 87-2 16, aepxnnft roap, Hnxnnfi aaaen, p. Mynyrya)fiaK' 2 - pocrp Ne 8?-217, 16 - eHg c aeaofi .toporrri); nonepeqHoe Cer{eHHe y BepunHbl aAbBeOJIbtlx 2' .
enf.
Crp' 76 . . . . . . . . . . . . . . . . . . 3-4. Holcobelus hirnsot,i Saclrs Sp. nov. 6prcugofi (3a BuA c Vprotlr-TyN{yc n-on toap, aepxnrnii 3 - roaorun l..1na7-218, aepxnufi roap, P'Ke- pocrp Ne 8?-219, cTopoHbt,36 - arAc npaaoft cTopoHbl); x 2, 46 - nonepeqHoe ceqeaJlbBeOJIbI, y BepuuHbl (q" CeqeHne nonepeqHoe rr*tp Hne B rrpnBepu]lnuofi qacrx).
crp' 76 ' ' ' ' . ' ' ' ' ' ' ' O n n . 5 - 8 . L e n c t b c l t t sl e r r e n s i . s G u . s t o n t e . s o r ' . cropoHbl' 6prcunofi (5a BHA c p' Keaxunp roap, aepxHxfi 5 - poctp Ne 87-161, 6 - pocrp I'ie87-1 62t 56 - ent co cnHHHoft croporor, 5a - BHa c aeeoft croposrr); - enA co cnnHlloii cro66 cTopoHbl' 6prorunoft ( BnA c 6a p. Keanltep Toap, BepxHnft roap, sona l)act,vHuxrmft 64, pocrp N 8?-1 7 polrbr, 6s - er{A c aeeofi cropoxu); - e,ua co cnuxrtofi lio<:eras comnlune, p' Bnaroft (ta - Bt'tA c 6pnuuoii cropoHbl' 76 oepxurft Toapt P' Ke- pocTp Ne 87-165, cTopoubr, ?B - eHA c npaaofi cTopoHbl); I r2, noneper{Hoe cer{eHue 86 a4'Beorblr rnni"p (8a - nonepeqHoe ceqeHge y BepuHHbI B cpeAHeft alacru PoctPa, '2 )'
t88
TAE/IHUA XIII ?
o n r ' 1 - 4 ' L e n o b e l u s ' u i ' l i g a( se an cs hi ss ) ... o............... o. c r p .g 6 1 - rororrn ]t 87-166, sepn*rf, P. Kearvep (la - .* . Srorunoi cr.po*r, 16 - snn co cmrnnofi c"opollbl' la "oap, - BnA'. o"r"n croponar);l - po"rp ls g?-167, aepxxnfi Toap' p' Kemruep (za - B.trA c 6proruror cropo'br, 2e - BE! crqnnor croponbr, 2s - Bnr c rpaaoil croponu); s -'p*rp ls gz_16g, eeprurtr "o roap, p. Kenr_ Mcp (ga - BuA c 6prcrunoftcroporrb[, 96 - BrrA co cmrnxofi cropot6t, BE _ BnA c reF sofi croporor); 4 - pocrp lS g7-l?q aepnn-l Toap, p. Kerurr,rqr (4a _ nonepeqgoe ce.,e'rre n6rrex Bepu''Hbr sJrbB€oJrbrlx2, 46 nonepe{H.e ceqeH,,e B cpe[trefi sacr' pocrpa, x 2 ). < D r n5 - 7 ' M e s o t e u t h i s i n o r n( P a thau l l i p s ) .... o o... o. o......... cttrt.47 5 - poctp lS 87-?1, HrxHnf, €rarreH,p. nend Kegon (Sa _ Etra c Srournol c1.gporrbl, 56 - sr! c npanofi ctoponsr); o - pocrp l.Ie 8Z-?g, eepxnnf dn*, Aua6apcrar ry6a ( 6a - BrA c 6proufiroftcropoHbt, 66 - st|n c npaao* croposu]; Z _ pocrp t* g7_24, aepxnnfi Toapr p. Kemrrrarp,loneperl'oe ceqeHre y Bepu,,Hbt arlbBeorDr. TAENHUA XIV Qxr. 1-4. Lenobelussrbiricus (sachs) . . . . : . . . o. . . . . crrp. g9 1- pocrp lrleBZ-127, t*txrrtrfr,aaJren,p. Keanurp (la _ . . . . . o. . BnA-c-6pnruuof, crolronbr, 16 - Bn! co cnHHHoficropot|blr ls - t"r. rp..ofi croporur); 2 -pocrpl\&g7-176, aepxnnfi T@pr p. Keanrraap(2a - B_HAc 6prculrof, cropotrbt, 26 _ s|tg co cmrnnof, cm_ pot6l, 2s - arA c npaaoft ctoponrr); 3 rononrn l& 8?-b, Hnxnrfi aaneg, p. E]rop(3a Sers{r - B.nAc 6prournoftcroponbr, 96 _ BHA co cnnnnofi cropoxbt, Ba _ 6gg . aeaoft cropolor); 4 - pocrp ]$ g7_lig, roap, p. Kelruqr (.la _ nonepeq_ Hoe ceqeHlte y Bepultl|bl aabBeo'blr x 2145 ".p*"rfi - nonepeqxo€ ceqerrne B np116epuxrnofi .1acrr pocrpa). Qur. g-7. MesoteuthissuDrostrilormis Voronez. . . ' 5 -- pocrp l$ 8z-g6, Hnxnufi roapr so'a z";;;";yfi,r" ur""ri;";", i;;"r:fi; i:_ 6a (5a - BtnA c 6prcurnoftcropotbr, 56 _ srA c_rpaaofi croponrr);6_poczpl\&g?_gg, aepxnxft Toap' p. Kemrvep (oa - BIiA c 6prourno[ctopotGr, -oo --BnA c rreBotrcropc. nH); 7 - pocrp ]t 87-s9r Hxxsnfi roapr .on Zu-jJ"",yiirJ" bn"uni"nu", Ana6apcraeny6a, nonepertnoe ceqenr{e y BepurnubraJIbBeoJIbr,x 2. TAENNUA XV onn' 1'3' Lenobelusminaeuae (sachs) . . . . . . . . . . . . . . . . . . . . . . crp. 94 1 - nonornn It g7-L79, aepxxnft Toap, p. Kennr,,rep( 1a _ ,"o t 6prournol cropotGr, 16 - sra co cruilrxoftcropoHblr,ls - r"a-. rpaaofi croponrr);-2 - pocrp l.fggT-1g1, nnxnnft aaJIeH' p' Eyop-3erq{r (la c 6iro'rHofi croporbr, 16 - ang co cmrxnofi ?"4 cropoHbr' ln - snA c npano* croporol); 3 - pocrp M g7_1g3, HrrxHnfraare', p.KeJIHMsp ( 3a - nonepeq'oe cerreHxe y Bepur'Hbr aabBeoJrbl, x 2; 86 - noneperHoe ceqeHr{e B npr{BepurnHnof,qacru pocrpa, x 2 }. Qnt' 4-7. Lenobelusuagt (sachs) . . . . . . . . . . . . . . . . . . . . o . . . . crp. 91 4 - roaotnn lt 8'7-L71, aepxnnft Toapr p. Keanraep (+a - Bra c 6pnunofi cropo'br, 16 - sua co cm{HHoficroponrr, 4s - *g-" npaBgft ctoporor); 5 - pocrp l.6 g7-172, eepxHufi Toap, p' Kelnvsp ( 5a c 6prorunoRcroponbr, s6 - s'a co cmr'Hofi ?HA cre poHbr' 5s - suA c npaeofi croponrr); 6 - pocrp r$ g7-173, aepxnnfi roap, p. Ke.nn_ Msp ( 6a - anA c 6prourHoftatlpo*r, 66 - s'a co cnlr'Hofi croporor, 6a - B,,A c [paaoft croponu);.7 - pocrp lt g?-lZS, HuxHnft aanert p. Eyop_eerr{T (7a _ nonepe.r_ Hoe ceqeHHey BepunHbraJrbBeoabt, x 2, 26 _ nonepeqHoeceqentaea cpeAHefiqacrn pocrpa, x t ), TAE NUUA X\I O x n .l - 4 . P . s e u d o d i t o e hl ii tdegsi e (t tSi a c h s ) .... ............. . . c T p .g ? 1 - noaotxn M 82-L, nnxnnfi aaJreH,.p. Eyop_Oerolr ( la _ gnA c 6prcuruoficropoxrr, 16 BHIrco cm{HHoftcroponu, 1a - sua c aenoft croponu); 2- _-pocrp }$ g7_1g7, xr'xuNft aaJIeH'p. Eyop-3ers{t ( 2a - sln c 6pr,ouHoficropoHbr, 26'-;;;;r-";;; cropo'br' 2s - au' c npaaoft cropoHrr); 3 - pocrp Ne gt-3, xnxsufr aareq p. Eyop_ Sexnr (3a - nonepeqHoeceqeHre y Bepu[Hbr aJrbBeoJrbt, x4, 86 _ nolepeqHoe ceqeHrre a cpeauefi qacrn pocrpa, xt ).
t89
' ctp'1O9 o " " P s e u d o d . i c o e l i t e s p r i m o r iS sachssp. nov." " " Onr.4-5. yprcHr-Tynnxnufi ro"p, eoua Dactylioceras commune, n-oB 4 - ronorun Ne g7-2O7, ' BHA c IIpa4a croporct, cmiHEQfi rJuA co 46 cTopoHblr sul c 6prcrurrofi Myc ( 4a rruxnuft Toapr soHa Dactylioceras commune, nofi croporur); 5 - pocTp ]$ 87-208, cToporblt 56 - sun co cnugHoft croponu' 6prcurHoft (Su BuA c n..oa YprcHr-Ty"ty. x2l' nonepeqnoe ceqeHl{e y BepuI{HbI arbB@Jlbt, Ss - BHA c npanofi cropogbr, 5r
TAEJlHUA
XVII
crp' 99 .. ' o .. .. " " O u r . 1 - 4 . P s e u d o d i c o e l i t e s p l a t y u e n t r i o s u sS a c h s ' " " p. Kennvnp ( 1a : Bt{A c 6prcunoft cTopoHbl' aaaeq aepxnufi ls 8?-189, roaorun 1 2 - poctp Ne 87-194 - anA c JreBofi croporbr); 16 - nuA co cnuHHofi cTopoHbr, ln
I ,
f
nnxtrrrfiaafleH'p.Fyop-Serolr(2a-nulc6prcrunoftcTopoHbl,26-snq'cocnngrrofi rurxnnfi aaIeH, P' Ke3 - pocrp }t 87-190, - Br{A c reBoft cropourt); ;;;;-i" 3s - BrA c Jlecropoult' cnnHgoft co ann 36 nr{Msp (bt - BuA c 6progruoft cTopoHblr nuxnufi aanelq p. Kerunvr*p (4a - nonepeqHoe 4 - po"tp l\trg87-193, oof, croponrr); - nonepe5Eoe CeEIeHUeB CpeAHef, qacTH ceqeHue s6JrI{oH Bepur]rHbr aJIbBeoJIbI, x 2r5, 46 Pocrpa, x 2r5), ' ' ' Crp'57 g s p-. S - 2 . M e s o t e u t h i s b r e u i r o s( dt rt rOs r b i g n y. .) . . . . . . . . . . . . . . - snA c rpa56 cTopoHbl' 6porunoft (5a c nxA Toap' p. IIurrAe 5 - pocrp l$ 8?-9O, p'JIemrxHl{ft Toap' so'a Dactylioceras commune, lrtrn87-93, 6 - pl"tp aofi croponrr); - pocrp 7 ctoponrt); BIiA c rtpaaoft 6prournofi BHA c Bbrft KeAoH (6a "topon"rr'66 x2 Toap, p. JbrHae: nonepetrHoe CeEIeHUe y BepuIuHbI a'ItaeoaH, ]..Ieg?-g4,
TAEJIHUA XVIII crp.1O4 i da. ce hs s s p . n o v . . . . . . . . . . . . . . . Qun. 1-4. pseudodicoelitesclauatoS (1a -, BBA c 6plqllnol cropop. Eyop-oerqr aaaen, unxnuft 8?-19?, Ne noroTHn I }t87-199' - s1rAcnesof,cTopoHH);2-pocrp nbr, 16 - BI{A co cru{H1loficropotot, 1g cTopolrbl, 26 - su1' co cm{HHofi cronuxrrnfi u.rr"", p. Kenuusp ( 2; - "oA c 6prcruuofi g - poctp I'le 8?-198, unxnxf; aaneq p' Eyop-3epoHbr, 2s - r}ul c neeoft croporrsr); 3e - BHA c rpacropotot' crrrrngofi BnA co 36 rs{T i3a - BuA c 6prcrunofi .topott"r, (44 - nonepeqnnxrurft aanell p. Byop-3e;olr 4 - pocTp N a?-2O1, aofi cropowrl; B [pnBeplutllF: cerreHle noleP€qHoe x2, 46 alrbBeonblr Hoe ceqegue e6nnsn BepuIrHbI Hofi .racrn pocTpa, x2). ctp'14 ' ' ' ' '' ' ' ' ' ' ' ' 't' H o l c o b e l u s g t a u i s( G u s t o m e s o v ) Qnr.5-6. - BIIA c 6prcunof croeepxHuft aaaen, Ana6apcrorft aarnrs ( 5a 5 - pocrp l$ 87-213. nnrcrnft aaaen, p' Eyop6 - pocrp It 8?-214 pbnu, 56 - aug c rrpanof, croponrt); -g"*, 6s - nouepeqHoe cropoHblr c npanoft eun ( 6a - bng c 6pnu$roft cropoHbr, 66 aarseonar). y Bepurrubt ceqenne
TA6 Nh UA XIX Our.
ctp' 1O2 . . . . . . . . o 1-4. Pseudpdicoelites hibotitoides Sachs,. . . . . . . . . cro( 6prcuxofi B}IA c 1a aaJn{a g5-6, Ana6apcrorfi aaJrert aepxHnft 1 - noaornn l$ ]$ 87-221' c IIpaDofi croponrr); 2-pocrp porbrr 16 - snq co cm{HHoft croponu, ls-auA 26 - Blnl co cnnnnofi Hnxrdrft aaJreg, p. Morop,ryna ( 2a - sul- c 6ptotunof, cropoHblr rnrrcrnfi aaIeEt p'Kepocrp lS 8?-195, 3 cropoHbr, 2s - auI- c npaaofi croponu); BilAeH Qparr.aoxox c cn@uot"t c aJIbBeoJIbt' o6racru s ceqenue (-Su ngnepeqHoe nr{Mnp 'lacfir pocrpa). 6prcurnofi cropoHbr, x4, 36 - nolepegHoe ceqeHue B cpeAHefi
' Crp'1O7 o. " ......... @ U n . 4 - 6 . P s e u d . o d i c o e l i t e s g u s t o m eSs ao cv hi s s P . n o v . ( 4a - BnA c 6prcutnofi eaJrEs g.7-2O2, Arra6apcrorft a€ureH, nepxuuft noaornn l$ 4 - pocTp croporrtr, 4s - BlrA c npanoft croponst); 5 cTopogbl, 46 - ;1nA "*""onp. Keanvsp (5a - BlrA c 6puunoft croponbl' 56 - BUA co "o roap, aepxtruft lt 87-204, eepxrurfi aalet4 6 - pocrp }$ 87-205' cm{HHofi cTopotlbl, 5e - aHA c aenofi ctoporst); y BepuJItHbI aJrbBeorbt' 66 - nonePeqHoe Ana6apcrcrfi sanus ( 6a - nonepegHoe cetreHHe ceqexne a cPeanefi tlacru PoctPa)'
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coAEP)ItAHUE
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BBEAEHI4E Haacerreftcrao Cylindroteuthaceae Sachs et N a l n j a e v a ,
Cel.reficrsoPassaloteuthidaeNaef, 1922 _
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1970
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floAcepreficrao Megateuthinae Sachs et Nalnjaeva, onpeaenurerbHac ra6rru[a poAoB ,roa..rlfi.rea
PoA AcrocoelitesLissajous, 19-15 PoA MesoteuthisLissajous, l9l5 .
196T Megat"uit,in*-
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.:'. Poa HolcobelusStolley, I92T ". ; ; , ' r e ; 6 HaaceveficreoDuvaliaceaeSachsetNalnjaeva,l970 Ceveficrso Duvaliidae Pavlow, |!ld . : . . florceueftcrso PseudodicoelitinaeSachs et Nalnjaeva, 1967 . . . . OnpeaeaurellbHasra6nuua BuaoB noaceueficraa Pseudodicoelitin ae Poa LenobelusGustomesov,1966 P o a p s e u d o d , i c o e l i t e s S a1c9h6sT, . .:....... CHCTENTATHKA IOPCKHX H ME IOBbIX EEJIEIV1HHTOB
7 7 7 7 8 I 29 59 72 80 81 81 82 83 96 LL2
CTPATIIIPAOI4T{ECKOE 3HAqEHHE P,U{HE- H CPEtrFIEIOPCKHX EEIIEMHHTOB.
O OHJIONEHHH MEGATEUTHIN,AE 14P S E U O O O I C O E L I T I N.A E . OE YCJIOBI4flX )I*13HU MEGATEUTHINAE 14PSEUDODICOELI TINAE . TEOTPAOHTIECKOE PACNPOCTPAHE HI4E MEGATEUTIIINAE H PSEUDODICOELITINAE.
}4(A3ATE'Ib BI4trOBTJX HA3BAHHHMEGATEUTHINAE H PSEUDODI,COELITINAE flHTEPATYPA .
i f
131 L4oL47 158
OOTOEAENHUbI
160 166
ObA.HCFIEHI4ffTAENHU I-XIX
185
Bnannrrp Hrroaaeam Catc, PAHHE-
TaGsEa l{garcara Hansrreaa
H CPEII}IEIOPCKHE EE'IEIT4HTITHCEBEPA C@P. MEGATEUTHINAEI,I PSEUMDI@ELTTINAE
Yraepxaelo r EeB&Tr t'hsrrrrnorr nF Drsr r reo{rm oraenerrr Aranorrr rayr C@P Cr6rpcuo Penarrop rsgaTqrtcrEa B.C. Bam. Xyucraernd fermrecnl PeaarroP F.tl. Kqerrra Ilorrcaso
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l(rrne lsaara orPerrnrr cro6oDr 'H4/Ia'. 103?1? IEf|, Ir|ocpa, |(-fl.2, Ibmoccgcrrt mp.' 21 }lsAn.reJecmo ,Hryra,. 199034, flergpag B-34' s-' |@r' 12 EsIa?g'trct.Da 1-r rruorpaltsr
